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1 ｨｬ : ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, ｨｨ05ｨ ｨｪ02ｨｪｨｺ ﾁ G R U M-G R Z H Y M A Y L O O.A., B I L A N E N K O E.N. MICROFUNGI AS A COMPONENT OF BOGS ECOSYSTEMS ｨｰｨｪ ｨｺ0609ｨｨ09ｨｰ02ｨｬ04 ｨｨ0008ｨ 06ｨｰ ｨ ｨｮ ｨｹ ｨ ｨ 05ｨｹｨｪ06ｨｬ ｨｺ08ｨｮ ｨｰ02 ｨｮ ｨ 09 ｨ ｨｨ ｨｪ06ｨｦ 09ｨｰ060806ｨｪ04, 06ｨｪｨｨ 06ｨ ﾂｨｨ09ｨ 06ｨｰ 09ｨｰ06ｨｺ ｨｮ ｨ ｨｨ04 ｨ ｨｰｨｬ , ｨｪｨ ｨｺｨ 0705ｨｨ09ｨ ｨｨ0102 ｨｰ ｨｪ ｨ ｨｦ, ｨｮ0006ｨｦ ｪ 0905ｨｮ03ｨ ｨｰ ｨｬ06ｨｴｨｪ04ｨｬ ｨｨ09ｨｰ06 ﾂｨｪｨｨｨｺ06ｨｬ 07ｨ 08ｨｪｨｨｨｺ ｨ ｪ ｨｬ02ｨｰｨ ｨｪｨ ｨｨ ｨｮ000502ｨｺｨｨ ｨ 04ｨ ｨｰｨｪ060302ｨｪｨｨ02 ｨｬ020301ｨｮ ｨｨｨｪｨｰ02ｨｪ09ｨｨ09ｨｪ0609ｨｰｨｹ ｨｰ06ｨｺ0609 ｨｮ ｨ ｨｰ ｨｮｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ06 ｨ ｨｰｨｪ ｨｺ0609ｨｨ09ｨｰ02ｨｬ, 09ｨｺ ｨｰｨｹ06 ｨｬｨｨｨｪ0208ｨ 05ｨｨ04ｨ 02ｨｨｨｨ 08ｨ 09ｨｰｨｨｨｰ0205ｨｹｨｪ ｨｰｨ ｨｰｨｺ0609, 0005ｨｮｨ ｨｨｨｪ06ｨｦ ｨｰ ｨｪ06ｨｦ 04ｨ ｨｨ, 00ｨｨ ｨｨ ﾂ0209ｨｺｨｨｨｬ ｨｨｨｬ06ｨｬ ｨ ｨｰ ｨｪ04ｨｦ ｨｨ ｨｬｨｪ ｨ ｨｺｨｰ0608ｨｪ04ｨｦ ｨｰ ｨ 08ｨ ｨ ｨｪｨｨ07 ｨｪ ｨｬ0603ｨｪ ｨｪ07ｨｰｨｹ ｨｨ 06ｨ ｨｲ0709ｨｪｨｨｨｰｨｹ, ｨｨ ｨｪ02 ｨｮ ﾂｨｨｨｰ0409ｨ ｨｰｨｹ ｨｰ0205ｨｹｨｪ0609ｨｰｨｹ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ ｨｰ08ｨｮｨｺｨｰ , ｨｺｨ ｨｺ ｨ ｨ ｨｺｨｰ0208ｨｨｨｦ, ｨｰｨ ｨｺ ｨｨ ｨｬｨｨｨｺ080609ｨｺ0607ｨｨ ﾂ0209ｨｺｨｨ ｨｨｨ ｨｮ ﾂ02ｨｪｨｨ ｨｨｨ ｨｰ ｨｪ ﾂ09ｨ ｨｴ02ｨｪ ｨ ｨ 06ｨｰ, 0601ｨｪｨ ｨｺ06 06ｨｰ09ｨｮｨｰ09ｨｰ09ｨｨ ｨｰｨｪ06ｨｦ ｨｺｨ 08ｨｰｨｨｨｪ04 ｨｬｨｨｨｺ0806ｨ ｨｨ ｨｨ ﾂ0209ｨｺ06ｨｦ ｨｰ08ｨｮｨｺ02ｨｨｨｨ 06ｨ ｨｮ ｨｪ06 ｨｪ ｨｰｨ ｨｰｨｺ06ｨｬ 01ｨ ｨｪｨｪ0401 ｨｺｨ ｨｺ 0706 ｨ ｨｨ0608ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨ ｨｨｨ ｨｪ ｨ ｨｴ0209ｨｰ ｨ 04ｨｪ0401 ｨｰｨｨ07ｨ 01 ｨｰ ｨｪ ﾂ09, ｨ 02ｨｪｨｪ ｨｪ ｨ ｨｦ06ｨｪ0609, ｨｰｨ ｨｺ ｨｨ ｨｮｨｪｨｺ02ｨｨ06ｨｪｨ 05ｨｹｨｪ06ｨｦ 04ｨｪｨ ﾂｨｨｨｬ0609ｨｰｨｨ 06ｨｰ010205ｨｹｨｪ ｨｨ ｨｨｨ 0609 ｨｪｨ 08ｨ 04ｨｪ ｨｰｨ 07ｨ 01 09ｨｮｨｺ ｨｨｨｨ ｨ ｨｪｨｨ ｨｨ00ｨｨｨｺｨｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ0609, ｨｬｨｨ08ｨｮ06ｨｴ ｨ 08ｨ ｨｺｨｰ0208ｨｪ ｨｰ ｨｪｨｨｨｺ0609 ｨｮ ｨｨ0701 ｨ ｨｪｨ ｨ ｨｨ0604ｨ, ｨｪｨｨ04ｨｺｨｨ01 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08, ｨｪｨｪ06ｨｦ ｨｺｨｨ090506ｨｰｨｪ0609ｨｰｨｨ ｨｨ 0905ｨ 03ｨｪ0609ｨｰｨｨ, ｨｪｨ 05ｨｨ ﾂｨｨ07 ｨ 0406ｨｰｨ 09 09ｨｨ ｨｪ ｨ ｨｪｨｨ ﾂ0209ｨｺｨｨ ｨｨｨｪ02ｨｪｨｨｨｦ ｨｨ ｨｨｨｪ00ｨｨｨ ｨｨ08ｨｮ06ｨｴ ｨｨ07ｨｪｨｨ ｨｪ0605ｨｹｨｪ ｨｨｨｪ02ｨｪｨｨｨｦ ｨｰｨ ｨｰ ｨｪ0206ｨ ｨｨｨｬ ｨｪ06 09 ｨｨ04ｨｮ ﾂ02ｨｪｨｨｨｨ ｨ ｨｰ ｨ 08ｨ ｨ ｨｪｨｨ ｨｬ. 04ｨ ｨｰ08ｨ ｨｪ02ｨｪｨｨ02 ｨｨ 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ09ｨｰｨｨｨｺｨ ｨ ｨｰ ｨｰｨ 04ｨ ｨｪｨｨｨｬｨ 06ｨｰ ｨｬｨｪ ｨｰ08ｨ ｨｪ09ｨｰ09ｨ ｨｪｨ 0402ｨｬｨｪ06ｨｬ 03ｨ ｨ ｨｴｨ ｨｴｨ 01ｨｹ ｨｨ09- ﾂｨｨ ｨｰ ｨｨｨ 05ｨｨ04ｨｨｨｰ0205ｨｹｨｪ ｨｬ05ｨｪ. 00ｨ (08ｨ 02, 1941) ｨｴｨ 01ｨｹ ｨｰ ｨｪ0401 ｨ ｨｰ ｨｨｨｨ ｨｰｨ ｨｰ 0708ｨｨｨｬ0208ｨｪ06 2/5 06ｨｰ ｨｬｨｨ ｨｦ (06ｨｪｨｨ030209ｨ, 2005). 05ｨｮｨｴ0209ｨｰ09ｨｮ02ｨｰ ｨｪ0209ｨｺ0605ｨｹｨｺ ｨｪｨｨｨｦ 0706ｨｪ07ｨｰｨｨ07 00ｨ ｨｰ0603, 01ｨ ｨｪｨｪ ｨ 04ｨｪ ｨｬ07 ｨｺ08ｨｮ07ｨｪ04ｨｬｨｨ ｨ ｨｰ ｨ ｨｬｨｨ. 0301ｨｨｨｪ09ｨｰ09ｨ 09 05ｨｰ06ｨｬ ｨｪ02ｨｰ. 0201ｨｪｨｨ ｨｨ ｨ ｨｰ0205ｨｨ ｨｪ02 09 ﾂｨｨｨｰｨ 06ｨｰ ｨｪｨ 05ｨｨ ﾂｨｨ02 ｨｰ060800ｨ 06ｨ 0704ｨ ｨｰ0205ｨｹｨｪ04ｨｬ 0708ｨｨ04ｨｪｨ ｨｺ06ｨｬ ｨ ｨｰｨ (05ｨｮｨｺｨ ﾂ0209, 1926; 08ｨ 02, 1941; 01ｨｨ0202ｨｪｨｺ06, 1967; ｨｨｨｪｨ ｨｨ 0108., 2003), 0108ｨｮ00ｨｨ02 ｨｪｨ ｨ 06ｨｰ ｨ ｨｰ06ｨｬ ｨｰ0605ｨｹｨｺ06 ｨｰｨ ｨｺ06ｨｦ 05ｨ ｨｪ0103ｨ 00ｨｰ, ｨｰ ｨｨｨ 06 ｨｮ0302 ｨｨｨｬ0202ｨｰ0907, 05ｨｨｨ ｨｬ02ｨｪｨｹ0302ｨｦ ｨｬ ｨｪｨ ﾂｨ 05 06ｨｰ05ｨ 00ｨ ｨｰｨｹ0907 ( ｨｬｨｪ0609, 1949) ｨｪｨｨ06, ｨｬｨｮ05ｨｨ080609ｨ ｨｪｨｪ06ｨｬｨｮ ｨｪ04ｨｬ ｨ 06ｨｰｨ ｨｪｨｨ ﾂ0209ｨｺｨｨｨｬ 06ｨ ｨｴ0209ｨｰ0906ｨｬ , 00...ｨ ｨｰ06 ｪ ｨｰｨｨ ｨｬｨｪ06ｨｦ ｨｪ0609ｨｰｨｨ, ｨｰ0607ｨｪｨｪ06 ｨｨ05ｨｨ 0105ｨｨｨｰ0205ｨｹｨｪ ｨｬ07 ｨｮ0905ｨ 03ｨｪ02ｨｪｨｪ06ｨｦ ｨｨ 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ04ｨｮ06ｨｴ02ｨｦ ｨｰ0902ｨｰ09ｨｰ0902ｨｪｨｪ04ｨｬ 0706 ﾂ090606ｨ 08ｨ ｨ ｨｰ0205ｨｹｨｪ04ｨｬ ｨｬ

2 1306 ﾂ02ｨｪｨｹ ｨｬｨｪ0600ｨｨ02 ｨ 09ｨｰ ﾂｨｨｨｰｨ 06ｨｰ ｨｪｨ 05ｨｨ ﾂｨｨ02 ｨｰ060800ｨ 06ｨ 0704ｨ ｨｰ0205ｨｹｨｪ04ｨｬ 0708ｨｨ04ｨｪｨ ｨｺ06ｨｬ ｨ ｨｰ ｪ 05ｨｰ ｨ ｨｪｨｨ ﾂ0209ｨｺｨ ｨ, ｨ ｨｴｨ 07 ｨｪ02 ｨ % ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨｴ0209ｨｰ09 (06ｨｰ ｨ ｨ ｨｰｨｪ06 09ｨｮ ｨｴ0209ｨｰ09ｨ ｨｰ060800ｨ ), 06ｨ 08ｨ ｨ 0903ｨ ｨｮ05ｨｹｨｰｨ ｨｰ02 06ｨｰｨｬｨｨ08ｨ ｨｪｨｨ07 ｨｨ ｨｪ ｨｪ ｨ 0907ｨ 01ｨ ｨ ｨｰｨｪ ｨ 09ｨｰ02ｨｪｨｨｨｦ 09 ｨｮ ｨｨ ｨｪｨｪ06ｨｦ 0905ｨ 03ｨｪ0609ｨｰｨｨ 0708ｨｨ ｨｪ ｨｰｨ ｨｰｨｺ02 ｨｺｨｨ ｨ ( ｨｬｨｪ0609, 1976) ｨｰｨ 08ｨ ｨｰ0907 ｨｪｨ ｨｰ08ｨｨ ｨｰｨｨ07ｨ : ｨｪｨｨ04ｨｨｨｪｨｪ0402, ｨｪ0402 ｨｨ (ｨｨ05ｨｨ ｨ 09ｨｰ06ｨｰ080600ｨｪ0402, ｨｬ020406ｨｰ080600ｨｪ0402 ｨｨ 0605ｨｨ0006ｨｰ080600ｨｪ0402) ｨｪｨｨ04ｨｨｨｪｨｪ04ｨｬｨｨ 0706ｨｪｨｨｨｬｨ 06ｨｰ ｨ ｨｰｨ, ｨｪ0609ｨｰｨｹ ｨｺ06ｨｰ ｨｮ0905ｨ 03ｨｪ0706ｨｰ , ｨ 0600ｨ ｨｰ0402 ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ04ｨｬｨｨ ｨｬｨｨ ｨｨ 07ｨｨｨｰｨ 06ｨｴｨｨ ｨｮｨｪｨｰ060904ｨｬｨｨ ｨ ｨｬｨｨ, ｨ ｨｬｨｨ ｪ ｨｮ0905ｨ 03ｨｪ0702ｨｬ ｨ ｨｬｨｨ, ｨ 0201ｨｪ04ｨｬｨｨ ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ04ｨｬｨｨ ｨｬｨｨ, ｨｰｨ ｨｬ04ｨｬｨｨ ｨ ｨｰｨｬ ｨｪ04ｨｬｨｨ 0609ｨ 01ｨｺｨ ｨｬｨｨ ｨｪ ｨｰｨ ｨｰ 0906ｨ 06ｨｦ ｨｬ0203ｨｮｨｰ06 ﾂｨｪｨｮ ｨｮ0707ｨｮ. 08 ｨｪｨｨ04ｨｨｨｪｨｪ04ｨｬ ｨｮ ｨｪ06 06ｨｰｨｪ060907ｨｰ0907 ｨ ｨｰｨ ｨｹｨｪ0609ｨｰｨｹ ｨｪ ｪ7 %, ｨｺ ｨｪ04ｨｬ ｪ ｨｹｨｪ0609ｨｰｨｹ06 06ｨｰ ｪ7 %, ｨ ｨｺ ｨｬ ｪ ｨｹｨｪ0609ｨｰｨｹ06 ｨｬ02ｨｪ %. 01ｨ ｨ ｨｰｨ 01 ｨｺｨｨ090506ｨｰｨｪ0609ｨｰｨｹ 07ｨｨｨｰｨ 06ｨｴｨｨ ｨｪ ｨ 09ｨｰ ｨｺｨ 07, ｨ ｨｪｨ ｨｪｨｨ04ｨｨｨｪｨｪ0401 ｪ ｨｪｨｨ04ｨｺｨ ｨ ｨｺ ｨｺｨ ｨｺ ｨｰｨｨ0704 ｨ ｨｰ 08ｨ 0405ｨｨ ﾂｨ 06ｨｰ ｨ 0600ｨ ｨｰ09ｨｰ09ｨｮ 07ｨｨｨｰｨ ｨｪｨｨ07, ｨｰ06 ｨｨ ｨｰｨ 09 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨｹ ｨｪ020601ｨｨｨｪｨ ｨｺ ｨｨｨｪ09ｨｰ0902ｨｪｨｪ ｨｰ0906 ｨ ｨｰｨｪ06ｨｦ ｨｰ06ｨｬ, ﾂｨｰ ｨｰｨｨ 08ｨ 09ｨｰ02ｨｪｨｨ ｨ 04ｨｪ06ｨｦ 09ｨｰ020702ｨｪｨｨ 0905ｨ ｨ ｨｨ0904 (01ｨｨ0202ｨｪｨｺ06, 1967). 0308ｨｨ ﾂｨｨｨｪ04 ｨｨ 06ｨ ｨｴｨｨ02 ｨｬ0201ｨ ｨｪｨｨ04ｨｬ04 06ｨ 08ｨ ｨ ｨｪｨｨ07 ｨｨ ｨｪｨ ｨｺ ｨｪｨｨ07 ｨｰ060800ｨ, ｨ ｨｰｨ ｨｺ0302 ｨｬ0209ｨｰ ｨｺｨ 05ｨｨ04ｨ 02ｨｨｨｨ ｨ ｨｰ ｨ 08ｨ ｨ ｨｪｨｨ07 09 ｨｰ ｨｪ06ｨｦ 04ｨ ｨｨ ｨｮ ｨｰ090608ｨｨｨｰ0205ｨｹｨｪ06 06ｨ ｨｲ0709ｨｪ0706ｨｰ 00ｨｨ010806ｨｰ0208ｨｬｨｨ ﾂ0209ｨｺｨ 07 00ｨｨ0706ｨｰ0204ｨ ｨｨ ｨｰ020608ｨｨ07 ｨｰ ｨｪｨｪ , 0601ｨｪｨ ｨｺ06 06ｨｪｨｨ ｨｪ02 06ｨ ｨｲ0709ｨｪ0706ｨｰ, 0706 ﾂ02ｨｬｨｮ ｨｰ ｨｨｨｬ0202ｨｰ 08ｨ 04ｨｪ ｨｨ ﾂｨｨｨｪ04 09ｨｰ020702ｨｪｨｨ 08ｨ ｨｪｨｨ07 ｨｨ 0706 ﾂ02ｨｬｨｮ 09ｨｰ020702ｨｪｨｹ 08ｨ ｨｪｨｨ ｨｨｨｮ0806 ﾂ02ｨｪｨ ｨｺ ｨ 06ｨｰｨ ｨｪｨｨ ﾂ0209ｨｺ06ｨｬｨｮ ｨｰｨ 09ｨｮ ｨｰ060800ｨ (09ｨ ｨｬｨ ｨ , 2005) ｨｰ ｨ 08ｨ ｨ ｨｪｨｨ07 ｨｪ02 06ｨ ｨｲ0709ｨｪ0706ｨｰ0907, ｨｨ ｨｨ00ｨｪ0608ｨｨ080609ｨ ｨｰｨｹ ｨｰ0205ｨｹｨｪ0609ｨｰｨｹ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ0609, ｨｺｨ ｨｺ060904ｨｬｨｨ ｨｰ0907 ｨ ｨ ｨｺｨｰ0208ｨｨｨｨ ｨｨ ｨｬｨｨｨｺ080609ｨｺ0607ｨｨ ﾂ0209ｨｺｨｨ ｨｨｨ ｨ ｨｰｨ ｨ ｨｰｨ ｨｴｨ 01ｨｨ ｨｨ 04ｨ 07ｨ 09ｨ ｨｬ ｨｰ060800ｨ 04ｨ ｨｪｨｨｨｬｨ 06ｨｰ ｨｨｨｨ ｨｬ0209ｨｰ ｨｨ 0108ｨｮ00ｨｨ01 ｨ ｨｰ. 02ｨｪｨｨ ｨｪ0609ｨｪ06ｨｬ 08ｨ ｨｰ08ｨ ｨｪ02ｨｪ04 09 ｨｰｨ ｨｦ0002, ｨｪ ｨｰ0802 ﾂｨ 06ｨｰ0907 ｨｨ 0603ｨｪ0202, ｨｰｨｹ ｨｪ04 09ｨｰ020702ｨｦ ｨ ｨｰ06 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ04ｨｮ02ｨｰ0907 ｨ 0201ｨｪ04ｨｬ ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ04ｨｬ 07ｨｨｨｰｨ ｨｪｨｨ02ｨｬ, 06ｨｪ ｨｬｨｨ08ｨｮ02ｨｰ ｨｮ ｨｨ ｨ 09ｨｰｨ ｨｨ09ｨ ｨｪｨｨ ｨｪ0609ｨｰｨｪ ｨｪｨ ｨｺｨｨ ｨｪｨｨ0302ｨｪｨｨ ｨ , ｨｰｨｨ05ｨ 02ｨｬ ｨｪ ｨｪｨｨ02ｨ 02ｨｬ04ｨｬｨｨ ｨ ｨｬｨｨ. 02ｨ 04 ﾂｨｪ ｨ ｨｰ06 ｨｪ ｨ ｨｪ ｨｮｨｪｨｰ060904ｨｬｨｨ ｨ ｨｬｨｨ ｨｨ 09ｨｮｨｴ0209ｨｰ09ｨｮ02ｨｰ 04ｨ 09 ﾂ02ｨｰ ｨｰｨｮ070502ｨｪｨｨ ｨ 00ｨｨ ｨｨ04 ｨ ｨｰｨｬ ｨｪ ｨ 01ｨｺ ﾂｨ 09ｨｰｨｨ02 ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ ｨ ｨｪｨｨｨｨ 08ｨ 09ｨｰｨｨｨｰ0205ｨｹｨｪ ｨｰｨ ｨｰｨｺ0609 ｨｰ ｨｪｨｨｨｺ ｨｨ05ｨｹｨｪ06 04ｨ ｨｰ08ｨｮ01ｨｪ02ｨｪ06 ｨｺｨ ｨｺ ｨｨ04-04ｨ ｨｨ00ｨｨ ﾂ0209ｨｺｨｨ01 ｨｮ ｨｨｨｦ ｨ ｨｰｨｪ ｨｺ0609ｨｨ09ｨｰ02ｨｬ (ｨｪｨｨ04ｨｺｨｨ02 04ｨｪｨ ﾂ02ｨｪｨｨ , ｨ ｨｪｨ ｨ ｨｪ0402 ｨｮ ｨｨ07, ｨｪｨｨ04ｨｺｨｨ02 04ｨｪｨ ﾂ02ｨｪｨｨ07 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0804), ｨｰｨ ｨｺ ｨｨ ｨｨ04-04ｨ ｨ ｨｨｨｬｨｨ ﾂ0209ｨｺ ｨｰｨ 09ｨ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ-ｨｰ ｨ 08ｨ ｨ ｨｰ020502ｨｦ (ｨｮ , 05ｨｨ07ｨｨ0104, ｨ 0205ｨｺｨｨ, 05ｨｨ00ｨｪｨｨｨｪ 09 09ｨ ｨｬ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｪ ﾂ02ｨｰｨ ｨｪｨｨ0701, ｨ ｨｰｨ ｨｺ ｨｴ0209ｨｰ09ｨ -ｨ ｨｪｨｰｨｨ090207ｨｰｨｨｨｺｨｨ ｪ 0002ｨｪ060504, 0002ｨｪ0605ｨｺｨ 08ｨ 06ｨｪ ｨｺｨｨ090506ｨｰ04) ｨ 02ｨｪｨｪ06 ｨｮ09ｨｰ06ｨｦ ﾂｨｨ0904ｨｬｨｨ ｨｺ 08ｨ ｨｪｨｨ06 ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬｨ ｨｬｨｨ ｨｰ ｨ 00ｨｪ04 ｪ 0106ｨｬｨｨｨｪｨｨ08ｨｮ06ｨｴｨｨ02 ｨｬ01ｨｨ ｨ ｨｰ. 01ｨ ｨｪｨｨ0701, ｨｺ06 ﾂｨｺｨ 01 ｨｨ ｨ ｨｮ0008ｨ 01 ｨ 0204 ｨｨ04ｨ 04ｨｰ06 ﾂｨｪ ｨｮ0905ｨ 03ｨｪ02ｨｪｨｨ ｨ 05ｨ 01ｨ 06ｨｰ Sphagnum fuscum ( ﾂｨ ｨｴ ｨｪ ｨ ｨｦ06ｨｪｨ 01) ｨｨ S. magellanicum ( ﾂｨ ｨｴ ｨｪ02ｨｦ , 09 ｨｮ ｨｨ0701 ｨ ｨｰ ｨｨ ｨｺ06ｨｪｨｰｨｨｨｪ02ｨｪｨｰｨ 05ｨｹｨｪ ｨｺ05ｨｨｨｬｨ ｨｰｨ ) ｨ 07ｨ 01ｨｪ ｨ ｨｪｨｨ ｨｨｨｨ 06ｨ ｨｨ05ｨｹｨｪ06 08ｨ 0409ｨｨ09ｨ 02ｨｰ0907 S. rubellum. 0009ｨｰ0802 ﾂｨ 02ｨｰ0907 ｨｰｨ ｨｺ0302 S. angustifolium, ｨｪ06 ﾂｨ ｨｴ ｨｨ ｨｨｨｬ0209ｨｨ ｨｺ ｨｮｨｴｨｨｨｬ. 0500ｨ 00ｨｪ ｨｬ01ｨｨ ｨ 06ｨｰ ｨｺ08ｨ ｨｦｨｪ02 ｨｪ02ｨ 05ｨ ｨｨ07ｨｰｨｪ0402 ｨｮ ｨｨ ｨｨ04ｨｪｨｨ 0108ｨｮ00ｨｨ ｨ ｨｪｨｨ04ｨｬ0609, 09 ｨｰ06ｨｬ ﾂｨｨ ｨｨ ｨｮ0202ｨｪｨｰ ｨｰ06ｨｬｨｮ 08ｨ ｨｪｨｨ02 06ｨｰｨｬ020803ｨｨ ｨｰｨ ｨｰｨｺ0609 ｨｪｨ ｨｰｨ ｨｺ06ｨｬ ｨ ｨｰ ｨｨｨｰ ｨｺ08ｨ ｨｦｨｪ02 04ｨ ｨｬ ｨｪｨｪ06, ﾂｨｰ ｨ 09ｨｰ09ｨｮ02ｨｰ ｨｰ0607ｨｪｨｪ06ｨｬｨｮ 06ｨｰ ｨｪｨｨ06 ｨｰ060800ｨ ( ｨｪ02ｨｬ 1 ｨｬｨｬ ) ｨｨ ｨｰｨｮ ｨｰ ｨｪｨｨｨｺｨ ( , html?mid= &s). 0308ｨｨ09ｨｮｨｰ09ｨｰ09ｨｮ06ｨｴｨｨ ｨｨｨｬｨｨ ﾂ0209ｨｺ06ｨｬ ｨｰｨ ｨ 00ｨｪｨｮｨｬｨ 0002ｨｪ0605ｨｹｨｪ ｨｨｨｪ02ｨｪｨｨ ｨｰ0907 ｨｰ06ｨｺ09ｨｨ ﾂｨｪ04ｨｬｨｨ ｨ 0605ｨｹ03ｨｨｨｪ09ｨｰ09ｨ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ0609, ｨｰ0605ｨｹｨｺ ｨｨｨ ｨｰｨｬ02 ﾂ02ｨｪｨ ｨ ｨｪ0609ｨｰｨｹ ｨｨ ﾂｨｪ06ｨｦ ｨｰ08ｨｮｨｺ02ｨｨｨｨ ｨｬ ｨ ｨｺｨｰ0208ｨｨｨｨ ｨｨ ｨｨ ｨｰ0907 ｨｪｨ ｨｮｨｺｨｰｨ 01 08ｨ 0408ｨｮ0302ｨｪｨｨ07 ｨｬ ｨｨｨ ｨ ｨｬｨｨ ｨｨ, ｨｨ ｨｹ04ｨｮ07 05ｨｰｨｨ ｨｮｨｺｨｰ04, ｨ 06ｨｰ ｨｨ01 01ｨ 05ｨｹｨｪ02ｨｦ0302ｨｬｨｮ 08ｨ ｨｪｨｨ06 (0206ｨ ｨｹ09ｨｺｨ 07 ｨｨ 0108., 2005). 486

3 ｨ ｨｪｨｨ07 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨ ｨｰ ｨ ｨｰｨ ｨ 02ｨｪｨｪ06 03ｨｨ0806ｨｺ06 08ｨ ｨｰ08ｨ ｨｪ02ｨｪ ｨｰｨ 0203ｨｪ06ｨｦ ｨｨ ｨｰｨｮｨｪ ｨｦ 0406ｨｪｨ ｨｹ 06ｨｪｨｨ 04ｨ ｨｪｨｨｨｬｨ 06ｨｰ ｪ80 % 0402ｨｬｨｪ06ｨｦ ｨｪ0609ｨｰｨｨ ｨｬ06ｨｰ0807 ｨｪｨ ｨ ｨｺｨｰｨｨ09ｨｪ0602 ｨｮ ﾂｨ 09ｨｰｨｨ02 ｨ ｨｰ ｨ ｨ 05ｨｹｨｪ06ｨｬ ｨｺ08ｨｮ ｨｰ02 ｨｮ ｨ, ｨｪｨ ｨｪ0703ｨｪｨｨｨｦ 0102ｨｪｨｹ 06ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ06 ｨｬｨ ｨ ｨｪｨｪ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨｨｨ 0008ｨｨｨ 0609 ｨｨ ｨｨ ｨｨ 09 ｨ ｨｰｨｪ ｨｺ0609ｨｨ09ｨｰ02ｨｬｨ 01, ｨ ｨｰｨ ｨｺ ｨ ｨｨ01 ｨｮ ﾂｨ 09ｨｰｨｨｨｨ ｨｰ ｨ 08ｨ ｨ ｨｪｨｨ07 (Thormann, 2006). 06ｨｬ0206ｨｴｨｨ ｨｨｨｰ0208ｨ ｨｰｨｮ ｨｪｨｨ ｨｰ 06ｨ 0604ｨｪｨ ﾂｨｨｨｰｨｹ ｨｪ0209ｨｺ0605ｨｹｨｺ06 ｨｪｨ 0708ｨ ｨｪｨｨｨｦ 09 ｨｨ ｨ ｨｪｨｨｨｨ 0008ｨｨｨ 0609 ｨｰ ｨｪ ﾂ09. 08ｨ ｨｺ ｨｮ 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ0401, ｨｰｨ ｨｺ ｨｨ ｨｮ 04ｨ 08ｨｮｨ 0203ｨｪ0401 ｨｨ ｨ ｨｰ020502ｨｦ 0609ｨｪ0609ｨｪ04ｨｬｨｨ ｨｪｨ 0708ｨ ｨｪｨｨ07ｨｬｨｨ 09 ｨｨ04ｨｮ ﾂ02ｨｪｨｨｨｨ 0008ｨｨｨ ｨｪ ｨ ｨｴ0209ｨｰ09 ｨｰ ｨｪｨｨｨｺ ｨ ｨｰ ｨｰ0907: ｨｪｨｨ02 09ｨｨ ｨｰｨ 09ｨ 0008ｨｨｨ 0609, ｨｨ01 ｨｺ0605ｨｨ ﾂ0209ｨｰ0902ｨｪｨｪ ｨｰｨｪ060302ｨｪｨｨ07, ｨｰ08ｨ ｨｪ09ｨｰ0902ｨｪｨｪ ｨｨ 0005ｨｮｨ ｨｨｨｪｨｪ ｨ ｨｪｨｨ07, ｨｬ ｨｨ ﾂ0209ｨｺ06ｨｦ 09ｨｰ08ｨｮｨｺｨｰｨｮ0804 (ｨｰ ｨｰｨｪ060302ｨｪｨｨ ｨｨ ｨｬｨｨ020205ｨｨ07), ｨｪｨｪ0401 ｨｺ060502ｨ ｨ ｨｪｨｨｨｦ ｨｰｨ ｨｨ ｨｨ ｨｨ01 ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ, 09ｨｮｨｺ ｨｨｨｨ 09ｨｨ010609, ｨｨ ｨ ｨｪ0609ｨｰｨｨ ｨｺ 08ｨ ｨｪｨｨ06 09ｨｮｨ 09ｨｰ08ｨ ｨｰｨ ｨｨ ｨ 09ｨｨ09ｨｨｨｬ0609ｨｰｨｨ 06ｨｰ 09ｨｪ0203ｨｪｨｨ01 00ｨ ｨｺｨｰ (ｨｰｨ ｨｺｨｨ01 ｨｺｨ ｨｺ 0905ｨ 03ｨｪ0609ｨｰｨｹ ｨｨ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨ ), 0904ｨ ｨｨｨｬ0606ｨｰｨｪ060302ｨｪｨｨｨｦ 0008ｨｨｨ ｨ 09ｨｰ02ｨｪｨｨ07ｨｬｨｨ ｨ ｨｬｨｨ (ｨｬ01ｨ ｨｬｨｨ) ｨｰ ｨ ｨ 06ｨｰ ｨｨ ｨ ｨｪｨｨ ｨ ｨｴ0209ｨｰ09 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ0609 ｨｰ ｨｪｨｨｨｺ ｨ 00ｨｪ ｨｨ ｨｪｨｨ04ｨｨｨｪｨｪ0401 ｨ ｨｰ, 04ｨ ｨ ﾂ02ｨｪｨｪ0401 ｨｨ ｨｰ ｨｪｨｨ09ｨｰ0401 ｨｬ0209ｨｰｨｪ0609ｨｰ02ｨｦ , 0804ｨｨｨｨ, ｨｪ06ｨｦ ｨｨ 0703ｨｪ06ｨｦ 08ｨｬ0208ｨｨｨｺ ｨｬ ｨｨ01ｨ 01 ｨｰ ｨｪｨｨｨｺ ｨｬｨｨｨｪｨｨ08ｨｮ06ｨｰ ｨ ｨｪｨ ｨｬ ｨ 09ｨｺ06ｨｬｨｨ0202ｨｰ0609 (ｨ % 06ｨｰ ｨｨ ｨｨｨ 0609). 08ｨ ｨｪｨ 0109ｨｺｨｨｨｦ ｨｨ ｨ ｨｰ0205ｨｹ ｨｬｨ ｨｪ (Thormann, 2006) ｨｦ 08ｨ ｨ 06ｨｰ ｨｹ 0008ｨｨｨ ｨ ｨｰｨ 01 ｨ ｨ 05ｨｹｨｪ06ｨｦ 0406ｨｪ ｨｨ090601ｨｨｨｰ 01ｨ ｨｪｨｪ ｨｰ06ｨｬ, ﾂｨｰ06 ｨｬ0603ｨｪ ｨｨｨｰｨｹ 0707ｨｰｨｹ 0008ｨｮ ｨｨｨ 0609, 0609ｨｮｨｴ0209ｨｰ ｨｴｨｨ01 08ｨ ｨｪｨｨ02 08ｨ 09ｨｰｨｨｨｰ0205ｨｹｨｪ ｨｰｨ ｨｰｨｺ ｨｰ ｨｪｨｨｨｺｨ ｨｦ 0008ｨｮ ｨｪ 06ｨｰｨｪ0609ｨｨｨｰ 07ｨ ｨｰ060002ｨｪ0609 ｨｨ 0905ｨ ｨ ｨ 08ｨ 04ｨｨｨｰ ｨ 09ｨｰ02ｨｪｨｨｨｦ. 06ｨｰ06 09ｨｨ Cladosporium, Aureobasidium, Botrytis ｨｨ Alternaria. 05ｨ ｨｰ02ｨｬ, ｨｰｨ 0802ｨｪｨｨ07 ｨｨ 06ｨｰｨｬｨｨ08ｨ ｨｪｨｨ07 08ｨ 09ｨｰ02ｨｪｨｨ07 ｨｪｨ ｨ ｨｪｨｨ ﾂ0209ｨｺ06ｨｬ ｨｬｨ ｨｰ0208ｨｨｨ ｨ ｨｰ0205ｨｹｨｪ ｨｰ ｨｨ06ｨｪ0208ｨｪ ｨ ｨｰ (09ｨｰ0608ｨ ｨｮ0707ｨ ｪ ｨｪ0609ｨｪ06ｨｬ 05ｨｰ06 04ｨｨ0006ｨｬｨｨ0202ｨｰ04, 09ｨｨ Mucor, Umbelopsis ｨｨ Mortierella), ｨｺ06ｨｰ ｨｬ02ｨｪ0706ｨｰ ｨｨｨ ｨ ｨｬｨｨ, 08ｨ 0405ｨ 00ｨ 06ｨｴｨｨｨｬｨｨ ｨｰ ｨｨｨｬ (ｨｰ0802ｨｰｨｹ ｨｮ0707ｨ ｪ 09ｨｨ Fusarium, Chaetomium, Trichoderma ｨｨ Penicillium) ｨｺ06ｨｪ ﾂｨ ｨｰ0205ｨｹｨｪｨｮ ｨ 01ｨ 02ｨｨ ｨ 06ｨｴｨｨ ｨ ｨｪｨｨ ｨｨｨｬ ｨｮｨｴ0209ｨｰ ｨｰ 0008ｨｨｨ 04 ﾂ02ｨｰ090208ｨｰ06ｨｦ 0008ｨｮ070704, ｨｺ ｨｺ06ｨｰ060804ｨｬ 06ｨｰｨｪ060907ｨｰ0907 ｨ ｨ 04ｨｨ01ｨｨ06ｨｬｨｨ0202ｨｰ04 ｨｨ ｨｪ02ｨｺ06ｨｰ ｨ 09ｨｺ06ｨｬｨｨ0202ｨｰ ｨｪ ｨｮ0707ｨ, 09ｨｺ0506 ﾂｨ 06ｨｴｨ 07 00ｨｨｨ ｨｺｨｨ02 09ｨ ｨｰ , ｨｰｨ ｨｺｨｨ02 ｨｺｨ ｨｺ 09ｨｨ Pythium, Mortierella ｨｨ ｨｪ02ｨｺ06ｨｰ ｨｮ00ｨｨ02, ｨｪ0609ｨｰｨｹ06 04ｨ ｨ 02ｨｰ ｨ ｨｪｨｨ07. 06ｨ ｨｺｨｨｨｬ 06ｨ 08ｨ 0406ｨｬ, 06ｨｰ ｨｪｨ - ﾂｨ 05ｨ 09ｨｰｨ 0802ｨｪｨｨ ｨｪ ｨ 0907ｨ 01ｨ 08ｨ 09ｨｰｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｨ 0008ｨｮ ｨ ｨｰ ｨｨ00ｨｨ ﾂ0209ｨｺｨｨｨｬｨｨ ｨｺ 08ｨ 0405ｨｨ ﾂｨｪ04ｨｬ ｨｮ ｨｪ04ｨｬ ｨｨｨｪ02ｨｪｨｨ07ｨｬ ｨｪｨ ｨ 0608ｨ ｨｬｨｨ ｨｬ02ｨｪｨｰ ｨｰ0907 ｨｪｨ ｨ ｨｪｨｨ ﾂ0209ｨｺ06ｨｬ 0902ｨｴ0209ｨｰ0902 ｨｨ, 09ｨｬ02ｨｪ ｨｮ ｨｮ00ｨ, 06ｨ 08ｨ 04ｨｮ06ｨｰ 09ｨｮｨｺ ｨｨ06. 02ｨ ｨｪｨｪ ｨｨ ｨｬｨｮ ｨｰｨ 09ｨｮ ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨ ｨｰ ｨｮ 08ｨ 04ｨｪ0401 ｨ 09ｨｰ ｨｪ04. 02ｨｰｨｬ02 ﾂ02ｨｪ ｨｺ ｨｨ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨ ｨ Penicillium, ｨｨ04 ｨｺ06ｨｰ ｨｬｨ ｨｬｨｨ ｨｺｨ ｨｺ 0706 ﾂｨ 09ｨｰ06ｨｰ ｨｰ0802 ﾂｨ 02ｨｬ0609ｨｰｨｨ, ｨｰｨ ｨｺ ｨｨ ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ06ｨｬｨｮ 06ｨ ｨｨ05ｨｨ ｨｰ ｨｨ0104 P. funiculosum, P. lividum, P. thomii, P. spinulosum, ｨｬ ｨｦ 0005ｨｮｨ ｨｨｨｪ02 ｨｰ ｨｪ06ｨｦ 04ｨ ｨｨ ｨｺｨｨ ｨｪｨ ｨ ｨｪｨｨｨｦ ｨｨ00ｨｨ ﾂ0209ｨｺｨｨｨｬ 09ｨｨ01ｨ ｨｬ 01ｨ ｨｪｨｪ0401 ｨ ｨｨ ｨｪ ｨｮ ﾂ02ｨｪ ｨｰｨｪ060907ｨｰ 09ｨｨ Oidiodendron (O. maius, O. griseum, O. tenuissimum, O. scytaloides). 0108ｨｨｨ 04 05ｨｰ ｨ ｨ ｨｪ04 ｨｺ 08ｨ ｨｪｨｨ ｨ 00ｨｪｨｮｨｬｨ ｨｨ 06ｨｰｨｬ02 ﾂ02ｨｪ04 ｨｺｨ ｨｺ ｨｬｨｨｨｺ0608ｨｨ040606ｨ 08ｨ ｨ ｨｰ0205ｨｨ ｨｪｨ ｨｺ ｨ 09ｨｰ02ｨｪｨｨ0701 (Thormann, 2006) ｨｬ02 ｨｰ060006, ｨｨ ｨ ｨｰ 09 ｨ 0605ｨｹ03ｨｨ01 ｨｺ0605ｨｨ ﾂ0209ｨｰ09ｨ ｨｪ ｨｮ06ｨｴｨｨ02 09ｨｨ0104: Trichoderma koningii, T. viride, T. harzianum, Tilachlidium microsporum, Trichosporella cerebriformis, Mortierella minutissima, 00. horticola, M. renispora M. ramanniana, Mucor hiemalis, Aspergillus niger, A. versicolor, Verticillium psalliotae, Penicillium montanense, Pochonia bulbillosa, Alternaria alternata, Cladosporium spp., Mycelia sterilia (Latter et al., 1967; 0502ｨｬ02ｨｪ0609 ｨｨ 0108., 1995; 09ｨ ｨ ｨｨｨ ｨｮ0505ｨｨｨｪｨ, 2000; ﾂ02ｨｪｨｺ06 ｨｨ 0108., 2002; Thormann et al., 2004; 08ｨ ﾂｨ 05ｨｺｨｨｨｪ ｨｨ 0108., 2005; 0108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008) ｨ ｨ 06ｨｰ 0706ｨｺｨ 04ｨ ｨｪ06, ﾂｨｰ ｨｦ ｨｰ ｨｪ06ｨｦ 0706 ﾂ ｨｪ0609ｨｪ ｨ 07ｨ ｨｨｨ ｨｪ06ｨｦ ｨ ｨｨ06ｨｬｨ ｨｰ06 ﾂ02ｨｪ ﾂ ｨ 00ｨｪｨｮｨｬｨ. 05ｨ 07ｨ 0904 ｨｮｨ 0409ｨ 06ｨｰ 09 3 ｪ5 08ｨ 04 09ｨｪｨｨ ｨｨ0506, ｨｺｨ ｨｺ 09 03ｨｨ0906ｨｬ ｨｬ0102, ｨｰｨ ｨｺ ｨｨ 09 ｨｰ ｨｪ ｨｨ0406ｨｪｨｰｨ ｨｰｨｨｨｺｨ 05ｨｹｨｪ06ｨｦ 09ｨｰ08ｨｮｨｺｨｰｨｮ

4 13ｨｰ ｨｪ ﾂ09 ｨｨ04ｨｬ02ｨｪ0702ｨｰ0907 ｨｨ ｨ ｨｨ06ｨｬ ｨｨ ﾂ0209ｨｺｨ 07 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨ, ｨｰ ｨｰｨｪ060302ｨｪｨｨ ｨｨｨ ｨｪ ｨｬｨｨ020205ｨｨ07 ｨｨ ｨｨ020205ｨｨｨｦ 0106ｨｬｨｨｨｪｨｨ08ｨｮ02ｨｰ , ｨｺ0806ｨｬ02 ｨｪｨｨ03ｨｪ020006, 09 ｨｺ06ｨｰ060806ｨｬ ｨ 05ｨ 01ｨ 06ｨｰ ｨｨｨ 0609, ｨｨ ｨｰｨ /3 06ｨｰ 0008ｨｨｨ ｨｪ06ｨｦ ｨ ｨｨ06ｨｬｨ (08ｨ ﾂｨ 05ｨｺｨｨｨｪ ｨｨ 0108., 2005). 01ｨ 05ｨｨ ﾂｨｨ02 ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪ ｨｺ0605ｨｨ ﾂ0209ｨｰ09ｨ 0008ｨｨｨ ﾂ ｨ 00ｨｪｨｮｨｬｨ ｨｪｨ ｨｰ0605ｨｺｨｪｨｮ0506 ｨｨ ｨ ｨｰ020502ｨｦ ｨｪｨ ｨｨ04ｨｮ ﾂ02ｨｪｨｨ ｨ ｨｰ08ｨｮｨｺ02ｨｨｨｨ 03ｨｨ0906ｨｦ ﾂｨ 09ｨｰｨｨ 0900ｨ 00ｨｪｨｮｨｬ ｨｪ02ｨｺ06ｨｰ ｨｺ0502ｨｰｨｺｨ ｨ 00ｨｪｨｮｨｬｨ 06ｨ ｨｪｨ 08ｨｮ0302ｨｪ 07ｨ 08ｨ 04ｨｨｨｰｨｨ ﾂ0209ｨｺｨｨｨｦ 0008ｨｨｨ Lyophyllum palustre, ｨｺ06ｨｰ060804ｨｦ 06ｨｰｨｪ0609ｨｨｨｰ0907 ｨｺ ｨ ｨ 04ｨｨ01ｨｨ06ｨｬｨｨ0202ｨｰｨ ｨｬ (Tricholomataceae) (Untiedt, Muelter, 2001). 06ｨｰｨｨ 07ｨ 08ｨ 04ｨｨｨｰｨｨ ﾂ0209ｨｺｨｨ02 06ｨｰｨｪ060302ｨｪｨｨ07 ｨｨ ｨ 05ｨｨ09ｨｹ 09 ｨｬ0209ｨｰｨ 01 ｨｺ06ｨｪｨｰｨ ｨｺｨｰ ｨｨ00 09 ｨｺ0502ｨｰｨｺ06ｨｦ ｨｬ01ｨ. 03ｨ 08ｨ 04ｨｨｨｰ ｨｪｨｨｨｺｨ 02ｨｰ 09 03ｨｨ09ｨｮ06 ｨｺ0502ｨｰｨｺｨｮ ｨｨｨｪｨ, 08ｨ 09ｨｰ ｨｺ0502ｨｰ06 ﾂｨｪｨｮ06 09ｨｰ02ｨｪｨｺｨｮ ｨｪｨｨｨｺｨ 06ｨｴｨｨｨｬ ｨｺ06ｨｪ ﾂｨｨｨｺ06ｨｬ 00ｨｨ ｨｰｨ ｨｺ0609ｨ ｨｪｨｪｨ 07 ｨｺ0502ｨｰｨｺｨ, 0601ｨｪｨ ｨｺ06, ｨｪ ｨ 04ｨｮ 06ｨｰｨｬｨｨ08ｨ 02ｨｰ, ｨ ｨｨ ｨｨｨｰ 08ｨ ｨｪｨｨ02 ｨｺ0502ｨｰ06 ﾂｨｪ06ｨｦ 09ｨｰ02ｨｪｨｺｨｨ ｨｪｨｨｨｺｨｪ060902ｨｪｨｨ07 09 ｨｺ0502ｨｰｨｺｨｮ 07ｨ 08ｨ 04ｨｨｨｰ ｨｨ04ｨｬ02ｨｪ0702ｨｰ ｨｰ060705ｨ 04ｨｬｨｮ ｨ 0302ｨｪｨｪ0402 ｨｺ0502ｨｰｨｺｨｨ 09 ｨｨｨｰ060002, 0209ｨｰ0209ｨｰ0902ｨｪｨｪ06, 06ｨ ﾂｨｨ09ｨ 06ｨｰ ｨｰｨ ｨｪ0609ｨｨｨｰ0907 ｨ ｨｪ04ｨｬ ｨｨ 08ｨ 0408ｨｮ03ｨ 02ｨｰ ｨｰ060906ｨｬ ｨｬｨｨｨｺ080609ｨｺ ｨｨ01ｨｪ04 00ｨｨ0004, 06ｨｺｨ ｨｦｨｬ050706ｨｴｨｨ02 08ｨ 0907ｨ 0903ｨｨ ｨｺ0502ｨｰ06 ﾂｨｪ ｨｰ02ｨｪｨｺｨｨ ｨｪｨ 0008ｨ 00ｨｬ02ｨｪｨｰ04. 06ｨ ｨｺｨｨｨｬ 06ｨ 08ｨ 0406ｨｬ, ｨ 00ｨ 02ｨｰ ｨ 03ｨｪｨ ｨｹ ｨ ｨ 04ｨｨ01ｨｨ06ｨｬｨｨ0202ｨｰｨ L. palustre ｨｺｨ ｨｺ ｨｰ08ｨｮｨｺｨｰ0608ｨ 0900ｨ 00ｨｪ ｨｺ0502ｨｰ06ｨｺ, ｨｺ06ｨｰ , ｨｺｨ ｨｺ ｨｨ ｨｰｨｪ06, 09 ｨｰ08ｨｮ0106ｨｬ ｨ 06ｨｰ ｨｪｨｨ02ｨｪｨｨ06 ｨ 05ｨ ｨ ｨｬｨｮ 01ｨｨｨｬｨｨ ﾂ0209ｨｺ06ｨｬｨｮ ｨｰｨ 09ｨｮ ｨｨｨ ｨｪ ｨ ｨｪｨｨ07 ｨｺｨ ｨｺｨｨ01-05ｨｨｨ 06 09ｨｮｨ 09ｨｰ08ｨ ｨｰ ｨ 04 ﾂｨｪ06 ｨｮ ﾂｨ 09ｨｰ09ｨｮ06ｨｰ 08ｨ 04ｨｪ ｨｮ ｨｨｨ 0609, ｨｺ06ｨｰ ｨ ｨｰ0205ｨｹｨｪ06 09ｨｬ02ｨｪ0706ｨｰ 0108ｨｮ ｨｮ00ｨ (09ｨｮｨｺ ｨｨ07) ｨ ｨｪｨｨｨｨ 0900ｨ 00ｨｪ ﾂ0209ｨ ｨｨ05ｨｨ ﾂ02ｨｰｨｺｨｮ06 09ｨｮｨｺ ｨｨ06: ｨｪｨ 08ｨ ｨｪｨｪｨｨ01 ｨｨ ｨｪｨｨ01 09ｨｰｨ 01ｨｨ ｨｬｨｨｨｪｨｨ080609ｨ 05ｨｨ 09ｨｨ Mortierella ｨｨ Verticillium, ｨｪｨ ｨ ｨｪｨｨ01 ｨｨ01 09ｨｬ02ｨｪ0705ｨｨ Sporothrix sp. ｨｨ Trichoderma viride. 05ｨｬ02ｨｪｨ ｨ ｨｴ0209ｨｰ09 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ0609 ｨｺ ｨｨ080609ｨ 05ｨ 09 ｨｮ090205ｨｨ ﾂｨｨ09ｨ 06ｨｴ02ｨｦ0907 ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨ02ｨｦ ｨｮ ｨ ｨｨ ｨ ｨｨ ｨｪｨｨ02ｨｬ ｨｺｨｨ090506ｨｰｨｪ0609ｨｰｨｨ ｨｪｨ ｨｺ06 ｨｺｨ 08ｨｰｨｨｨｪｨ ｨｺ05ｨ 0909ｨｨ ﾂ0209ｨｺ ｨ 08ｨｨｨ ｨｪｨｰｨ 09ｨｮｨｺ ｨｨｨｨ ｨ 0405ｨ ｨｪｨ 08ｨｮ0302ｨｪｨ 0708ｨｨ09ｨｮｨｰ09ｨｰ09ｨｨ02ｨｬ 09 ｨｰ02 ﾂ02ｨｪｨｨ ｨｨ0601ｨ ｨｨ ｨ ｨｪｨｨ07 (0109ｨ ｨ ) 09ｨｨ010609, ｨ ｨｪ ｨ 0408ｨｮ03ｨ ｨｰｨｹ ｨｮ ｨｨ 05ｨｨ00ｨｪｨｨｨｪ (Oidiodendron maius, Acremonium cf. curvum ｨｨ Scleroconidioma sphagnicola) ｨ 00ｨｨ080609ｨ 05ｨｨ ｨｪｨ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 01ｨｨｨｬｨｨ04ｨｬｨ ｨｰｨｨ05ｨｺｨｨ Penicillium sp. ｨｨ Mucor sp., ｨｺ06ｨｰ ｨｨ09ｨｮｨｰ09ｨｰ090609ｨ 05ｨｨ ｨｪｨ ｨｰｨ 01ｨｨ ｨ ｨｪｨｨ ｨｬ0603ｨｪ06, ｨｰｨ ｨｺｨｨ ｨｮ05ｨｹｨｰｨ ｨｰ ｨｮ ﾂｨｨ05ｨｨ09ｨｹ ｨｨ04-04ｨ ｨｪ ｨｰｨ ｨｰ06 ﾂｨｪ ｨｨｨｰ0205ｨｹｨｪ ｨｨ0601ｨ ｨｨ ｨ ｨｪｨｨ07 (Thormann, 2003) ｨ ｨｪｨｨ ｨ ｨｪ0609ｨｰｨｨ 0008ｨｨｨ 0609, ｨｪｨｪ ｨｨ ｨｨ 06ｨｰｨｬｨｨ08ｨ 06ｨｴｨｨ01 ｨｰｨｺｨ ｨｪ02ｨｦ Sphagnum fuscum, 08ｨ 0405ｨ 00ｨ ｨｰｨｹ 08ｨ 04ｨｪ0402 ｨｮ ｨｪ ｨｮｨ 09ｨｰ08ｨ ｨｰ04 (ｨｴ0207ｨｺｨｨ ｨｨ ｨｦ ｨｨｨｪ04 ｨｨ 08ｨ 09ｨｰ02ｨｪｨｨ07 S. fuscum), 0706ｨｺｨ 04ｨ 05ｨｨ, ﾂｨｰ06 09ｨｨ0104 Sordaria fimicola, cf. Bjerkandera adusta, Acremonium cf. 09urvulum, Oidiodendron maius, Mucor hiemalis, Oidiodendron scytaloides, Penicillium thomii ｨｨ Trichoderma viride ｨ ｨｪ04 ｨｺ 08ｨ ｨｪｨｨ ; Sporormiella intermedia, Acremonium cf. 09urvum, Oidiodendron maius, Mucor hiemalis, Oidiodendron scytaloides, Penicillium thomii ｨｨ Trichoderma viride 08ｨ 0405ｨ 00ｨ 06ｨｰ ｨｺ08ｨ 01ｨｬｨ 05, ｨ ｨｺ 08ｨ 0408ｨｮ0302ｨｪｨｨ06 01ｨｮｨ ｨｨ05ｨｹｨｪ0401 ｨｺｨｨ090506ｨｰ ｨ ｨｪ04 ｨｰ0605ｨｹｨｺ ｨ 09ｨｨ01ｨ Oidiodendron: O. maius (09 ｨｪｨ ｨｨｨ 0605ｨｹ0302ｨｦ 09ｨｰ020702ｨｪｨｨ) ｨｨ O. scytaloides (0905ｨ ｨ 0202). 0305ｨｹ ｨ 0409ｨｰ ｨ 0408ｨｮ03ｨ 05ｨ 09ｨｹ ｨ ｨ 04ｨｨ01ｨｨ06ｨｬｨｨ0202ｨｰ06ｨｬ cf. Bjerkandera adusta, ｨ ｨｬ ｨｪｨｪ ｪ 02. scytaloides. 0500ｨ 00ｨｪｨｮｨｬ 08ｨ 0405ｨ 00ｨ ｨ ｨｮ0302, ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪ ｨ 0408ｨｮ0302ｨｪｨｨ ｨｨ ｨｨ050609ｨｹ ｨ 09ｨｺ06ｨｬｨｨ0202ｨｰ06ｨｬ Sordaria fimicola, ｨｬｨｨｨｪｨｨｨｬｨ 05ｨｹｨｪ0602 ｪ 04ｨｨ0006ｨｬｨｨ0202ｨｰ06ｨｬ Mucor hiemalis (Thormann et al., 2001, 2002b, 2003). 00ｨｨ0104 Sordaria fimicola, Sporormiella intermedia ｨｨ cf. Bjerkandera adusta ｨ 0405ｨｨ ｨｪ04 c 03ｨｨ ｨｰｨｺｨ ｨｪ02ｨｦ ｨｬ01ｨ, ｨ Acremonium cf. curvulum ｨｨ Oidiodendron maius ｪ 09 08ｨ 0405ｨ 00ｨ 06ｨｴ ｨ 00ｨｪｨｮｨｬｨ. 00ｨｨ0104 Mucor hiemalis, Oidiodendron scytaloides, Penicillium thomii ｨｨ Trichoderma viride ｨｨ05ｨｨ ｨｨ 09 03ｨｨ , ｨｨ 09 08ｨ 0405ｨ 00ｨ 06ｨｴ ｨｬ01ｨ. 0108ｨｨｨ Scleroconidioma sphagnicola 06ｨｰｨｬ02 ﾂ02ｨｪ ｨｺｨ ｨｺ ｨｨ00ｨｨ ﾂ0209ｨｺｨｨｨｦ ｨｰ08ｨｮｨｺｨｰ ｨ 00ｨｪｨｮｨｬｨ, ｨｨ ｨ ｨｪ ｨｨｨｺ05 08ｨ 0409ｨｨｨｰｨｨ07, ｨｬ ｨｪ0204 ｨｨ ｨｬ060800ｨｪ04ｨｦ ｨｺ06ｨｪｨｨ01ｨｨ060002ｨｪ0204 (Tsuneda et al., 2000, 2001). 00ｨｨ0104 Acremonium cf. curvum ｨｨ Oidiodendron maius ｨｰｨ ｨｺ0302 ｨｬ0600ｨｮｨｰ 08ｨ 0408ｨｮ03ｨ ｨｰｨｹ ｨｰ ｨｦｨｪｨｮ06 ｨｺ0502ｨｰ06 ﾂｨｪｨｮ06 09ｨｰ02ｨｪｨｺｨｮ ｨｨ ｨｪｨｨｨｺｨ ｨｰｨｹ 09 ｨｺ0502ｨｰｨｺｨｨ 0900ｨ 00ｨｪｨｮｨｬｨ (Thormann et al., 2001). 06ｨｪｨｰ ｨｪ04 01ｨ ｨｪｨｪ ｨｺ ｨｨｨｬ02ｨｪｨｰ ｨｨ07ｨｪｨｨ ｨ 03ｨｪ0609ｨｰｨｨ ｨｪｨ ｨ 04ｨｪｨｮ06 ｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ0609, ｨ ｨｰｨ ｨｺ0302 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0804 ｨｨ ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨｬ02ｨｪｨｰ0609 ｨｪｨ ｨ 04ｨｪｨｮ06 ｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ ｨｨ 08ｨ 0409ｨｨｨｰｨｨ02 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨｬ ｨｰ (ｨ ｨｰ ｨｰ09ｨｺｨｨｨｦ 488

5 13ｨｬ060103, ｨｺｨ 07 06ｨ 05., 05ｨ 07ｨ 01ｨｪ ｨｨｨｪ09ｨｺｨｨｨｦ 08ｨ ｨｦ06ｨｪ) (0502ｨｬ02ｨｪ0609 ｨｨ 0108., 1995) ｨ ｨｰｨ ｨ 03ｨｪ0609ｨｰｨｹ ｨｨ0008ｨ 02ｨｰ ｨ ｨｮ ｨｹ ｨｰｨｨ01 05ｨｺ0609ｨｨ09ｨｰ02ｨｬｨ 01 08ｨ 09ｨｰｨｨｨｰ0205ｨｹｨｪ ｨｰｨ ｨｰｨｺｨｨ 09 ｨｰ02- ﾂ02ｨｪｨｨ02 04ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ ｨｬ02ｨｪｨｨ 04ｨ ｨｰ ｨｪ ｨｦ, 0601ｨｪｨ ｨｺ ｨｪ0707 ﾂｨ 09ｨｰｨｹ ｨｰ ｨｪ ｨｨ0507 ｨｨ090704ｨｰ0409ｨ 02ｨｰ ｨｬ02ｨｪｨｪ0602 ｨｨ0909ｨｮ0302ｨｪｨｨ ｨｪｨｨ ｨｮ03ｨｨ09ｨ ｨｪｨｨ07 ｪｨｮ0905ｨ 03ｨｪ02ｨｪｨｨ07 06ｨｺｨ ｨ 02ｨｰ 09ｨｨ05ｨｹｨｪ ｨｰｨｨｨｬｨｮ05ｨｨ08ｨｮ06ｨｴ ｨｨ07ｨｪｨｨ02 ｨｪｨ ｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ0609 ｨｨ 09ｨｺ ｨｰｨｹ 08ｨ ｨｪｨｨ ｨ ｨｪｨｨ ﾂ0209ｨｺ ｨｴ0209ｨｰ09ｨ ﾂ ｨ ｨｨ04ｨｮ ﾂ02ｨｪｨ 04ｨ 09ｨｨ09ｨｨｨｬ0609ｨｰｨｹ ｨ 04ｨｪ06ｨｦ ｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｨ (0008) ｨｰ060800ｨ 06ｨｰ 0905ｨ 03ｨｪ0609ｨｰｨｨ, ｨｨ09ｨ 02ｨｬ06ｨｦ ｨｪｨ ｨｰ0607ｨｪｨｪ06ｨｬ ｨｮ080609ｨｪ02, ｨｨ 09 02ｨｨｨｺ05ｨ ｨｮ03ｨｨ09ｨ ｨｪｨｨ07 ｪｨｮ0905ｨ 03ｨｪ02ｨｪｨｨ ｨｨ ｨｬ02ｨｪｨｪ06ｨｬ ｨ ｨｪｨｨｨｨ 0905ｨ 00ｨｨ 09 ｨｰ ｨ ｨｨｨ 0605ｨｹ03ｨ ｨ 0405ｨ 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪｨ 0708ｨｨ ｨ ｨｪｨｨｨｨ 0905ｨ 03ｨｪ0609ｨｰｨｨ ｨｰ060800ｨ ｨｪｨ 09ｨ ｨｬ06ｨｬ ｨｪｨｨ04ｨｺ06ｨｬ ｨｨ04 ｨｨ090704ｨｰｨ ｨｪｨｪ0401 ｨｮ080609ｨｪ02ｨｦ (ｨｺ060502ｨ ｨ ｨｪｨｨ ｨ ｨｪｨｨ ｨ 00ｨｨ 40 ｪ 60 %). 0308ｨｨ 05ｨｰ06ｨｬ ｨ 09ｨｰｨ 05ｨ ｨｰ020702ｨｪｨｪ06 09 ｨｰ02 ﾂ02ｨｪｨｨ02 2 ｨｪ ｨｹ ｨｰｨｨ0302ｨｪｨｨ02ｨｬ ｨｬｨ ｨｺ09ｨｨｨｬｨｮｨｬｨ ｨｪｨ ｨｮｨｰｨｺｨｨ. 06ｨｪｨｺｨｮｨ ｨ 02ｨｨ07 06ｨ 08ｨ ｨｰ060800ｨ 0708ｨｨ ｨｺ06ｨｦ 0905ｨ 03ｨｪ0609ｨｰｨｨ 92 ｪ93 % ｨｪ ｨ 05ｨ 09ｨｹ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07ｨｬｨｨ ｨ 0609ｨｪ0609ｨ ｨｪｨｨｨｨ 01ｨ ｨｪｨｪ ｨｨｨｪｨ ｨｬｨｨｨｺ ｨ ｨｪ , ﾂｨｰ ｨ 09ｨｰｨ ｨｪｨｨ ｨ ｨｪ06 ｨｪ ｨｰｨｨｨｬｨｨ04ｨ 02ｨｨ02ｨｦ 0905ｨ 03ｨｪ0609ｨｰｨｨ, ｨ ｨｬ ｨ ｨｺｨｰｨｨ09ｨ 02ｨｨｨｨ ｨｰｨ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ0609, 09 ｨｰ06ｨｬ ﾂｨｨ ｨｨ ｨｨｨｰｨｨｨｺ0609, ｨｦ09ｨｰ09ｨｨ02ｨｬ 09ｨｰ ｨ ｨｮ03ｨｨ09ｨ ｨｪｨｨ07 ｪ080200ｨｨ0108ｨ ｨｰｨ 02ｨｨｨｨ. 02ｨ ｨｪｨｪ04ｨｦ ｨ ｨｰ ｨｪ ｨｰ060806ｨｦ ｨｨｨｨ 05ｨｺ ｨｨｨｬ02ｨｪｨｰ0609, 09 ｨｺ06ｨｰ ｨ 08ｨ ｨｰ060800ｨ ｨ 05ｨｨ09ｨｹ 02ｨｨｨｺ05ｨｨ ﾂ0209ｨｺ06ｨｬｨｮ ｨｮ03ｨｨ09ｨ ｨｪｨｨ06 ｪ ｨｮ0905ｨ 03ｨｪ02ｨｪｨｨ06. 01ｨ ｨｨｨ 0605ｨｹ03ｨ ｨｨ 05ｨｰ06ｨｬ 06ｨ ｨｪｨ 08ｨｮ03ｨｨ09ｨ 05ｨ 09ｨｹ 0908ｨ 04ｨｮ ｨｨ0108ｨ ｨｰｨ 02ｨｨｨｨ ｨｮ0302ｨｪｨｪ ｨｰ060800ｨ ｨｮ06ｨｴ02ｨｦ ｨｨｨｪｨｺｨｮｨ ｨ 02ｨｨｨｨ 0708ｨｨ ｨｰ0607ｨｪｨｪ06ｨｦ ｨｺ06ｨｦ 0905ｨ 03ｨｪ0609ｨｰｨｨ (92 ｪ93 %) ｨｰｨ 09ｨ 05ｨ 09ｨｹ ｨｺ06ｨｦ ｨｨ 06ｨ ｨｪｨ 08ｨｮ03ｨｨ09ｨ 05ｨ ｨｪ0204ｨ ｨｰｨｮ01ｨ 06ｨｴｨｨ ｨｨ ｨｨｨｨ 06ｨｰ ｨｬ ｨｮ02ｨｨ08ｨｮ06ｨｴｨｨ01 09ｨ 01ｨ /( ﾂ1 705) (01ｨｨ ｨ ｨｨ 0108., 1995). 06ｨｰ06ｨｦ ｨｮ070706ｨｦ ｨｮ ﾂ02ｨｪ0401 ｨ 0405ｨ ｨｪｨ 01ｨｨｨｪｨ ｨｬｨｨｨｺｨ 08ｨ ｨｪｨｨ07 ｨｰ060800ｨ 0708ｨｨ 08ｨ 04ｨｪ ｨ 01 ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨｦ ｨｨ 08ｨ 04ｨｪ06ｨｦ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0802 ｨｨｨｪｨｺｨｮｨ ｨ 02ｨｨｨｨ. 06ｨｬｨｨ ｨｪｨ 0008 ｨｨ ｨｪｨ 09ｨｬ02ｨｪｨ 09ｨｨ ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨ 08ｨ 0402ｨ ｨｰ060800ｨ 0708ｨｨ 09ｨｪ020902ｨｪｨｨｨｨ ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨｬ02ｨｪｨｰ0609 (N, P, K, S) ｨｨ 08ｨ 04ｨｪ0401 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨ 01 (25 ｨｨ 38 紊). 01ｨ ｨｨｨ ｨｪｨｨ04ｨｺｨ , ｨｪ06 ｨｪｨ ｨｨｨ 0605ｨｹ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨ02 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨ ｨｪｨ 08ｨｮ0302ｨｪ ｨｰ09ｨｮｨｰ09ｨｰ09ｨｨ02 ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨ ｨ 0906ｨｺ 0708ｨｨ 25 紊 ｨｪｨｨ02 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ 紊 ｨｨ 09ｨｪ020902ｨｪｨｨ02 ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨｦ 0708ｨｨ090601ｨｨ0506 ｨｺ ｨ 09ｨｰｨ ｨｪｨｨ ｨｨ ｨｺ 09ｨｪｨｨ0302ｨｪｨｨ06 ｨｨ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ06 08ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨ ｨｬ0401 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨｪｨｨ ｨｨ 08ｨ 04ｨｪ0401 ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨ0701 ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨｦ (50, 250, 1000 ｨｬ00 N/ｨｺ00) ｨｨ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ 紊 0706ｨｺｨ 04ｨ 0506, ﾂｨｰ06 ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪｨ ｨ 0405ｨ 0601ｨｨｨｪｨ ｨｺ060906ｨｦ ｨ 08ｨｨｨ ｨｪｨｰｨ 01, ｨ ｨｪｨ ｨｨｨ ｨｨｨｰ0205ｨｹｨｪ04ｨｦ 09ｨｰｨｨｨｬｨｮ05ｨｨ08ｨｮ06ｨｴｨｨｨｦ ｨｺｨｰ ｨｪｨ ｨ ｨｪｨ 08ｨｮ0302ｨｪ 0708ｨｨ ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨｨｨ ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ ｨｦ 1000 ｨｬ00 N/ｨｺ00. 00ｨｪ020902ｨｪｨｨ02 Cu ｨｨ Zn 06ｨｺｨ ｨ 0506 ｨｪ0204ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ ｨｨ07ｨｪｨｨ02 ｨｨ ｨ ｨ 04ｨ ｨ 09ｨｪ02ｨｪｨｨ06 09 ｨｺ06ｨｪｨｰ ｨｬ (0502ｨｬ02ｨｪ0609 ｨｨ 0108., 1995) ｨ ｨｪｨｨ07 04ｨ 09ｨｨ09ｨｨｨｬ0609ｨｰｨｨ 09ｨｺ ｨｰｨｨ 08ｨ ｨｪｨｨ07 Carex aquatilis ｨｨ Sphagnum fuscum 06ｨｰ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ ｨｺｨ 04ｨ 05ｨｨ, ﾂｨｰ ｨｨｨ ｨｪ ｨ ｨｪｨｨ ｨ ｨｰｨ 01 ｨｨ0102ｨｰ ｨｬ ｨｪｨｪ ｨｪｨｨ02ｨｬ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ ｨｰ 紊 (Thormann et al., 2004). 00 ｨ ｨｰｨｪ ｨ ｨｴ0209ｨｰ09ｨ 01 06ｨ ｨｪｨ 08ｨｮ0302ｨｪ04 ｨ ｨｨｨ 02ｨｨｨｨ ｨｬ01ｨ Polytrichum commune ｨｨｨ ｨ ｨｬｨｨ, ｨｨ01 06ｨｰｨｪ060302ｨｪｨｨ07 08ｨ ｨｪｨｨ09ｨ 06ｨｰ ｨｺｨ ｨｺ 09ｨｨｨｬｨ ｨｨ06ｨｰｨｨ ﾂ0209ｨｺｨｨ02, 00ｨｨ ｨｨｨ ｨ ｨｪｨ ｨ ｨ 05ｨｨ ｨｪ0609ｨｪ0401 ｨｺ0502ｨｰｨｺｨ 01 ｨｬ01ｨ, ｨｺ06ｨｰ060804ｨｦ 0708ｨｨ 05ｨｰ06ｨｬ ｨｪ02ｨｮ00ｨｪ02ｨｰ02ｨｪｨｪ04ｨｬ (Gross, Robbins, 2000) ｨｪｨｨｨｺｨｨ 08ｨ 04ｨｪ06ｨｦ ｨｬ06ｨｴｨｪ0609ｨｰｨｨ 06ｨｰ05ｨｨ ﾂｨ 06ｨｰ ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨｨｨ 09 ｨｪｨｨ01 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ0609 ｨｨ 01ｨ 08ｨ ｨｺｨｰ0208ｨｮ 08ｨ ｨｪｨｨ ｨｨｨ ｨｪ06ｨｦ ｨｨ ｨｬｨｨｨｺ0806ｨ ｨｪ06ｨｦ ｨ ｨｨ06ｨｬｨ ｨｨ ｨｬｨ 0506ｨｬ06ｨｴｨｪ06ｨｬ ｨｰ ｨｪｨｨｨｺ02 09ｨｮｨｬｨｬｨ 08ｨｪｨ 07 ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨ07 05ｨｮｨｺｨ 08ｨｨ06ｨｰｨｪ0401 ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ , ﾂ02ｨｬ 09 ｨｬ06ｨｴｨｪ0401 ｨｰ ｨｪｨｨｨｺｨ 01, ｨｨ ｨｰ06 ﾂ02ｨｪｨ ｨｪ02ｨｦ ｨｮｨｬ02ｨｰ ｨｦ ｨｰ0605ｨｴ ｨｬ06ｨｴｨｪ0401 ｨｰ ｨｪｨｨｨｺｨ 01 06ｨｪｨ 08ｨ ｨｪｨ ｨｨ ｨ 09ｨｪ06ｨｬ0208ｨｪ06 ( ﾂ02ｨｪｨｺ06 ｨｨ 0108., 2002) ｨｪｨｪｨ 07 01ｨｨｨｪｨ ｨｬｨｨｨｺｨ ｨｬｨｨ020205ｨｨｨ 05ｨｹｨｪ ｨｮｨｺｨ 08ｨｨ06ｨｰｨｪ ｨｬ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ0609 ｨ 0405ｨ ｨ 0302ｨｪｨ ｨｰ0605ｨｹｨｺ06 09 ｨｪｨｨ04ｨｨｨｪｨｪ0401 ｨｰ ｨｪｨｨｨｺｨ 01 ( ﾂ02ｨｪｨｺ06 ｨｨ 0108., 2002). 00 ｨｮ ｨｨ ｨ ｨｰｨ, ｨｨ04-04ｨ ｨｰ ﾂｨｰ06 08ｨ ｨｪｨｨ ｨｰ02ｨｺｨ 02ｨｰ ｨｪ02ｨｨｨｪｨｰ02ｨｪ09ｨｨ09ｨｪ06, 06ｨｪ06 ｨｨｨｬ0202ｨｰ 08ｨ 09ｨｰ07ｨｪｨｮｨｰ0602 ｨｪｨ ﾂｨ 0506 ｨｨ ｨｪ02 01ｨ 02ｨｰ ﾂ02ｨｰｨｺｨｨ ｨｪｨｪ ｨｺ (05ｨｬｨ 00ｨｨｨｪｨ, 1988). 07 ﾂｨ 09ｨｰｨｨ ｨｨｨ ｨｰ08ｨｮｨｺ02ｨｨｨｨ ｨｮ03ｨｪ0401 ﾂｨ 09ｨｰ02ｨｦ 0900ｨ 00ｨｪｨｮｨｬｨ 0706ｨｺｨ 04ｨ ｨｪ06 ｨｪ02ｨｺ06ｨｰ060804ｨｬｨｨ ｨｨ ｨ ｨｰ020507ｨｬｨｨ, ｨｬ06ｨｪ09ｨｰ08ｨｨ080609ｨ ｨｪｨ ｨ ｨｪ0609ｨｰｨｹ 0008ｨｨｨ 0609, ｨｪｨｪ ｨｬ01ｨ, 08ｨ 0408ｨｮ03ｨ ｨｰｨｹ ｨｮ, ｨｺ08ｨ 01ｨｬｨ 05, 01ｨｮｨ ｨｨ05ｨｹｨｪ0402 ｨｺｨｨ090506ｨｰ04 (Tsuneda et al., 2001). 0201ｨｪｨ ｨｺ06 ｨｨ ｨｰｨｪ06, ﾂｨｰ06 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨｰ ｨｨ04 06ｨ 08ｨ , 06ｨｰ06ｨ 08ｨ ｨｪｨｪ ｨｬｨｮ ｨｨ0506 ｨｰ ｨｪｨｨｨｺ0609, 0708ｨｨ ﾂ02ｨｬ ﾂｨ 09ｨｰ06 ｨｺ0605ｨｨ ﾂ0209ｨｰ0906 ｨｺ060506ｨｪｨｨ0206ｨ 08ｨ 04ｨｮ06ｨｴｨｨ ｨｨｨｪｨｨ02 (080203) ｨｨ ｨｮｨ ｨｨｨｪ- 489

6 13ｨｪ ｨ 08ｨ ｨｰ ｨｪｨｨ02 ｨｬ0603ｨｪ06 06ｨ ｨｲ0709ｨｪｨｨｨｰｨｹ 09ｨｬ040906ｨｬ ｨｨ ｨｪ0609ｨｰｨｪ , ｨｪ ｨｰｨ 02ｨｰ ｨｰｨｺ0804ｨｰ04ｨｬ ｨｰ0607ｨｪｨｨｨｨ ｨｨ ﾂｨ 09ｨｰ02ｨｦ ｨｬｨｨ020205ｨｨ07 ｨｪｨ 0005ｨｮｨ ｨｨｨｪ02: ｨｪｨ ｨｰ ｨｨ 06ｨｪｨｨ 09 09ｨｰｨ 01ｨｨｨｨ 0706ｨｺ0607, ｨｨ05ｨｨ ｨ ｨｪ04 ｨｺ ｨｰｨｮ ｨｨ 00ｨｮｨｪｨｺ02ｨｨ06ｨｪｨｨ080609ｨ ｨｪｨｨ06 09 ｨｮ ｨｨ ｨｨ02ｨｨｨｰｨ ｨｺｨｨ ｨ. 04ｨ ｨ 06ｨｰ ｨｨ ｨ ｨｪｨｨ06 08ｨ 0409ｨｨｨｰｨｨ ｨｨｨ ｨｮ ｨｨ0701 ｨｪ ｨｰｨ ｨｰｨｺｨ ｨｺｨｨ ｨ ｨｨ05ｨｨ09ｨｹ 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ04ｨｬｨｨ ｨｨ ｨ ｨｰ020507ｨｬｨｨ ｪ (08ｨｮ08ｨ ｨｺ0609 ｨｨ 0108., 2007; ｨｺｨｨｨｪｨ, 08ｨｮ08ｨ ｨｺ0609, 2007; 0108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008; ｨ ｨ ｨｺ0609ｨ ｨｨ 0108., 2008) ｨ ｨｪｨｪ0401 ｨｨ ｨ ｨｪｨｨ ｨｬ06ｨｪ09ｨｰ08ｨｨ080609ｨ ｨｪｨ ｨ ｨｪ0609ｨｰｨｹ ｨｺ 00ｨｮｨｪｨｺ02ｨｨ06ｨｪｨｨ080609ｨ ｨｪｨｨ06 09 ｨｮ ｨｨ0701 ｨｪ ｨｰｨ ｨｰｨｺｨ ｨｺｨｨ ｨ ｨ 09ｨｨ ｨｨｨ 0609, ﾂｨｰ ｨｰ ｨｨｨｰｨｹ ｨｨ ｨｬ0603ｨｪ0602 ｨｮ ﾂｨ 09ｨｰｨｨ02 09 ｨｺ06ｨｬ070502ｨｺ ｨｪ ｨｰ ｨ 08ｨ ｨ ｨｪｨｨ07, ｨｨ ｨｴｨｨ01 ｨｪ02 ｨｰ0605ｨｹｨｺ06 ｨｪｨ ｨｪ0609ｨｰｨｨ, ｨｪ06 ｨｨ ｨｮｨ ｨｨｨｪｨｪ ｨｰ ｨｪｨｨｨｺ ｨｺ ｨｨ ﾂ0209ｨｺｨｨ ｨ 02ｨｪｨｪ0609ｨｰｨｨ ｨｪｨ ｨｨｨ ｨ ｨｰ08ｨ ｨｪ02ｨｪｨｪ ｨｨ ｨｨｨ ｨｰ ｨｪｨｨｨｺｨ ｨ ｨｰ ｨｰｨ 0203ｨｪ06ｨｦ 0406ｨｪ ｨ ｨｪｨｨ07 ｨｰ ｨｪｨｨｨｺ ｨ ｨｰ ｨｪ06ｨｰｨ 0203ｨｪ06ｨｦ 0406ｨｪ ｨｺｨ 04ｨ 05ｨｨ, ﾂｨｰ , ｨｺ06ｨｰ ｨｬｨｨｨｪｨｨ08ｨｮ06ｨｰ 09 01ｨ ｨｪｨｪ ｨ ｨｴ0209ｨｰ09ｨ 01 (Geomyces, Cladosporium ｨｨ Penicillium), ｨ 01ｨ 06ｨｰ ｨ ｨｬｨｨ, ｨｪｨ ｨｨｨ ﾂｨ 09ｨｰ ｨｰ0802 ﾂｨ 06ｨｴｨｨｨｬｨｨ ﾂ0902ｨｪｨｪ ｨ 08ｨ 0402ｨ ｨｪ ﾂｨｪ06ｨｦ ｨｬ ｨｰ04 (0108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008; 0806 ﾂｨｺｨｨｨｪｨ ｨｨ 0108., 2008) ｨｦ 09ｨｨ01ｨｨｨｬ0609ｨｰｨｨ, 05ｨｰｨｨ 0008ｨｨｨ ｨｰ0907 ﾂｨ 09ｨｰｨｹ ｨｨ ｨｨ05ｨｹｨｪ ｨ ｨｴ0209ｨｰ09 ｨｨ ｨ ｨｪ04 ｨｺ ｨ ｨｺｨｰｨｨ09ｨｪ06ｨｬｨｮ 09ｨｮｨｴ0209ｨｰ090609ｨ ｨｪｨｨ ｨｨ 0905ｨ ｨ ｨｰ08ｨｨ02ｨ ｨｰ0205ｨｹｨｪ0401 ｨｨ ｨｨｨｰ0205ｨｹｨｪ0401 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨ ｨｰｨ ｨｪ0609ｨｨｨｬ0907 ｨ ｨ ｨｪ06 ｨｪｨ 09ｨ ｨｬ ｨ ｨｰ08ｨ ｨｪ02ｨｪｨｪ ｨｪ0401 ｨｰ ｨｪｨｨｨｺｨ 01 09ｨｨ01ｨ 01 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨ 0405ｨｨ ﾂｨｪ ｨ 00ｨｨ ﾂ0209ｨｺｨｨ01 ｨｰ06 ﾂｨｺｨ ｨ 0605ｨｹ0306ｨｬ ｨｺ0605ｨｨ ﾂ0209ｨｰ0902 ｨｨ ｨ ｨｰ ｨｨ 0706 ﾂ ｨｰ 09ｨｨ01 Penicillium spinulosum ｨｪ06ｨｦ 08ｨ ｨｪｨ ｨｨ05ｨｨ09ｨｹ ｨｨ ｨ ｨｪｨｨ07 09ｨｨ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨ ｨｨｨ ｨｨｨｺ0608ｨｪ020906ｨｦ 0406ｨｪ ｨｨ (Picea mariana), ｨｺ06ｨｰ0608ｨ 07 06ｨ ｨｨｨｰｨ 02ｨｰ ｨｨ09ｨｰ06ｨｦ 0706 ﾂ0902 ｨｨ 0706 ｨｺ08ｨ 07ｨｬ ｨ ｨｰ ｨ ｨｰ0205ｨｨ ｨｪｨ 0409ｨ 05ｨｨ P. spinulosum ｨｰｨｨ07ｨｨ ﾂｨｪ04ｨｬ 09ｨｨ0106ｨｬ 00ｨｮｨｬｨｮ ｨｨ0406ｨｪｨｰｨ ｨｪ ｨｨ ｨ ｨｰ ( ｨ ｨｨ 0108., 2004; Summerbell, 2005). 0604ｨｮ ﾂ02ｨｪｨｨ ﾂ ｨｪ ｨ ｨｴ0209ｨｰ ｨｺ080209ｨｰｨｪ0609ｨｰ ｨｬ060809ｨｺ06ｨｦ ｨ ｨｨ ｨｨ ﾂ0209ｨｺ06ｨｦ 09ｨｰｨ ｨｪ02ｨｨｨｨ (090905) ｨｨｨｬ ｨ , 0706ｨｺｨ 04ｨ 0506, ﾂｨｰ06 P. spinulosum ｨｰｨ ｨｺ ｨ 05ｨ 01ｨ ﾂｨ 09ｨｰ06ｨｰ ｨｰ0802 ﾂｨ 02ｨｬ0609ｨｰｨｨ 09 00ｨｮｨｬｨｮ ｨｬ ｨｨ0406ｨｪｨｰ02 (09ｨｮｨ ｨｪ0609ｨ, 2005; 0108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008) ｨ ｨｪｨｪ06ｨｦ 06ｨ 05ｨ 09ｨｰｨｨ ｨｰｨｨ07ｨｨ ﾂｨｪ ｨｨ09ｨｰ ﾂ0904, 06ｨｪｨｨ, ｨｺｨ ｨｺ ｨｰ ｨｪｨｨｨｺｨｨ ｪ ｨｨｨｬ0206ｨｰ ｨｪｨｨ04ｨｺｨｨ02 04ｨｪｨ ﾂ02ｨｪｨｨ ｨｨｨｰ0607ｨｪ04ｨｦ 09ｨｨ01 P. spinulosum ｨｰｨｨ07ｨｨ ﾂｨｪ04ｨｦ 06ｨ ｨｨｨｰｨ ｨｰ0205ｨｹ ｨｪ ﾂ09. 02ｨｪ 08ｨ ｨｰ08ｨ ｨｪ02ｨｪ ﾂ09ｨ ｨｪ ｨ 05ｨ 09ｨｰ02ｨｦ, 06ｨｺ ｨｬ060807, 09 08ｨ ｨｪｨ 0102, 0106ｨｰ05ｨ ｨｪ01ｨｨｨｨ ｨｨ ｨｰ. 01., 09 ｨｰｨｮｨｪ ｨｦ ｨｨ ｨｰｨ 0203ｨｪ06ｨｦ 0406ｨｪｨ 01, ﾂ09ｨ ｨｨｨｺ0608ｨｪ ｨｪ ｨｪ04 ｨｨ 0205ｨｨ, ｨ ｨｰｨ ｨｺ ｨ ｨｰｨ 01 08ｨ 04ｨｪ0401 ｨｰｨｨ ｨｰｨｬ02 ﾂ02ｨｪｨ ｨｺｨ ｨｰ0802 ﾂｨ 02ｨｬ0609ｨｰｨｹ P. spinulosum ﾂ09ｨ ｨｪｨｪ06ｨｦ ｨｺｨｨ090506ｨｰｨｪ0609ｨｰｨｹ ｨｨｨ 08ｨ 09ｨｰ02ｨｰ 09 03ｨｨ0806ｨｺ06ｨｬ 01ｨｨｨ 07ｨ 0406ｨｪ02 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08: 06ｨｰ 5 ｪ ｪ42 紊 ( ｨｪｨｨ ｨｰ ｨｪｨｨｨｺ ｨ 08ｨ ｨｺｨｰ0208ｨｪ04 09ｨｨ05ｨｹｨｪ ｨ 0104 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08) 0708ｨｨ 0607ｨｰｨｨｨｬｨ 05ｨｹｨｪ06ｨｬ 04ｨｪｨ ﾂ02ｨｪｨｨｨｨ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ ｨｰｨ 09 ｨｨｨｪｨｰ020809ｨ ｪ28 紊. 06ｨｰ ｨｨｨｪ ｨｨ04 ｨｪ02ｨｬｨｪ0600ｨｨ01 09ｨｨ010609, ｨｺ06ｨｰ ｨｰｨｬ02 ﾂ02ｨｪｨ ｨ ｨｪ0609ｨｰｨｹ ｨｺ 08ｨ ｨｪｨｨ ｨ 00ｨｪ ｨｬ (Domsch et al., 1980, 2007). 06ｨ ｨｺｨｨ02 01ｨ ｨｪｨｪ ｨｴ02 08ｨ ｨｰ ｨ 06ｨｰ 04ｨ ｨｺ06ｨｪ06ｨｬ0208ｨｪ0609ｨｰｨｹ ｨｰ0607ｨｪｨｪ ｨ ｨｪｨ 08ｨｮ0302ｨｪｨｨ07 09ｨｨ01ｨ P. spinulosum 09 ｨｰ ｨｪｨｨｨｺｨ 01 ｨｨ ｨｪ ｨｪｨｨｨｬｨｨ 0706 ｨｮ ｨｨ07ｨｬ ｨｬ0209ｨｰ0606ｨ ｨｨｨｰｨ ｨｪｨｨ ｨｮ00ｨｨ02 09ｨｨ ｨ Penicillium, ｨｨ040605ｨｨ080609ｨ ｨｪｨｪ0402 ｨｨ ｨ ｨｰ020507ｨｬｨｨ 09 ｨ 0605ｨｹ0306ｨｬ ｨｺ0605ｨｨ- ﾂ0209ｨｰ0902 ｨｨ04 ｨｰ ｨｪｨｨｨｺ ｨ ｨｰ, ｨｰｨ ｨｺ0302 ｨｪ ｨｮ ﾂｨ ｨｦｨｪ ｨ ｨｴ0209ｨｰ09ｨ 01 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ0609, ｨｪｨｪ0401 ｨｨ04 ｨｺｨｨ ﾂ09, ｨｮｨｺｨ 04ｨ ｨｪ P. thomii ( ｨｺｨ 07, 1980; Domsch et al., 1980, 2007; 09ｨ ｨ ｨｹ0209ｨ, 0502ｨｪ0609ｨ, 1989). 06ｨｰ06ｨｰ 09ｨｨ01 ﾂｨ 09ｨｰ ｨｰ0802 ﾂｨ 02ｨｰ ｨｰｨｮｨｪ , ｨｨ09ｨｰ ﾂ09ｨ 01, ｨ ｨｰｨ ｨｺ ｨｰ ｨｪｨｨｨｺｨ 01 ｨｨ ｨｪｨ 08ｨ 09ｨｰ02ｨｪｨｨ ｨ ｨｰ. 02ｨｪ ｨ 02ｨｪ 08ｨ 0409ｨｨ09ｨ ｨｰｨｹ ｨｨ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨ 01 06ｨｰ 紊 ｨｬ ｨｨｨｰｨｪ04ｨｦ 09ｨｨ01 P. funiculosum, ｨ ｨｪ04ｨｦ ｨｺ ｨｰｨｮ ｨｨ 08ｨ 0409ｨｨｨｰｨｨ ｨｨ0806ｨｺ06ｨｬ 01ｨｨｨ 07ｨ 0406ｨｪ02 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08 (5 ｪ42 紊), 06ｨ ｨｨｨｰｨ 02ｨｰ ｨｰｨ ｨｺ ｨｺ09ｨｰ0802ｨｬｨ 05ｨｹｨｪ ｨｪ0401 ｨｮ ｨｨ0701 ｨｨ ｨｪｨ ｨｺｨｨ ｨｮｨ 09ｨｰ08ｨ ｨｰｨ 01, 0909ｨｰ0802 ﾂｨ 02ｨｰ ｨｰｨｨ07ｨ ﾂ09, 09 ｨｰ ｨｪｨｨｨｺｨ ｨ ｨｰ ｨｨ ｨｬｨ ﾂ09ｨ 01 ｨｰｨ ｨｦ00ｨｨ (Domsch et al., 1980, 2007). P. funiculosum ｨｰｨ ｨｺ ｨｰｨｬ02 ﾂｨ 02ｨｰ0907 ｨｮ ｨｬｨｪ0600ｨｨ01 ｨｨ ｨ ｨｰ020502ｨｦ 09 08ｨ ｨ 06ｨｰｨ ｨｨ04ｨｮ ﾂ02ｨｪｨｨ06 09ｨｨ ｨｰｨ 09ｨ 0008ｨｨｨ 0609, 06ｨ ｨｨｨｰｨ 06ｨｴｨｨ01 09 ｨｰ ｨｪｨｨｨｺｨ 01 (Thorman et al., 2004; 490

7 130108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008) 05ｨｮｨｴ0209ｨｰ09ｨｮ06ｨｰ 01ｨ ｨｪｨｪ ｨｰ06ｨｬ, ﾂｨｰ06 P. funiculosum ｨ 02ｨｪ 08ｨ 0408ｨｮ03ｨ ｨｰｨｹ 0900ｨ 00ｨｪ ｨｬ01ｨｨ (Domsch et al., 1980, 2007). 00ｨｨ0104 P. glabrum ｨｨ P. implicatum ｨｰｨ ｨｺ ｨ 08ｨ ｨｺｨｰ0208ｨｪ ｨｨ09ｨｰ0401 ｨｨ 0108ｨｮ00ｨｨ01 ｨｺｨｨ ﾂ09, ｨｪｨｨ04ｨｨｨｪｨｪ0401 ｨｨ ｨ ｨｰ. P. glabrum 06 ﾂ02ｨｪｨｹ 08ｨ ｨｰ08ｨ ｨｪ02ｨｪ 09 08ｨ 04ｨｪ ｨｨ06ｨｪｨ ｨｬ05ｨｨ ｨｨ ﾂｨ 09ｨｰ06 06ｨｰｨｬ02 ﾂｨ 02ｨｰ ｨｪ ｨ 05ｨ 09ｨｰ0701, 09 ｨｰ06ｨｬ ﾂｨｨ ﾂ09ｨ 01 ｨ 08ｨｺｨｰｨｨ ﾂ0209ｨｺ06ｨｦ ｨｰｨｮｨｪ010804, 09 06ｨｺ080209ｨｰｨｪ0609ｨｰ ｨｬ060807, 00ｨｮ08ｨｬｨ ｨｪ09ｨｺｨ, ｨｪ06ｨｦ ｨｨｨｨ (Domsch et al., 1980, 2007; Nilsson et al., 1992). 00ｨｨ ｨ Oidiodendron ｨｨ ｨｰｨｪ04 ｨｺｨ ｨｺ 06ｨ 04 ﾂｨｪ ｨ ｨｨｨｰｨ ｨｰ0205ｨｨ ｨｰ ｨｪｨｨｨｺ0609, ｨ ｨｪ0402 ｨｺ 08ｨ 0408ｨｮ0302ｨｪｨｨ ｨ 00ｨｪ ｨｬ (Thormann et al., 2001, 2002b, 2004; Sigler et al., 2005). 02ｨｪｨｨ ｨｬ0600ｨｮｨｰ 08ｨ 09ｨｰｨｨ ｨｨ 08ｨ 0409ｨｨ09ｨ ｨｰｨｹ ｨｨ 04ｨｪｨ ﾂ02ｨｪｨｨ ｨｪｨｨ (Gross, Robbins, 2000). 00 ｨｨ ｨ ｨｪｨｨ0701 ｨ ｨｰ ｨｪ ｨｨ0806ｨｰ O. griseum ｪ 0601ｨｨｨｪ ｨｨ ｨｬｨｨｨｪｨｨ08ｨｮ06ｨｴｨｨ ｨｨ ｨｰ0802 ﾂｨ 06ｨｴｨｨ ｨｹ 09ｨｨ ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ0609 (Thorman et al., 2004, 2006; 0108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008) ｨｮ05ｨｹｨｰｨ ｨｰ02 05ｨｺ ｨｨｨｬ02ｨｪｨｰｨ ｨｪ02ｨｪｨｨ ｨ ｨｪ0609ｨｰｨｨ ｨｺ ｨｰｨｮ ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ｨｨ ｨｪｨｨ04ｨｺｨｨ01 04ｨｪｨ ﾂ02ｨｪｨｨ ｨｺｨ 04ｨ ｨｹ, ﾂｨｰ06 O. griseum ｨｬ060302ｨｰ 08ｨ 0409ｨｨ09ｨ ｨｰｨｹ ｨ ｨｬ06ｨｬ 03ｨｨ0806ｨｺ06ｨｬ ｨｨ ｨｨ ｨ ｨｪｨｪ ｨｨ ｨ ｨｪｨｪ ｨ 01ｨｨｨ 07ｨ 0406ｨｪ : 06ｨｰ (Gross et al., 2000). 00ｨｨ0104 O. griseum ｨｨ O. tenuissimum ｨｰ ｨ 04 ﾂｨｪ04ｨｬｨｨ ｨｺ06ｨｬ0706ｨｪ02ｨｪｨｰｨ ｨｬｨｨ 0708ｨｨｨｺ0608ｨｪ020906ｨｦ 0406ｨｪ ｨｺ ｨ 09ｨｰ02ｨｪｨｨｨｦ, ｨ ｨｰｨ ｨｺ0302 ｨｰ ｨｪ0401 ｨｨ ｨｪ ｨｨ09ｨｰ ﾂ09 (Sigler et al., 2005; Thormann, 2006). 00ｨｨ01 O. priconioides ｨｰｨ ｨｺ0302 ﾂｨ 09ｨｰ ｨｰ0907 ｨｨ ｨ 00ｨｪ ｨ ｨｰ (Rice, Currah, 2005). 00ｨｨ01 Geomyces pannorum, 0706 ｨｬｨｪ02ｨｪｨｨ ｨ ｨｨ ｨ ｨｰ020502ｨｦ, 06ｨｰｨｪ0609ｨｨｨｰ0907 ｨｺ ｨｬ ｨｨ05ｨ ｨｬ, ｨｪ06 09 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07ｨｬｨｨ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨｪ ｨ ｨｪｨｨ ｨｰ060806ｨｪｨｮ 0709ｨｨ0106ｨｰ ｨ ｨｪｨｰｨｪ0609ｨｰｨｨ. 02ｨｪ 06ｨ ｨｨｨｰｨ 02ｨｰ ｨｨｨｬｨｮｨｴ0209ｨｰ0902ｨｪｨｪ ｨｪ ﾂ09ｨ 01 ｨｰｨｮｨｪ ｨｮ ｨｨ ｨｰ0209ｨｰ0902ｨｪｨｪ06ｨｦ ｨｺ08ｨｨ06ｨｺ06ｨｪ ｨ 02ｨｨｨｨ (00ｨｨ08 ﾂｨｨｨｪｨｺ, 1976; 0806 ﾂｨｺｨｨｨｪｨ ｨｨ 0108., 2008). 02ｨ ｨｪｨｪ04ｨｦ 09ｨｨ01 ｨｨ ｨｰ02ｨｪ ｨｺｨ ｨｺ 05ｨｺ09ｨｰ0802ｨｬ06ｨｰ ｨ ｨｪｨｰｨｪ04ｨｦ, ｨｺ06ｨｰ060804ｨｦ ｨｬ060302ｨｰ 08ｨ 0409ｨｨ09ｨ ｨｰｨｹ0907, ｨｪｨ ｨｹ ｨｦ09ｨｰ09ｨｨ02ｨｬ ｨｬｨｪ ｨｰ0902ｨｪｨｪ ｨｰ , ｨｰｨ ｨｺｨｨ01 ｨｺｨ ｨｺ ｨｪｨｨ04ｨｺｨｨ02 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0804, ｨｪｨｨ04ｨｺｨ 07 ｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ ｨｨ (0806 ﾂｨｺｨｨｨｪｨ ｨｨ 0108., 2008). 06ｨｰ06ｨｰ 09ｨｨ01 03ｨｨ04ｨｪ ｨ 02ｨｪ 09 03ｨｨ0806ｨｺ06ｨｬ 01ｨｨｨ 07ｨ 0406ｨｪ02 04ｨｪｨ ﾂ02ｨｪｨｨｨｦ 0801 ｪ 3.5 ｪ8.0 (Oorschot, 1980). 0108ｨｨｨ ｨ Geomyces ｨｰｨ 09ｨｨ05ｨｨ 0609ｨｪ0609ｨｪｨｮ06 ﾂｨ 09ｨｰｨｹ ｨｬｨｨｨｺ06ｨ ｨｨ06ｨｰ04, 06ｨ ｨｪｨ 08ｨｮ0302ｨｪｨｪ06ｨｦ 09 ｨｬｨｪ ｨｰｨｪ02ｨｬ ｨｰ ｨｪｨｨ ｨｺｨｰｨｨｨｺｨｨ (020208ｨ ｨ ｨｺ0609ｨ ｨｨ 0108., 2008) ｨｹ ｨｪｨｨ07 09ｨｰ08ｨ ｨｰ0200ｨｨｨｦ ｨｨ09ｨ ｨｪｨｨ07 ｨｺｨｮ05ｨｹｨｰｨｮ08 G. pannorum, ｨｪｨｪ0401 ｨｨ04 ｨｬｨｪ ｨｰｨｪ02ｨｦ ｨｬ ｨｰ04, 01ｨ ｨｪｨｪ04ｨｬｨｨ ｨｨ ｨ ｨｰ020507ｨｬｨｨ ｨ 0405ｨ ｨｪｨ ｨ ｨｪ0609ｨｰｨｹ ｨ 08ｨｺｨｰｨｨ ﾂ0209ｨｺ ｨｨ ｨｰｨ ｨｨ 03ｨｰｨ ｨｬｨｬｨ, ｨｪｨｪ ｨｨ ｨｰｨｨ ｨｦ ｨｺｨｨ ｨｪ02ｨｦ ｨｨｨｨ, ｨｺ ｨｰｨｮ 0708ｨｨ ｨ ｨｪｨｨ ﾂ02ｨｪｨｨｨｨ ｨｺｨｨ ｨ ｨｨ 08ｨ 0405ｨｨ ﾂｨｪ ｨｺｨｨ0905ｨｨｨｰ0205ｨｹｨｪ ｨｰｨ ｨｪ0609ｨｨｨｰ0205ｨｹｨｪ0401 ｨｮ ｨｨｨｦ ｨｮ05ｨｹｨｰｨ ｨｰ ｨｺｨ 04ｨ 05ｨｨ, ﾂｨｰ06 09ｨｪｨｨ0302ｨｪｨｨ02 ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨｨｨ ｨｺｨｨ ｨ 09 00ｨ ｨｦ 00ｨ ｪ ｪ2 % ｨｪ02 ｨｨｨｪ00ｨｨｨ ｨｨ080609ｨ ｨｰ ｨｨ ｨｮ02ｨｬ ｨｰｨ ｨｬｨｬ ｨｬｨｪ02ｨｪｨｨ06 ｨ 09ｨｰ , ｨｪｨｪ0402 ｨｪ ｨｨ04ｨｨ ｨｨ ﾂ0209ｨｺｨｨ ｨ 02ｨｪｨｪ0609ｨｰｨｨ 06ｨ ｨｲ0709ｨｪ0706ｨｰ ｨ ｨｪ0609ｨｰｨｹ 0008ｨｨｨ 0609 G. pannorum ｨｨ09ｨ ｨｰｨｹ 09 ｨ ｨｪｨ ｨ ｨｪ0401 ｨｮ ｨｨ0701, 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ ｨｺ08ｨｨ ﾂｨｪ06ｨｦ ｨｬ ｨｰ02 (020208ｨ ｨ ｨｺ0609ｨ ｨｨ 0108., 2008). 02ｨｰｨｬ02ｨｰｨｨｨｬ, ﾂｨｰ ｨｮｨ ｨｨｨｪｨｪ ｨｰ ｨｪｨｨｨｺ0609 ｨｰｨ ｨｺ ｨ 08ｨ ｨｺｨｰ0208ｨｪ04 ｨ ｨｪｨ ｨ ｨｪ0402 ｨｮ ｨｨ07 ｨｨ ｨｪｨｨ04ｨｺｨｨ02 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0804, ｨｰｨ ｨｨ 08ｨ 0409ｨｨｨｰｨｨ07 09 ｨｺ06ｨｰ ｨｰ0802ｨ ｨｮ06ｨｰ0907 ｨｰ ｨｨ04ｨｨ ｨｨ ﾂ0209ｨｺｨｨ ｨ 02ｨｪｨｪ0609ｨｰｨｨ, ｨｨ ｨｪｨｨ02 ｨｪｨ ｨｬｨｨ 01ｨ ｨｪｨｪ ｨｨ01ｨ ｨｮｨ ｨｨｨｪ04 1 ｨｬ ｨｨ04 06ｨ 08ｨ 0402ｨ ｨｰ060800ｨ ｨｪ ｨ ｨｰｨ ｨｺ06ｨｦ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｹ06 ( ｨｺｨ ｨｪｨ ｨｮ0106ｨｦ ｨｬｨ ｨ 08ｨ 0402ｨ ) ｨｪ ｨｮ ﾂｨ ｨｦｨｪ06. 00ｨｨ01 Tolypocladium inflatum 0106ｨｬｨｨｨｪｨｨ08ｨｮ02ｨｰ 09 ｨｪ02ｨｺ06ｨｰ ｨｰ ｨｪｨｨｨｺｨ ｨ ｨｰ (0108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008). 00ｨｬ0209ｨｰ ｨｨ01ｨ ｨｬｨｨ T. geodes ｨｨ Beauveria bassiana, ﾂｨ 09ｨｰ04ｨｬｨｨ 09 01ｨ ｨｪｨｪ ｨ ｨｴ0209ｨｰ09ｨ 01, T. inflatum ｨｰ0907 ｨｰｨｨ07ｨｨ ﾂｨｪ04ｨｬ 09ｨ ｨｰ ｨｬ, ﾂｨ 09ｨｰ ｨｬ04ｨｬ ｨｨ ﾂ ｨｪ ｨ 05ｨ 09ｨｰ02ｨｦ. 0309ｨｰｨｹ 01ｨ ｨｪｨｪ ｨｨ010806ｨｰ ｨ ｨｪｨｰｨｪ0609ｨｰｨｨ 09ｨｨ ｨ Tolypocladium (Bisset, 1982) ｨｬ02 ｨｰ060006, ﾂｨｨ090502ｨｪｨｪ ｨｨ0104 ｨｨ ｨｰｨｪ04 ｨｺｨ ｨｺ 05ｨｪｨｰ06ｨｬ0607ｨ ｨｰ060002ｨｪｨｪ0402, ｨｪ02ｨｺ06ｨｰ ｨｨ ｨｹ04ｨｮ06ｨｰ ｨ ｨｨ06ｨｺ06ｨｪｨｰ T. inflatum ｨｮｨｺｨ ｨ 02ｨｰ ｨ ｨｪ0609ｨｰｨｹ ｨｺ ｨ ｨｺｨｰｨｨ09ｨｪ06ｨｬｨｮ 07ｨ 08ｨ 04ｨｨｨｰｨｨ04ｨｬｨｮ ｨｪｨ 05ｨｨ ﾂｨｨｨｪｨｺｨ 01 ｨｺ06ｨｬｨ ｨｨ 09 05ｨｰｨｨｨｬ 0601ｨｪ06ｨｦ ｨｨ ｨｬ0603ｨｪ ｨｨ ﾂｨｨｨｪ 06ｨｰｨｬ02 ﾂｨ 02ｨｬ06ｨｦ 04ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ06ｨｦ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ 09ｨｨ ｨ Tolypocladium, ｨ ｨｰｨ ｨｺ0302 B. bassiana 09 ｨｰ ｨｪｨｨｨｺｨ 01 ｨｬ060302ｨｰ ｨ 04ｨｰｨｹ ｨｺｨ ｨｰｨｪ0609ｨｰｨｹ ｨ ｨｪ06 ﾂｨｪ ｨｨ0906ｨｰｨｪ0401, 09 ｨｰ06ｨｬ ﾂｨｨ ｨｨ ﾂｨｨｨｪ06ｨｺ ｨｺ06ｨｬｨ , ｨｪｨ ｨｺ06ｨｰ ｨｬ0600ｨｮｨｰ 08ｨ 0409ｨｨ09ｨ ｨｰｨｹ ｨ ｨｪｨｪ ｨｨ ｨｨｨ 0609 (09ｨｮｨ ｨｪ0609ｨ, 2005). 00ｨｨ0104 Mucor hiemalis, Lecanicillium muscarium, Umbelopsis ramanniana. 06ｨ 04 ﾂｨｪ ｨｪ ｨｨ09ｨｰ ﾂ09, 09 ｨｰ06ｨｬ ﾂｨｨ ｨｺ080209ｨｰｨｪ0609ｨｰ0701 ｨｰ ｨｪｨｨｨｺ0609 (00ｨｨ08 ﾂｨｨｨｪｨｺ, 1976; 09ｨ ｨ ｨｹ0209ｨ ｨｨ 0108., 1989; 09ｨｮｨ ｨｪ0609ｨ, 2005; 0108ｨｮｨｬ ｨｨｨｬｨ ｨｦ0506, 09ｨｨ05ｨ ｨｪ02ｨｪｨｺ06, 2008). 491

8 ｨｰｨｪ06, ﾂｨｰ06 09ｨｨ01 Mucor hiemalis 0909ｨｰ0802 ﾂｨ 02ｨｰ ｨ ｨｬ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｪ0401 ｨｬ0209ｨｰｨ 01 06ｨ ｨｨｨｰｨ ｨｪｨｨ07 ｨｨ ｨｰｨｨ07ｨ ﾂ09. 02ｨｪ ｨｰｨ ｨｺ ｨ ｨｨｨｰｨ 02ｨｰ ｨｨ09ｨｰ ﾂ09ｨ 01 ｨｨ 09 05ｨｺ09ｨｰ0802ｨｬｨ 05ｨｹｨｪ0401 ｨｮ ｨｨ0701, 09 ｨｰ06ｨｬ ﾂｨｨ ｨｰ ｨｪｨｨｨｺｨ 01, ｨ 02ｨｪ ｨｺ ｨｰｨｮ 09 03ｨｨ0806ｨｺ06ｨｬ 01ｨｨｨ 07ｨ 0406ｨｪ ｨｨ ｨｰ ｨ ｨｪｨｰ02ｨｪ ｨｺ ｨ ｨｪｨ ｨ ｨｪ04ｨｬ ｨｮ ｨｨ07ｨｬ ｨｬｨｪ0600ｨｨ01 08ｨ ｨ 06ｨｰｨ 01 06ｨｪ ｨｮ0706ｨｬｨｨｨｪｨ 02ｨｰ0907 ｨｺｨ ｨｺ 06ｨ 04 ﾂｨｪ04ｨｦ 09ｨｨ ｨｰ ｨｪｨｨｨｺ0609, ｨ ｨｪ04ｨｦ ｨｺ ｨｰ08ｨｮｨｺ02ｨｨｨｨ 0106ｨｬｨｨｨｪｨｨ08ｨｮ06ｨｴ ｨｨ01ｨ ｨｬ ｨ ｨｰ Sphagnum fuscum (Domsch et al., 1980; Gross, Robbins, 2000; Thormann et al., 2000, 2002, 2004; Domsch et al., 2007). 00ｨｨ01 Umbelopsis ramanniana ｪ 0601ｨｨｨｪ ｨｨ04 ｨｰｨｨ07ｨｨ ﾂｨｪ0401 ｨｬｨｨｨｺ0806ｨｬｨｨ0202ｨｰ ﾂ ｨｦｨｪ04ｨｬｨｨ ｨ ｨｬｨｨ ｨｰｨ 0203ｨｪ06ｨｦ 0406ｨｪ04, 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ04ｨｦ ｨｨｨｪ01ｨｨｨｺｨ ｨｰ0608 ｨｪｨ ｨｨ09ｨｰ04ｨｦ ｨｰｨｨ ﾂ090606ｨ 08ｨ ｨ ｨｪｨｨ07, ｨｰｨ ｨｺ ｨ 04 ﾂ02ｨｪ ｨｰ ｨｪｨｨｨｺ ｨ ｨｰ (00ｨｨ08 ﾂｨｨｨｪｨｺ, 1976; 09ｨ ｨ ｨｹ0209ｨ, 0502ｨｪ0609ｨ, 1989) ｨｹ0407 ｨｪ02 06ｨｰｨｬ02ｨｰｨｨｨｰｨｹ ｨｪｨ 05ｨｨ ﾂｨｨ02 09 ｨｰ ｨｪｨｨｨｺｨ 01 ｨ 0605ｨｹ ｨｺ0605ｨｨ ﾂ0209ｨｰ09ｨ 09ｨｰ0208ｨｨ05ｨｹｨｪ ｨｬｨｨ020205ｨｨ07. 06ｨｰ06ｨｰ 00ｨ ｨｺｨｰ ｨｨｨｰ 06 ｨｰ06ｨｬ, ﾂｨｰ ｨｰ ｨ 08ｨ ｨ ｨｪｨｨ07 ｨｮ ﾂｨ 09ｨｰ09ｨｮ06ｨｰ 09ｨ ｨｬ ｨ 04ｨｪ ｨｮ ｨｨｨ 0609, ｨｨ0102ｨｪｨｰｨｨ00ｨｨｨｺｨ 02ｨｨ ｨ ｨｺ06ｨｰ ｨｬ0603ｨｪｨ ｨｬ06ｨｴｨｹ06 ｨｬ060502ｨｺｨｮ050708ｨｪ ｨｪ02ｨｰｨｨ ﾂ0209ｨｺｨｨ01 ｨｬ02ｨｰ ｨｹ ｨｰ ｨｪｨｨｨｺ ｨ ｨ 05ｨｹｨｪ ｨ 01 ｨ ｨｨ ｨｨ ｨ ｨｺｨｰｨｮｨ 05ｨｹｨｪ0609ｨｰｨｹ ｨｨ ｨ ｨｪｨｨｨｦ ｨ ｨｰｨｪ ｨｺ0609ｨｨ09ｨｰ02ｨｬ 0209ｨｪ0609ｨｪｨ ｨｹ ｨ ｨｰｨｪ ｨｺ0609ｨｨ09ｨｰ02ｨｬ ｨ ｨ 05ｨｹｨｪ ｨ 01 ｨ ｨｨ ｪ 05ｨｰ06 ｨｨ01 ｨｮ ﾂｨ 09ｨｰｨｨ02 09 ｨｺ08ｨｮ ｨｰ02 ｨｮ ｨ ｨｰｨ ｨｪｨ ｨｺｨ 0705ｨｨ09ｨ 06ｨｰ ｨｮ ｨｨ04 ｨ ｨｰｨｬ ｨｰ ｨｪ ｨ ｨ 01 ｨｨ 0601ｨｪ ｨｬ02ｨｪｨｪ ｨｰ 07ｨ 08ｨｪｨｨｨｺ ｨ ｨｪ04ｨｦ ｨ ｨ 05ｨ ｨｪ ｨ 06ｨｦ 05ｨｺ0609ｨｨ09ｨｰ02ｨｬ04 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ04ｨｮ02ｨｰ ﾂｨｨ09ｨｰ04ｨｦ ｨｮ ｨｪ04ｨｦ 0706ｨｰ06ｨｺ, ｨｰｨ ｨｴｨｨｨｦ 0906ｨ 06ｨｦ 09ｨｮｨｬｨｬｨｮ 09ｨ ｨｦ ｨｨ ﾂｨｪ06ｨｦ ｨｮｨｺ02ｨｨｨｨ ｨｨ 09ｨ ｨ ｨｪｨｨ07 ｨｺｨ ｨｺ 08ｨ 04ｨｪ06ｨｪｨ 0708ｨ ｨｪｨｪ ｨｰ06ｨｺ ｨ 02ｨｪｨｪ0609ｨｰｨｹ ｨ ｨｰ 06ｨ ｨｮ ｨｪｨ ｨｪ0204ｨ ｨｬｨｺｨｪｨｮｨｰ0609ｨｰｨｹ06 ｨｺ08ｨｮ ｨｰｨ 0902ｨｴ0209ｨｰ09, ｨｺ060001ｨ 09ｨｨ09ｨｰ02ｨｬ ｨ ｨｴｨ 06ｨｰ 09 ｨ ｨｨ ｨｮ ｨｬ02ｨｪｨｹ ｨｴ0209ｨｰ09, ﾂ02ｨｬ 04ｨ ｨ ｨｨ08ｨ 06ｨｰ ｨｨ04 ｨｪ ｨｬｨ ｨｰｨｨ ﾂｨｪ06 ｨ ｨ 05ｨ ｨｪ09 ｨｮ ｨ 09 ｨ ｨｰｨｪ ｨｺ0609ｨｨ09ｨｰ02ｨｬｨ 01 ｨｬ0603ｨｪ ｨｰｨ 09ｨｨｨｰｨｹ 09 09ｨｨ ｨｨ09ｨｰ02ｨｬ04 ｨ 0506ｨｺ0609 ｨｨ 0706ｨｰ06ｨｺ ｨｺｨ ｨｬｨｨ ｨｮ ｨ ｨｰ0907: 04ｨ 07ｨ 09 ｨｮ ｨ 09 09ｨｨ ｨｨｨｰ06ｨｬｨ , 09 ｨｰ ｨｪ06ｨｦ 04ｨ ｨｨ, ｨ ｨｰｨｪ06ｨｦ , ｨ ｨｰｨｬ ｨｰ06ｨｺｨｨ, ｨ 06ｨｴｨｨ02 05ｨｰｨｨ ｨ 0506ｨｺｨｨ, ｨ 03ｨ 06ｨｰ ｨｰ0609ｨｨｨｪｨｰ02ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ ｨｮｨｺ02ｨｨ02ｨｦ ｨｮ ｨ ( ﾂｨｨ09ｨｰｨ ｨｨ ﾂｨｪｨ ｨｮｨｺ02ｨｨ07), 05ｨｬｨｨ0909ｨｨ02ｨｦ ｨｮ000502ｨｺｨｨ ｨ 04ｨ ｨｨ ｨｬ02ｨｰｨ ｨｪｨ, 0904ｨｪ060906ｨｬ ｨｮ ｨ 09 ｨ ｨｰｨｪ04ｨｬｨｨ ｨ ｨｬｨｨ, 08ｨ ｨｪｨｨ02ｨｬ ｨｰｨｨ05ｨｺｨｨ ｨｨ ｨｰ060800ｨ, ｨ ｨｪｨｨ02ｨｬ ｨｺ0608ｨｪ02ｨｦ, ｨ ｨｰｨ ｨｺ ｨｬ ｨｮ ｨ ｨｨ ｨｪ ｨ 0506ｨｺｨ ｨｮ0006ｨｦ ｨｰ0906ｨｬ 06ｨｰｨｬｨｨ08ｨ ｨｪｨｨ07 ﾂｨ 09ｨｰ02ｨｦ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ, ｨ ｨｪｨｨ07 09ｨ ｨｬｨｨ01 08ｨ 09ｨｰ02ｨｪｨｨｨｦ, ｨｪ ｨｰ ｨｪｨｨ07 ｨｰ060800ｨ ｨｰ08ｨ ｨｪ ｨｨ05ｨｨ09ｨｹ ｨｨ ｨ ｨｪｨｨ ｨｪ0609ｨｪ0401 ｨ 0506ｨｺ0609 ｨｨ 0706ｨｰ06ｨｺ0609 ｨｮ ｨ ｨｪｨ 0605ｨｨ0006ｨｰ080600ｨｪ06ｨｬ ｨ ｨｰ02 ( ｨ 02ｨｺｨ 07, 2005), ｨｨ04ｨｮ ﾂｨ 05ｨｨ09ｨｹ ｨ 02ｨｪｨｪ0609ｨｰｨｨ 08ｨ ｨｪｨｨ ｨ 0408ｨｮ03ｨ 06ｨｴｨｨ01 ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ ｨｨ0006ｨｰ080600ｨｪ0401 ｨｰ ｨｪｨｨｨｺｨ 01 ( ｨ, 2005), ｨ ｨｰｨ ｨｺ ｨｨ07ｨｪｨｨ ｨｪｨｨ07 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0804 ｨｪｨ 08ｨ 0405ｨ 00ｨ 06ｨｴｨｮ06 ｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ0609 ｨｨ, ｨｺｨ ｨｺ ｨｰ09ｨｨ02, ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ02 09ｨｺ ｨｰｨｨ ｨｰｨｮ070502ｨｪｨｨ07 ｨｮ000502ｨｺｨｨ ｨ 04ｨ 09 ｨ ｨｰｨｬ ｨｮ (Thormann et al., 2002ｨ ) ｨ ｨｪｨｨｨｨ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ ｨｨ ｨｮ ﾂ02ｨｰ06ｨｬ ｨｰｨｨ07ｨ 04ｨ ｨｨ 06ｨｰｨｬ02 ﾂ02ｨｪｨ 06ｨ ｨｴｨ 07 04ｨ ｨｺ06ｨｪ06ｨｬ0208ｨｪ0609ｨｰｨｹ: ｨｪｨ ｨｨｨｬ02ｨｪｨｹ03ｨ 07 ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｹ ｨ 0408ｨｮ03ｨ 06ｨｴｨｨ01 ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ ｨ 08ｨ ｨｺｨｰ0208ｨｪｨ ｨ ｨｨ ｨｪｨｨ04ｨｨｨｪｨｪ ｨｰｨｨ07ｨ. 05ｨ ｨｹ ｨｰｨｨ07ｨ 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ04ｨｮ02ｨｰ0907 ｨｪｨ ｨｨｨ 0605ｨｹ03ｨｨｨｬ ｨ ｨｪｨｨ02ｨｬ 01ｨ ｨｪｨｪ06ｨｦ 0008ｨｮ ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ ｨｰｨｪ06, 05ｨｰ06 ｨｰｨ ｨｺ ｨ ｨｪ06 09 ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨ02ｨｦ ｨｮ ｨ ｨｴｨｨ ｨｴ0209ｨｰ09. 05ｨ ｨｹ ｨｪｨｨ04ｨｨｨｪｨｪ ｨｰｨｨ07ｨ ｨｺ06ｨｦ 09ｨｰ020702ｨｪｨｹ06 08ｨ ｨｪｨｨ ｨｨｨｰ ｨｬｨｨｨｪｨｨｨｬｨ 05ｨｹｨｪ0602 ｨｺ0605ｨｨ ﾂ0209ｨｰ ｨｺ0600ｨｨ ｨｨ04ｨｮ02ｨｬ ｨｴ0209ｨｰ09, ﾂｨｰ ｨｨ090601ｨｨｨｰ ｨｺ ｨｪ0204ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ06ｨｬｨｮ 08ｨ 0409ｨｨｨｰｨｨ ｨｨｨｰｨｨｨｺ ｨ 09ｨｪ02ｨｪｨｨ06 09 ｨｰ ｨｪ06ｨｦ 04ｨ ｨｹ ｨｰｨｨ07ｨ. 04ｨ ｨｪｨｨ ｨ 0408ｨｮ03ｨ 06ｨｴｨｨ01 ｨｬｨｨｨｺ ｨ ｨｪｨｨ04ｨｬ ｨ 06ｨｰ ｨｰｨ 0906ｨｬ ｨ ｨｪｨｨ ﾂ0209ｨｺ ｨｴ0209ｨｰ09ｨ ｨｰ ｨｪ06ｨｦ 04ｨ ｨｨ ｨｨ ｨｮ080609ｨｪ02ｨｬ ｨ ｨｰｨｪ (Thormann et al., 2002ｨ ; ｨ, 2005). 06ｨ ｨｺ ｨｺｨ 04ｨ ｨｪ06, ﾂｨｰ06 ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪ0602 ｨｮ ｨｪ0602 ｨｪｨ ｨｺ ｨｪｨｨ ｨｨ ｨｨｨｰ ｨｪｨ 06ｨｺ08ｨ ｨｨｨｪ02 ｨ ｨｰｨｪ ｨｬｨ 0909ｨｨ09ｨ ( ｨｺｨｨｨｦ 0807ｨｬ), ｨ ｨｬｨｨｨｪｨｨｨｬｨ 05ｨｹｨｪ0602 ｪ 09 06ｨｰｨｺ0804ｨｰ06ｨｦ ｨｰ0607ｨｨ ｨｺ0602 ｨｪｨ ｨｺ ｨｪｨｨ02 ｨｮ ｨ ｨｪｨ ｨｺ06ｨｬ 0807ｨｬ ｨｨ ｨｨｨｰ 04ｨ 09 ﾂ02ｨｰ ｨｺ06ｨｦ ｨｮｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｨ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨｨ ｨｪｨ 05ｨｨ ﾂｨｨ ｨｪ ｨｮ09ｨ, ｨｰ. 02. ｨｪｨ ｨｺ ｨｪｨｨ02 ｨｮ ｨ ｨｨ ｨｨｨｰ ｨｪ02 ｨｰ0605ｨｹｨｺ