櫻井2008p1-8v2
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1 The Bulletin of the Hobetsu Museum, no. 23, (March, 2008), p A review of the mosasaur fossils from Hobetsu and around area. Kazuhiko SAKURAI Hobetsu Museum; 80-6, Hobetsu, Mukawa-cho, Yufutsu-gun, Hokkaido, Japan Abstract This is a brief review of the mosasaur fossils in Hobetu Museum. Those are from Hobetsu and around area. In the nine specimens in the museum, seven ones are found from Hobetsu. The first study of the mosasaur fossil form Hobetsu area is Obata et. al. (1972). After opening of the museum (1982), Suzuki (1985a, b), Chitoku (1994), Sakurai et.al.(1999) reported some of them. I hope this report will be a basis of the further study for the mosasaurs in Japan and the related area. Key words- mosasaur, Hobetsu ab HMG-1075HMG-1076HMG Plotosaurus (?) sp. NSM-PV NSM-PV a HMG-10 3c HMG-11 3b HMG-12 3d 1985a 1984 HMG-12 Mosasaurus sp. indet HMG-10 Mosasaurinae gen. et sp. indet. HMG-11 Plioplatecarpinae? gen. et sp. indet. HMG-370 Mosasauridae gen. et sp. indet.hmg By The Hobetsu Museum 1
2 1985b HMG-12 Mosasaurus hobetsuensis Mosasaurinae Mosasaurus M.conodon M.conodon M.missouriensis HMG-10HMG-11HMG-12 HMG-370 HMG HMG-371 HMG-1 100m Santonian Campanian Campanian HMG HMG-371 Campanian 1994 HMG-371 Mosasaurus Plotosaurus Tylosaurus T.prorigerT.nepaeolicusT.haumuriensis Tylosaurus sp HMG-1065 Mosasaurus prismaticus Mosasaurinae Mosasaurus 2004 HMG-1076HMG-1077 HMG HMG-12HMG-1065 HMG-371HMG-11HMG-10HMG HMG-1075 HMG-1076HMG-1077 Caldwell and Konishi (2007) Plotosaurus (?) sp. NSM-PV15003 haemal arch Plotosaurus Russellosaurina Russellosaurina 1985 HMG HMG-371 HMG-371 Tylosaurus NSM-PV15003 HMG-1075HMG-1076HMG-1077 HMG-1075 HMG-1075 Mosasauridae, gen. et sp. indet. caudal vertebra intermediate region 4 ConiasianMaastrichtian The Bulletin of the Hobetsu Museum, no. 23, March 2008
3 Mosasauridae intermediate region Russell1967 Plioplatecarpinae Tylosaurinae HMG-1076 HMG-1076 Mosasauridae gen. et sp. indet. caudal vertebra pygal4 CampanianMaastrichtian Mosasauridae pygal 30 Russell1967 ClidastesPlotosaurusPlatecarpusTylosaurusM.conodon M.conodonPlatecarpus Ectenosaurus Mosasauridae gen. et sp. indet. HMG-1077 HMG-1077 Mosasaurus sp. cf. M. hobetsuensis pterygoid tooth 5(?) prefrontal (?)? 1 splenial1 dentary1 ulnare1 Campanian?Maastrichtian? Mosasauridae?? supraorbital process 6 7 V U angular mandibular channel Russell1967 1/3 8 HMG-12 intermedium The Bulletin of the Hobetsu Museum, no. 23, March
4 HMG-371Tylosaurus? sp.m Mosasaurus hobetsuensis HMG-12 M.hobetsuensis HMG-12 Mosasaurus sp. cf. M. hobetsuensis HMG-10 HMG-10 Mosasaurinae gen. et sp. indet. intermediate region Maastrichitian a a Mosasauridae Russell, 1967 Russell, Russell1967 Mosasaurinae ClidastesPlotosaurusPlatecarpus TylosaurusM.conodon 1985a Mosasaurinae HMG-11 HMG-11 Plioplatecarapinae? gen. et sp. indet. premaxilla 1 maxilla 2 marginal tooth1 angular1 atlas 1 cervical vertebra1 Coniasian Santonian1985a a a narial opening internarial bar atlas centrum atlas intercentrum atlas neural arch atlas synapophysis 4 The Bulletin of the Hobetsu Museum, no. 23, March 2008
5 hypapophysis Mosasauridae Russell1967, p.74 Platecarpus hypapophysis hypapophysis zygospene zygontrum Russell (1967) ClidastesMosasaurusEctenosaurus Clidastes EctenosaurusMosasaurus GlobidensPlatecarpus EctenosaurusPlioplatecarpus zygospene zygontru Clidastes Ectenosaurus 1985a Plioplatecarpina? gen. et sp. indet. Mosasaurinae Dr. Caldwell Platecarpus Platecarpus 1985a Plioplatecarpinae? gen. et sp. indet. HMG-370 HMG-370 Mosasaurinae gen. et sp. indet. cervical vertebra 1 Santonian 1985a a a hypapophysis Mosasauridae 1/2 Russell, 1967 zygosphene zygantrum (3) Russell (1967) ClidastesMosasaurusTylosaurus 1985a Mosasauridae gen. et sp. indet Mosasaurinae Caldwell, Mosasaurinae gen. et sp. indet. HMG-371 HMG-371 Tylosaurs sp. parietal 1 jugal1 pterygoid pterygoid teeth 3 marginal teeth 6anterior mandibular unit splenialdentary posterior mandibular unitangular surangularprearticular articular basisphenoid basioccipitaldorsal vertebra? 1 disit 1 Campanian parietal foramen overlap ectpterygoidal process The Bulletin of the Hobetsu Museum, no. 23, March
6 alar process ectpterygoidal basipterygoid process process basipterygoid process parasphenoid rostrum alar process occipital condyle basioccipital tuber basioccipital tuber basisphenoid neural arch hypapophysis perpendicular keel 1994 Tylosaurus mandibular channel prearticular anterior blade Tylosaurus sp Dr. Caldwell Tylosaurus coronoid Caldwell and Konishi, 2007Caldwell Russell1967; p.177, Text-fig.95 Williston1898; Pl.XVI Tylosarus Tylosaurs sp. HMG-1065 HMG-1065 meckelion fossa Mosasaurus prismaticus Sakurai, Chitoku and glenoid fossa Shibuya, 1999 marginal teeth1 parietal anterior blade 1 basioccipital basisphenoid parasphenoid - ophisthotic-exoccipital prootic supratemporal(?) epipterygoid?1 glenoid fossa rectangular process CampanianMaastrichtian meckelion fossa 6 The Bulletin of the Hobetsu Museum, no. 23, March 2008
7 (2) 1999 parietal foramen table occipital unit spraoccipital (?) Mosasaurus M. hobetsuensis 1999 Mosasaurus HMG-12 HMG-12 Mosasaurus hobetsuensis Suzuki, 1985 marginal tooth1 dorsal vertebra 10 dorsal rib coracoid1 scapula1 forelimb 1 interclavicle 1(?) clavicle? 1 carpus 1 metacarpals1 phalanges 6 Maastrichtian ab b 1985b M.conodon M.conodon M.missouriensis M.conodon M.missouriensis Lingham-Soliar1995HMG-12 Leiodon Dr. Caldwell Caldwell HMG-1077 Mosasaurus HMG-1075HMG-1076 HMG-1077 Mosasaurus Mosasaurinae University of Alberta Dr. Caldwell 2006 Dr. Caldwell Sternberg Museum Dr. Everhart 2002 HMG-1065 HMG-12 The Bulletin of the Hobetsu Museum, no. 23, March
8 Caldwell, M.W. and Konishi, T, 2007, Taxonomic re-assignment of the first-known mosasaur specimen from Japan, and a discussion of circum-pacific mosasaur paleobiogeography. Journal of Vertebrate Paleontology 27(2): p p p.29-36pl p.9-14pl p.1-7pl.3 Lingham-Soliar, T, 1995, The mosasaur Leiodon bares its teeth. in Sarjeant, W.A.S. ed. Vertebrate Fossils and the Evolution of Scientific Concepts. p p p Russell, D.A., 1967, Systematics and morphology of American mosasaurs. Bulletin of the Peabody Museum of Natural History 23: p Mosasaurus 15p P p a 2p pl b Mosasaurus 30p Williston, S.W., 1898, Mosasaurs. University of Kansas Geological Survey, 4, p , pls Kazuhiko SAKURAI, A review of the mosasaur fossils from Hobetsu and around area. The Bulletin of the Hobetsu Museum, 23, HMG-1075HMG-1076HMG ab The Bulletin of the Hobetsu Museum, no. 23, March 2008
9 穂 別 とその 周 辺 地 域 のモササウルス 化 石 の 再 検 討 b a a b 図 1 HMG-1075 産 出 部 位 断 片 的 な 尾 椎 ( 中 位 部 )の 椎 体 3 点 と, 分 離 したV 字 骨 の 遠 位 部 からなる. (a,b:hmg-1075を 異 なった 方 向 から 見 る) Fig.1 Preserved portion of HMG-1075 Three fragmental caudal vertebra (intermedial region) and an isolated chevron (distal portion). (a, b: different view of HMG-1075) a a b 図 2 HMG-1076 産 出 部 位 断 片 的 な 尾 椎 ( 基 部 )の 椎 体 3 点 と, 断 片 的 な 横 突 起 からなる. (a,b:hmg-1075を 異 なった 方 向 から 見 る) Fig.2 Preserved portion of HMG-1076 Three fragmental caudal vertebra (pygal) and a fragment of transverce process. (a, b: different view of HMG-1076) The Bulletin of the Hobetsu Museum, no.23, March b
10 櫻 井 和 彦 図 3 HMG-1077 産 出 部 位 ( 翼 状 骨 歯 ) 分 離 した 翼 状 骨 歯 の 歯 冠 部 4 点. Fig.3 Preserved portion of HMG-1077 (pterygoid tooth) Four isolated pterygoid teeth (crown). 図 4 HMG-1077 産 出 部 位 ( 翼 状 骨 歯 ) 4 点 の 翼 状 骨 歯 のうち, 最 も 保 存 状 態 の 良 い 歯. Fig.4 Preserved portion of HMG-1077 (pterygoid tooth) One of the pterygoid teeth (the best condition). 図 5 HMG-1077 産 出 部 位 ( 翼 状 骨 歯 ) 単 独 の 翼 状 骨 歯.ほぼ 完 全 な 歯 根 と 輪 郭 状 の 歯 からなる. Fig.5 Preserved portion of HMG-1077 (pterygoid tooth) An isolated pterygoid tooth (nearly complete root and outline of crown). p? a b 図 6 HMG-1077 産 出 部 位 ( 前 前 頭 骨?) 分 離 した 前 前 頭 骨?( 右?).a- 外 側 観,b- 内 側 観.p?: supraorbital process? Fig.6 Preserved portion of HMG-1077 (prefrontal?) Isolated prefrontal? (R?). a-lateral, b-medial. p?: supraorbital process? 10 The Bulletin of the Hobetsu Museum, no.23, March 2008
11 穂 別 とその 周 辺 地 域 のモササウルス 化 石 の 再 検 討 a b c 図 7 HMG-1077 産 出 部 位 ( 下 顎 前 位 部 ) 部 分 的 な 夾 板 骨 ( 左 )と 歯 骨 ( 左 ). a- 夾 板 骨,b- 歯 骨,c- 重 ね 合 わせた 状 態 ( 本 来 の 接 合 状 態 ). 全 て 外 側 観. Fig.7 Preserved portion of HMG-1077 (anterior mandibular unit) Partial splenial (L) and dentary (L). a-splenial, b-dentary, c-dentary on splenial. All phots are lateral view. a b c 図 8 HMG-1077 産 出 部 位 ( 尺 腕 骨 ) 一 部 を 欠 損 する 尺 腕 骨 ( 右 ).HMG-12(Mosasaurus hobetsuensis Suzuki, 1985 模 式 標 本 )のものに 類 似 する. a- 内 側 観,b- 外 側 観,c- 遠 位 観 ( 写 真 下 方 が 外 側 ) Fig.8 Preserved portion of HMG-1077 (ulnare) Partially lacked ulnare (R). Similar to one in HMG-12 (Holotype of Mosasaurus hobetsuensis Suzuki, 1985). a-medial, b-lateral, c-distal (lower is lateral) The Bulletin of the Hobetsu Museum, no.23, March
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