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1 , Department of Anatomy and Histology, University of Sydeney NIDCR, NIH, USA NIDCRNIHUSA University of California, Davis, Dept. of Animal Sci ISAS/JAXA , IBDML, France 1 2 1,2 1, ) ) Univ. of Minnesota, GCD , , , 1,2 1 2 PD 1, PD Department of Cellular Biology, University of Georgia

2 , Laboratory of Cell Signaling, Stazione Zoologica Anton Dohrn RCAI JT , SPring-8JASRI ,

3 JSF , , Department of Pathology, Emory University 1745 CDB

4 ABRC , , , University of MarburgAnimal Physiology 1, , ,2 1 2 PRESTO

5 , , ,2 1, University of Rouen University of Hawaii Slovak Academy of Sciences HHMI/University of Minnesota

6 Univ. of Minnesota ,2 1 1 National Institute of Child Health and Human Development Rahman Farhana Shahjahan MD , , , , Universidad de La Habana

2 R K/S K/S K/S K/S K/S K/S K/SR R K/S K/S K/S K S R K/S K/S K/S K/S K/S K/S

2 R K/S K/S K/S K/S K/S K/S K/SR R K/S K/S K/S K S R K/S K/S K/S K/S K/S K/S Graduate School of Environment and Information Sciences, Yokohama National University, Yokohama 240 8501 Faculty of Education and Human Sciences, Yokohama National University, Yokohama 240 8501 Keywords:

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