гомелюк.p65

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1 ｨｰｨｨ ｨｬ ｨC ｨC ｨｬ020506ｨｺ ｨ 08ｨｰｨ ｨｬ02ｨｪｨｰ 0708ｨｨ080601ｨｪ ｨｮ , 06ｨｺ08ｨｮ03ｨ 06ｨｴ02ｨｦ ｨｨ ｨｨ09ｨｺｨｮ0909ｨｰ09, 0308ｨ 09ｨｨｨｰ0205ｨｹ09ｨｰ ｨｪ06ｨｦ ｨｨｨｰ0608ｨｨｨｨ, ｨｰ08ｨ 05ｨｨ ｨｪｨ 07 01ｨｨ09ｨｰｨ ｨｪ02ｨｨ06ｨｪｨｪｨ 07 09ｨｨ ｨｨ09ｨｰ02ｨｬｨ ｨｨｨｬｨ ｨｪｨｺ06ｨｦ ｨ 0405ｨ ｨｨ ｨｹ040609ｨ ｨｪｨ ｨ 09ｨｪ02ｨｪｨｨ07 09ｨｨ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨ07 ｨｨ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ 0804ｨ 09 ｨｰ ｨｬ0205ｨｺ ｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ 01 ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ ( ｨｺ06ｨｦ 07ｨ 08ｨｺ 0806ｨ ｨｮ0800, ｨｪｨ ｨｨｨｰ0608ｨｨ07, ｨｰ08ｨ 05ｨｨ07). 0508ｨ 09ｨｪｨｨ09ｨ 05ｨｨ09ｨｹ ﾂｨ ｨｪｨ 07 ｨ ｨ ｨｪｨｺｨ, ｨｬ0208ｨｰ0904ｨｦ ｨｺ0608ｨ ｨｦ 08ｨｨ00 ｨｨ 09ｨｺｨ 05ｨｨ09ｨｰ04ｨｦ 08ｨｨ ｨ 04ｨｪ04ｨｬｨｨ 0706ｨｺｨ 04ｨ ｨｰ020507ｨｬｨｨ ﾂ02ｨｪｨｪ0609ｨｰｨｨ ｨｹ0200ｨ 01ｨｪｨ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨｨｪｨｨ080609ｨ 05ｨｨ ｨｰｨ 09ｨｨｨｰ0205ｨｨ 0902ｨｬ. Carangidae, ｨｰｨ ｨｨ02 35 % ｨ, 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪｨｪ ｨｬ07 09ｨｨ010206ｨｮ ﾂ02ｨｰ ｨ ｨｬ0602 ｨｪｨｨ04ｨｺ0602 ﾂｨｨ ｨｨ (28) ｨ ｨ ｨｪｨ 08ｨｮ0302ｨｪ06 ｨｪｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺ02, 09ｨ ｨｬ ｨｺ0602 (42) ｪ ｨｪｨ 09ｨｺｨ 05ｨｹｨｪ06ｨｬ 08ｨｨ0002, ｨ ｨｬ0203ｨｮｨｰ06 ﾂｨｪ0602 (38) ｪ ｨｪｨ ｨｬ0208ｨｰ0906ｨｬ ｨｺ0608ｨ ｨｬ 08ｨｨ ｨ ｨｨｨ 0605ｨｹ ﾂｨｨ ｨ ｨ ｨｮ ﾂｨｰ02ｨｪ06 ｨｪｨ ｨｬ0208ｨｰ0906ｨｬ ｨｺ0608ｨ ｨｬ 08ｨｨ0002 (554), ｨｬ02ｨｪｨｹ0302 ｪ ｨｪｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺ02 (283) ｨｨ 09ｨｺｨ 05ｨｹｨｪ06ｨｬ 08ｨｨ0002 (278) ｨ ｨｴ0209ｨｰ09ｨ 0804ｨ 06ｨｰ ﾂ02ｨｰ05ｨｨ ｨｰ05ｨｨ ﾂｨ 05ｨｨ09ｨｹ 0108ｨｮ00 06ｨｰ 0108ｨｮ00ｨ ｨｪｨ ｨｺ06ｨｬ ｨｮ080609ｨｪ02 04ｨｪｨ ﾂｨｨｨｬ0609ｨｰｨｨ, ｨｪ06 ｨｪ0209ｨｺ0605ｨｹｨｺ ｨｺ080409ｨ 05ｨｨ09ｨｹ ｨｺ ｨｺ080409ｨ ｨｪｨｨ02 ｨ ｨｪｨ ｨｦ0102ｨｪ06 ｨｬ020301ｨｮ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺｨｨ ｨｨ 09ｨｺｨ 05ｨｨ09ｨｰ ｨｨ00ｨ ｨｮ02ｨｬ ｨ ｨｴ0209ｨｰ09ｨ 04ｨ ｨｬ02ｨｰｨｪ06 06ｨｰ05ｨｨ ﾂｨ 05ｨｨ09ｨｹ 06ｨｰ ｨｰｨｨ07ｨｨ ﾂｨｪ ｨ ｨｪ ｨ ｨｴ0209ｨｰ09 ｨｺ0608ｨ ｨｨ ｨｰ08ｨｨ ｨ ｨｴ0209ｨｰ09ｨ ｨ 05ｨｨ 09ｨｨ0104, 06ｨ ｨｨｨｰｨ 06ｨｴｨｨ02 ｨｪｨ ｨｺ0608ｨ 0505ｨ 01, 0601ｨｪｨ ｨｺ06 05ｨｰｨ 00ｨ ｨｮｨｪｨ ｨ 0405ｨ ｨ 0201ｨｪ0202, ﾂ02ｨｬ ｨｰｨ, ﾂｨｰ06 ｨｪｨ ｨ ｨ 05ｨ 09ｨｹ ｨｪｨ ｨｺ0608ｨ ｨｨ00ｨ ｨｺｨｮｨｰ060906ｨｦ ﾂｨ 09ｨｰｨｨ 04ｨ 05ｨｨ09ｨ, ｨ ｨ ｨｴ0209ｨｰ09ｨ 0804ｨ 04ｨｪｨ ﾂｨｨｨｬ06 06ｨｰ05ｨｨ ﾂｨ 05ｨｨ09ｨｹ 06ｨｰ ｨｰｨｨ07ｨｨ ﾂｨｪ0401 ｨｺ0608ｨ ｨｨ01ｨｰｨｨ060202ｨｪ ｨｨ 0601ｨｪ06 ｨｨ04 ｨｨ04ｨｮ ﾂ02ｨｪｨｪ ｨ ｨｴ0209ｨｰ09 ｨｪ02 ｨ ｨｲ ｨｪ ｨｬ060002ｨｪｨｪ06ｨｦ 0008ｨｮ070706ｨｦ 09ｨｨ010609, 0708ｨｨ090704ｨ ｨｪｨｪ0401 ｨｨ09ｨｺ0506 ﾂｨｨｨｰ0205ｨｹｨｪ06 ｨｺ 0601ｨｪ06ｨｬｨｮ ｨｬ0209ｨｰ0606ｨ ｨｨｨｰｨ ｨｪｨｨ ｨ ｨｪｨｪ ｨｨ00ｨ ｨｬｨｨ 09ｨｨ0104 ｨｨｨｬ0205ｨｨ ｨ 0605ｨｹ03ｨｮ ｨｪｨ ｨｬ0208ｨｰ0906ｨｬ 08ｨｨ0002 ｨｨ ｨｪｨ 09ｨｺｨ 05ｨｹｨｪ06ｨｬ 08ｨｨ0002, ｨｪ0609ｨｰｨｹ ｨ 0405ｨ , 0601ｨｪｨ ｨｺ06 ｨｰｨ ｨｬ 0909ｨｰ0802 ﾂｨ 05ｨｨ09ｨｹ ｨｨ ｨｰｨｨ07ｨｨ ﾂｨｪ ｨ ｨｨｨｰｨ ｨｰ0205ｨｨ ｨｬ0700ｨｺｨｨ ｨｮｨｪｨｰ ｨｪ04ｨｬ 06ｨ 08ｨ 0406ｨｬ 0804ｨ 04, 06ｨ 04 ﾂｨｪ06 06ｨｰｨｪ0609ｨｨｨｬ0402 ｨｺ 0008ｨｮ ｰ08ｨｨ ｱ, ｨ 0405ｨｨ ｨｪｨ ｨｦ0102ｨｪ04 ｨｪｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺ02. 00ｨｨ ｨｨ0806ｨｺｨｨｨｬ 08ｨ ｨｰ08ｨ ｨｪ02ｨｪｨｨ02ｨｬ ｨｨ ｨｪｨｨ04ｨｺ06ｨｦ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ ﾂｨｰ02ｨｪｨｨｨｨ ｨ ｨｨ06ｨｰ ｨｰｨ ｨｨ 09ｨｮｨｴ0209ｨｰ0902ｨｪｨｪｨｮ06 ﾂｨ 09ｨｰｨｹ ｨ ｨｴ0209ｨｰ ｨｰ ｨ ｨｨ06ｨｰ0607ｨ ｨｨ ｨｨ09ｨｰ02ｨｬｨ 06 ﾂ02ｨｪｨｹ ｨｮ ｨｪ06 08ｨ ｨ 06ｨｰｨ 05ｨ 09 ｨ ｨｮ01ｨｰ02, ｨｪｨｨ ｨｪ0401 ｨｮ ﾂ02ｨｰ0609 ｨｪ ｨｬ0603ｨｪ ｨｨ05ｨｮ ｨｪｨｨ04ｨｺ06ｨｦ 09ｨｨ01ｨｨｨｬ0609ｨｰｨｨ ｨｦ, ｨｺ0806ｨｬ02 ｨｰ060006, 06ｨｪｨ ｨｰ ｨｨｨｰｨｹ ｨｮ ﾂ02ｨｰ04 ｨ 0204 ｨｨ04ｨｲ07ｨｰｨｨ ｨ 04 ｨｨ05ｨｨ 0108ｨｮ00ｨｨ01 00ｨｨ010806ｨ ｨｨ06ｨｪｨｰ0609 ｨｨ ｨｰ06ｨｬｨｮ ｨｬ060302ｨｰ ｨ 04ｨｰｨｹ ｨｨ ｨｹ040609ｨ ｨｪｨ ｨｬ02ｨｪｨｪ ｨｬ06ｨｪｨｨｨｰ0608ｨｨｨｪ00ｨ ｨｪｨ 0601ｨｪｨｨ01 ｨｨ ｨｰ ｨｰ0607ｨｪｨｪ0401 ｨｮ ﾂｨ 09ｨｰｨｺｨ 01. Gomeluk V.E. Study of fish communities in biotopes of the Arafura Sea shallows by means of baited remote underwater video system // Izv. TINRO. ｪ ｪ Vol ｪ P. 181ｨC200. Baited remote underwater video technique (BRUVS) was used to compare fish assemblages at three sites with different habitat characteristics: sandy bank (SB), rock reef (RR) and degraded coral reef (DCR) in Port Essington, Garig Gunak Barlu 181

2 13National Park, Northern Territory, Australia. Carangidae family dominated, representing 35 % of all recorded fishes. The highest species number was recorded at RR (42) followed by DCR (38) and SB (28). The highest total fish number was recorded at DCR (554) followed by SB (283) and RR (278). Fish assemblages from all three sites were clearly different at high confidence level, but still overlapping. The higher overlapping was found between sandy bank and rocky reef. Fish assemblages at all three studied sites contained some coral-associated species, however fish fauna was less rich comparing to coral reefs in the mouth of the bay and fish assemblages were noticeably different from typical coral fish communities. None of fish assemblages at studied sites were presented by the ｰpure ｱ, single habitat-associated group of species ｪ at all three sites the fish communities were a mixture of fishes with various habitat preferences. Reef-associated species had larger proportion at DCR and RR where habitat complexity was higher. Still, soft-bottom habitat fishes made substantial fractions in fish assemblages at both DCR and RR. Similarly, fishes usually identified as ｰreef associated ｱ were found in notable proportions on sandy bank. Fishes with wide distribution and low selectivity in habitat preferences comprised a significant part in fish assemblages at all studied sites. In this study BRUVS technique worked well in the area where diving visual surveys was impossible to implement because of high water turbidity. Another advantage of this method is non-impact nature of visual survey; they can be used in long-term monitoring of fish assemblages at reference sites ｨｪｨｨ02 08ｨ ｨｮｨｪｨ ｨｰ080607ｨｨ ﾂ0209ｨｺｨｨ01 ｨｬ060809ｨｺｨｨ ｨ ｨｺ ｨ 08ｨｺｨ 0806ｨ ｨｮ0800 (Cobourg Marine Park, Garig Gunak Barlu National Park) ｨ 0600ｨ ｨｰｨ, 08ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｪｨ ｨｨ 09ｨｺ0506 ﾂｨ 02ｨｰ ｨ 07 ｨ ｨｨ ｨ, 0708ｨｨｨｪｨ ｨ ｨｴｨｨ01 ｨｺ ｨｬ02ｨｦ09ｨｰ09ｨ ｨｬ (Larson, Williams, 1997). 00ｨｬ0209ｨｰ02 09 ｨｰ02ｨｬ ｨｨ ｨ ｨｪｨｨ ｨｰ0607ｨｪｨｨ07 04ｨ 07ｨ ｨｨ ｨｪｨｺｨｨ 0905ｨｨ07ｨｪｨｨ07 ｨｪｨ ｨｪｨｨ ｨ ｨｨｨｰ0205ｨｹ09ｨｺ ｨ ｨｰ09ｨ 09 ｨｬ060809ｨｺ06ｨｬ 07ｨ 08ｨｺ ｨ ｨｮ ｨｨ ｨｪ ｨｨ05ｨｨ09ｨｹ. 06ｨｰ06 06ｨ ｨｲ0709ｨｪ0702ｨｰ0907 ｨｨ ｨｪｨｨ04ｨｺoｨｦ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ ｨｰ02ｨｦ ｨｨ ｨｰ08ｨ ｨｮ ｨｨ ｨｨ000609, ｨｨ 06ｨｰ08ｨｨ02ｨ ｨｰ0205ｨｹｨｪ04ｨｬ 06ｨｰｨｪ060302ｨｪｨｨ02ｨｬ ｨ ｨ 0608ｨｨ0002ｨｪ0609, ｨｴｨｨ ｨｰ 0706 ｨｮ0708ｨ ｨｪｨｨ06 ｨｪｨ 02ｨｨ06ｨｪｨ 05ｨｹｨｪ04ｨｬ 07ｨ 08ｨｺ06ｨｬ, ｨｺ 0708ｨｨｨｬ02ｨｪ02ｨｪｨｨ06 ｨｨ01ｨｰｨｨ0602ｨｨ ｨｨ ｨ ｨｴ02 ｨｺ y 0804ｨ 04 ｨｪ ｨｮ0706ｨｰ0802ｨ 0502ｨｪｨｨ ｨｨｨｴｨｮ. Bｨｨ01ｨｨｨｬ0609ｨｰｨｹ ｨｦ 09 ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ 09 ｨｰ02 ﾂ02ｨｪｨｨ ｨ ｨｰｨ ｨｰ 1,5ｨC2,0 ｨｬ ｨｨ ｨｬ02ｨｪｨｹ ｨｰ06 06ｨ 09ｨｰ0607ｨｰ0205ｨｹ09ｨｰ0906 ｨｨ 0708ｨｨ09ｨｮｨｰ09ｨｰ09ｨｨ02 ｨｺ08ｨｮ07ｨｪ0401 ｨｺ0806ｨｺ0601ｨｨ Crocodilus porosus ｨ 06ｨｰ ｨｪ ｨｬ0603ｨｪ04ｨｬｨｨ ｨ 04ｨｪ ｨ ｨ ｨｪｨｨ ｨｮ0707ｨｨ ｨｺ 0804ｨ. C ｨｪｨｨ ｨｰｨ 0902 ｨｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ ｨ ｨｴ0209ｨｰ ｨ, ｨｪｨ ｨｴｨｨ01 ｨｺ0608ｨ ｨｨ 09ｨｺｨ 05ｨｹｨｪ ｨｨ0004 ｨｨ 0108ｨｮ00ｨｨ02 ｨ ｨｨ06ｨｰ ｨｨｨ ｨｪ06ｨｦ 0406ｨｪ02 ｨｬ060802ｨｦ, 06ｨｬ0409ｨ 06ｨｴｨｨ ｨｪｨｮ ｨｨｨｰ0608ｨｨ06, 02ｨｴ ｨｹｨｪ06 09ｨｺｨｮ01ｨｪ04, 0106ｨｰ07 ｨｮ ｨ 08ｨ ｨｪ 04ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ04ｨｦ ｨｬｨ ｨｰ0208ｨｨｨ ｨ ｨｴ0209ｨｰ09ｨ ｨｬ 0804ｨ ｨｰｨｮｨ 08ｨｨ0701, ｨｬｨ ｨｪ x 04ｨ x ｨｨ 0708ｨｨ050200ｨ 06ｨｴｨｨ01 ｨｺ ｨｪｨｨｨｬ 08ｨ ｨｦ06ｨｪｨ 01 (Blaber et al., 1985, 1989, 1990; Blaber, 1986; Robertson, Duke, 1987; Sheaves, 1995; Larson, 1999), 09 04ｨ x ｨｬ060809ｨｺｨｨ01 ｨｰ08ｨ 09 (Blaber et al., 1992) ｨｨ 06ｨｰｨｺ0804ｨｰ ｨ ｨｦ06ｨｪｨ x ｨｬ060802ｨｦ (Russel, Houston 1989; Blaber et al., 1990, 1994, 1995, 2005). 0209ｨｪ0609ｨｪ ｨｪｨｨｨｬｨ ｨｪｨｨ02 ｨｨ ｨ ｨｰ020502ｨｦ, 08ｨ ｨ 06ｨｰｨ 0903ｨｨ01 09 ｨｰ080607ｨｨ ﾂ0209ｨｺｨｨ01 ｨｬ , ｨ ｨｨ ﾂ02ｨｪ06 ｨｺ 00ｨ ｨｮｨｪ ｨ ｨｺ0608ｨ ｨｨ (Randall, 1967; Luckhurst, Luckhurst, 1978; Gladfelter et al., 1980; Hixon, Beets, 1993; Sale et al., 1994; Sale 2006). 00 ｨｰ ｨｬ ｨ ｨｴ0209ｨｰ09ｨ 0804ｨ 09ｨｺｨ 05ｨｹｨｪ ｨｨ000609, 04ｨ ｨｪｨｨｨｬｨ 06ｨｴｨｨ01 04ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ ｨｴｨ 01ｨｨ ｨｨｨ ｨｪ06ｨｦ 0406ｨｪ02 ｨｰ080607ｨｨｨｺo09, ｨｰ0605ｨｹｨｺ ｨｪ ｨｬ07 09ｨｰｨ 05ｨｨ ｨｨｨｪｨｰ ｨ ｨｰｨｹ ｨｮ ﾂ02ｨｪ0401 (Falcon et al., 1996; Arburto-Oropeza, Balart, 2001; Ferreira et al., 2001; Dominici-Arosemena, Wolff, 2006). 01ｨｨ0806ｨｺｨｨ ｨｺｨｰｨｨ0904 ｨｨ ｨ ｨｪｨｨｨｦ 0804ｨ ｨｪ ｨ ｨｴ0209ｨｰ09 06ｨｰｨｺ080409ｨ 02ｨｰ ｨｨ ｨｹ040609ｨ ｨｪｨｨ02 08ｨ 0408ｨ ｨ 06ｨｰｨ ｨｪｨｪ ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ06 ｨｪ0201ｨ 09ｨｪ ｨｪ ｨｨ09ｨｰｨ ｨｪ02ｨｨ06ｨｪｨｪ ｨｨ ｨｨ09ｨｰ02ｨｬ ｨｨｨｬｨ ｨｪｨｺ06ｨｦ ( ) ｨｨ05ｨｨ, ｨｺｨ ｨｺ 06ｨｪｨｨ ｨｪｨ ｨ 02ｨｰ ｨ ｨｪ ﾂｨｪ06ｨｦ 05ｨｨｨｰ0208ｨ ｨｰｨｮ0802, baited remote underwater video systems (BRUVS) , ｨ ｨｴｨ ｨｪ ｨｰo00o ｨｬ02ｨｰ0601ｨ, ｨｰｨ : 0804ｨ 04, 0708ｨｨ ﾂ02ｨｪｨｪ04e 0708ｨｨｨｬｨ ｨｪｨｺ06ｨｦ, 09ｨｪｨｨｨｬｨ 06ｨｰ ｨｨ010206ｨｺｨ ｨｬ020806ｨｦ ｨ 0204 ｨｺｨ ｨｺ ｨｨｨ 06 ｨｮ ﾂｨ 09ｨｰｨｨ ｨ ｨｰ0608ｨ ｨｮ06ｨｴｨｨｨｬ ｨ ｨｪｨ 05ｨｨ0406ｨｬ ｨｮ ﾂ02ｨｪｨｪ ｨｨ ｨ 07ｨｨ0902ｨｦ 09 05ｨ ｨ 0608ｨ ｨｰ0608ｨｨｨｨ (Priede, Merret, 1996; Willis, Babcock, 2000; Cappo et al., 2004, 2006; Watson et al., 2005; Stobart et al., 2007). 182

3 ｨｮ0706ｨｬ07ｨｪｨｮｨｰ ｨ ｨｴ02ｨｪｨｨ0701 ｨｪ02 ｨ ｨｰ0608ｨｪ ｨｨ ｨ ｨｺ, ｨｰ ｨｰ06ｨｦ ｨ ｨｬ06ｨｦ ｨｰ06 ﾂｨｺｨｨ, ｨｰｨ ｨｺ ｨｺｨ ｨｺ 05ｨｰ06 ｨｪ ｨｰ02ｨｺｨ 0506 ｨｨ04 04ｨ 01ｨ ﾂ ｨｨ ｨ ｨｪｨｨｨｦ; 05ｨｨｨ ｨｨ09ｨ ｨｪｨｨ ｨ ｨｪ ｨ ｨｴ0209ｨｰ09 06ｨ 03ｨｨ08ｨｪ ｨｨ06ｨｪｨ ｨｨ 0908ｨ 09ｨｪ02ｨｪｨｨ ｨ ｨｴ0209ｨｰ09 ｨｨ04 08ｨ 0405ｨｨ ﾂｨｪ ｨｨ06ｨｪ0609 ｨｬ020301ｨｮ 0906ｨ 06ｨｦ (Watson et al., 2005; Stobart et al., 2007), 05ｨｨｨ ｨ 09ｨｪ02ｨｪｨｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｨ ｨｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｨ 08ｨ 0405ｨｨ ﾂｨｪ0401 ｨｬ02ｨｰ ｨｮ ﾂ02ｨｰ0609 (Willis, Babcock, 2000; Cappo et al., 2004; Watson et al., 2005) ｨ 0502ｨｪｨｨ06, 0708ｨｨ 0607ｨｨ09ｨ ｨｪｨｨｨｨ 0804ｨ ｨｪ ｨｪｨ ｨｪｨｨ07 06ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ06 ｨｺ08ｨｮ07ｨｪ ｨｨ06ｨｪｨ 0904ｨ 08ｨ ｨｪｨｪ04ｨｦ ｨｮｨｴｨｨｨｬｨｨ ｨｨ ｨ ｨｰ020507ｨｬｨｨ ｪ 08ｨ ｨｪ0106ｨｬｨｨ04ｨｨ080609ｨ ｨｪｨｪ ｨ ｨｪｨｨ ｨ ｨｺ ｨ 01 06ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ06 ｨ 0605ｨｹ0306ｨｦ ｨｴｨ 01ｨｨ, ｨｪ ｨｰ ｨｴｨｨ ｨ ｨｪ06ｨｦ 09ｨｰｨ ｨｪ02ｨｨｨｨ ｪ ｨｪ ｨｰ ｨ ｨ ｨｪ ｨｨ09ｨ ｨｰｨｹ 0008ｨｮ0707ｨｨ080609ｨｺｨｨ 0804ｨ 09 ｨｬｨｨｨｺ0806- ｨｨ ｨｬ ｨｺｨ 0502 (Sale et al., 1994; Forrester et al., 2002), ｨ 01 06ｨｰ010205ｨｹｨｪ ｨｨ00ｨ, 0008ｨｮ ｨｪ02ｨ 0605ｨｹ03ｨｨ01 08ｨｨ ｨｪｨｨ07, ｨｮ ﾂ02ｨｪｨｪ ｨｬ ｨｺｨ 05e, 06 ﾂ02ｨｪｨｹ 09ｨ 03ｨｪ ｨｨ ｨｪｨｺ ｨｨ090704ｨ ｨｪｨｪ0609ｨｰｨｨ 06ｨｰ010205ｨｹｨｪ ｨｨ ｨｨ ｨ ｨｴ0209ｨｰ ｨ ｨｺ ｨｰ02ｨｬ ｨｨ05ｨｨ ｨｨｨｪ04ｨｬ ｨ ｨｨ06ｨｰ0607ｨ ｨｬ ｨｨ 08ｨ ｨｪｨｨ06 09 ｨｪｨｨ ｨ 01ｨ ﾂｨｨ ｨｪｨ 09ｨｰ0607ｨｴ ｨｨ ｨ ｨｪｨｨ ｨｨ05ｨｨ 0607ｨｨ09ｨ ｨｪｨｨ ｨｰｨ 09ｨ ｨｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ0804 ｨｨ 0908ｨ 09ｨｪ02ｨｪｨｨ ｨ ｨｪ ｨ ｨｴ0209ｨｰ09 09 ｨｰ ｨ 08ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ ｨｺ06 08ｨ 0405ｨｨ ﾂｨｪ04ｨｬｨｨ 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ09ｨｰｨｨｨｺｨ ｨｬｨｨ ｪ ｨｪｨ ﾂｨ ｨｪ06ｨｦ 06ｨｰｨｬ0205ｨｨ, 06ｨｺ08ｨｮ0302ｨｪｨｪ06ｨｦ ｨ ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ04ｨｬｨｨ 0005ｨｮｨ ｨｨｨｪｨ ｨｬｨｨ, ｨｪｨ ｨｬ0208ｨｰ0906ｨｬ ｨｺ0608ｨ oｨｬ 08ｨｨ0002 ｨｨ ｨｪｨ 09ｨｺｨ 05ｨｨ09ｨｰ06ｨｬ 08ｨｨ ｨｹ040609ｨ ｨｪｨｨ02 ｨｬｨｪ060006ｨｺ08ｨ ｨｰｨｪ ｨｨ ｨｲ02ｨｬ06ｨｺ 0601ｨｪ ｨｨ ｨｰ ｨｮ ﾂｨ 09ｨｰｨｺｨ ｨｹ06 ｨｮｨｰ06 ﾂｨｪ02ｨｪｨｨ ｨｪ ﾂｨｨ0905ｨ 09ｨｨ ｨｲ02ｨｬ06ｨｺ ｨｪ0206ｨ ｨｨｨｬ ｨｨ09ｨ ｨｪｨｨ07 09ｨｨ ｨ 04ｨｪ0606ｨ 08ｨ 04ｨｨ ｨ ｨｴ0209ｨｰ ｨ ｨｨ ｨｪｨｺｨｨ ｨ ｨｪ0609ｨｰｨｨ 0708ｨｨｨｬ02ｨｪ02ｨｪｨｪ ｨｬ02ｨｰ0601ｨ ｨｮ050609ｨｨｨｰｨｹ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ % 06ｨ ｨｴ02ｨｦ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ 0804ｨ 09 01ｨ ｨｪｨｪ06ｨｬ ｨ ｨｴ0209ｨｰ ｨｬ06ｨｴｨｪ0609ｨｰｨｹ06 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺ ｨｺ08ｨｨｨｰ0208ｨｨ ｨｪｨｨｨｨ 08ｨ 0405ｨｨ ﾂｨｨｨｦ ﾝ 0,8. 00ｨ ｨｰ0208ｨｨｨ 0504 ｨｨ ｨｬ02ｨｰ ｨ ｨｪｨｨ ｨｨ050609ｨｹ 09 ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ, 08ｨ ｨｪｨｪ06ｨｦ ｨｪｨｰ08ｨ 05ｨｹｨｪ06ｨｦ ﾂｨ 09ｨｰｨｨ ｨ 0806ｨ ｨｮ0800, ｨｬ ｨ 00ｨｮ08ｨ, ｨｰ08ｨ 05ｨｨ07 (08ｨｨ09. 1), 09 06ｨｺｨｰ07ｨ 0802 ｪ ｨｬｨ 08ｨｰ ｨｨ 0802ｨｺ06ｨｬ02ｨｪ010609ｨ ｨｪｨｪ ｨ ｨ ｨｬ-0506ｨ ｨｨｨｰ020507ｨｬ ｨｮ ﾂｨ 09ｨｰｨｺｨ ｨ 0405ｨｨ 0904ｨ 08ｨ ｨｪ04 ｨｨ04 07ｨｮｨ 05ｨｨｨｺｨ 02ｨｨｨｦ 09 03ｨｮ08ｨｪｨ 0502 "03ｨｮｨｰ ｨｨｨｰ0205ｨｹ 0804ｨ ｨｪ06ｨｦ ﾂｨ 09ｨｰｨｨ ｨｰ08ｨ 05ｨｨｨｨ" (North Australian Fish Finder Magazine), 09 ｨｺ06ｨｰ060806ｨｬ 0708ｨｨ ｨｰ ｨ 00ｨｨ- ﾂ0209ｨｺｨｨ02 ｨｺ ｨｨｨｪｨ ｨｰ04 09 ｨｨ ｨｹ040609ｨ ｨｪｨｨ02ｨｬ 0907ｨｮｨｰｨｪｨｨｨｺ060906ｨｦ ｨｪｨ 09ｨｨ00ｨ 02ｨｨ06ｨｪｨｪ06ｨｦ 09ｨｨ09ｨｰ02ｨｬ04 GPS ｨｺ0605ｨｹｨｺｨｮ 0904ｨ 0608 ｨ ｨｨ06ｨｰ ｨ 0405 ｨｪ ｨｮ ﾂｨ 02ｨｪ, 09ｨｰaｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨｨｦ ｨ ｨｪｨ 05ｨｨ04 ｨｪｨ 03ｨｨ ｨｮ05ｨｹｨｰｨ ｨｰ ｨｨ080609ｨ 05 ｨｰｨ ｨｺ ｨｪｨ ｨ 02ｨｬ04ｨｬｨｨ "00ｨｨｨｺ09ｨｨ080609ｨ ｨｪｨｪ04ｨｬｨｨ 00ｨ ｨｺｨｰ0608ｨ ｨｬｨｨ" (Quinn, Keough, 2002) ｪ ｨｺ06ｨｪｨｺ0802ｨｰｨｪ04ｨｬｨｨ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ 0804ｨ ｨ 08ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ ｨ ｨｪｨｪ06ｨｬ ｨｨ ｨ ｨｪｨｨｨｨ ｨｰ0605ｨｹｨｺ06 06ｨｪｨｨ ｨｰｨ ｨｨ ｨｨｨｪｨｰ , ｨｨ ｨｮ05ｨｹｨｰｨ ｨｰ04 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺ ｨ ｨｪｨ 05ｨｨ04ｨ ｨｨ 09 05ｨｰｨｨｨｬ ｨｪ02 ｨｬ0600ｨｮｨｰ ｨ 04ｨｰｨｹ 05ｨｺ09ｨｰ08ｨ ｨｨ080609ｨ ｨｪ04 ｨｪｨ 0108ｨｮ00ｨｨ02 ｨ ｨｨ06ｨｰ ｨｨ06ｨｰ ｪ ｨｬ0205ｨｺ ｨｪｨ ﾂｨ ｨｪｨ 07 ｨ ｨ ｨｪｨｺｨ ｨｴｨ 01ｨｹ06 06ｨｺ ｨｬ 2 (ｨｺ09ｨ 0108ｨ ｨｰｨｮ08ｨｪ04ｨｦ 0708ｨｨ05ｨｨ09), 06ｨｺ08ｨｮ0302ｨｪｨｪｨ ｨｮｨ ｨｨｨｪｨ ｨｬｨｨ 10ｨC12 ｨｬ. 0206ｨｪｨｪ ｨ 01ｨｺｨｨ ｨｰｨ ｨｪ04 09ｨｨ05ｨｹｨｪ06 04ｨ ｨｨ0502ｨｪｨｪ04ｨｬ ｨｺ06ｨｬ 09 06ｨｰ010205ｨｹｨｪ04ｨｬｨｨ ｨｺｨｮ09ｨｺｨ ｨｬｨｨ ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ 08ｨ ｨｺｨｮ0302ｨｦ. 09ｨｨ06ｨｰ ｪ ｨｺ08ｨ 0209ｨ ｨｪｨ ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ00ｨ, 06ｨ ｨｴｨ ｨｴｨ 01ｨｹ ｨｺ06ｨｰ ｪ ｨｬ 2. 02ｨｪ06 09 ｨ 06ｨｺ09ｨｨｨｰ060904ｨｬｨｨ 09ｨｺｨ 05ｨ ｨｬｨｨ ｨｨ ｨｬｨ 0909ｨｨ09ｨｪ04ｨｬｨｨ ｨｬ0208ｨｰ0904ｨｬｨｨ ｨｺ0608ｨ ｨｬｨｨ ｨｺ060506ｨｪｨｨ07ｨｬｨｨ, 0706ｨｺ0804ｨｰ06 00ｨｨ010806ｨｨ01ｨ ｨｬｨｨ, 00ｨｮｨ ｨｺｨ ｨｬｨｨ ｨｨ 06ｨｰ010205ｨｹｨｪ04ｨｬｨｨ 03ｨｨ0904ｨｬｨｨ ｨｺ060506ｨｪｨｨ07ｨｬｨｨ ｨｺ0608ｨ , 0106ｨｪｨｪ ｨ 01ｨｺｨｨ ｪ 04ｨ ｨｨ0502ｨｪｨｪ04ｨｦ ｨｺ 09 ｨｺｨｮ09ｨｺｨ ｨｬｨｨ ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ06ｨｰ ｪ 09ｨｺｨ 05ｨｹｨｪ04ｨｦ 08ｨｨ ｨｬ 06ｨｰ 09ｨｺｨ 05ｨｨ09ｨｰ ｨ ｨｹ07. 01ｨ 01ｨｪ ｨｰ0802- ﾂｨ 05ｨｨ09ｨｹ ｨｺｨ 05, 0706ｨｺ0804ｨｰ ｨｨｨ 02ｨｪｨｰ060906ｨｬ, 06ｨｺ08ｨｮ0302ｨｪｨｪ0402 ｨｮ ﾂｨ 09ｨｰｨｺｨ ｨｬｨｨ 04ｨ ｨｨ0502ｨｪｨｪ ｨｺｨ, 09 06ｨｰ010205ｨｹｨｪ04ｨｬｨｨ 0507ｨｨｨ ｨｨ06ｨｪｨｰｨ ｨｬｨｨ ｨｨ ｨｪ04ｨｬｨｨ ｨｨ ｨ ｨｮ0804ｨｬｨｨ ｨｬｨｨ ｨ ｨｪ ｨｨ09ｨ ｨｪｨｨ ｨ 0405ｨｨ ﾂｨｪ0401 ｨｺ06ｨｪ09ｨｰ08ｨｮｨｺ02ｨｨｨｦ 0708ｨｨ090601ｨｨｨｰ ｨｬ ｨ 09ｨｰ (Willis, Babcock, 2000; Cappo et al., 2004, 2006). 02ｨ ｨｴｨｨｨｦ 0708ｨｨｨｪ02ｨｨ07 ｨｮ09ｨｰ0806ｨｦ09ｨｰ09ｨ 0601ｨｨｨｪｨ ｨｺ0609: 09ｨｨ010206ｨｺｨ ｨｬ0208ｨ ｨｰ06ｨｬ (ｨ ｨ ｨｪ0609 ｨｮ0708ｨ ｨｪｨｨ07) ｨｬ02ｨｰｨｨ ﾂｨｪ06ｨｬ ｨ 06ｨｺ0902, ｨｨ040006ｨｰ ｨｪｨｪ06ｨｬ ｨｨ04 ｨｰ060509ｨｰ0609ｨｰ02ｨｪｨｪ06ｨｦ ｨｨ ｨｨｨｪｨｨ ｨｦ ｨｰ08ｨｮｨ 04, 09 ｨ ｨｺ08ｨｨ ｨｬｨｨ ｨｪ02ｨｦ ｨｨ 04ｨ 01ｨｪ02ｨｦ ｨｺ080403ｨｺｨ ｨｬｨｨ, 0706ｨｬ02ｨｴｨ 02ｨｰ ｨｨｨ 05ｨｹｨｪｨｮ06 ｨｬ02ｨｰｨ 0505ｨｨ ﾂ0209ｨｺｨｮ ｨ 08ｨｪｨｮ06 08ｨ ｨｬｨｮ, 04ｨ ｨｴｨｨｨｴｨ 06ｨｴｨｮ06 ｨ 06ｨｺ09 ｨｨ 09ｨｨ010206ｨｺｨ ｨｬ0208ｨｮ 06ｨｰ ｨｬ0201ｨ ｨｪｨｨ ﾂ0209ｨｺｨｨ ｨｪｨｨｨｦ. 04ｨ ｨｬｨ 0609ｨｪｨ ｨｴ02ｨｪｨ 0705ｨ 09ｨｰｨｨｨｺ060904ｨｬ ｨｰ06ｨｬ 0105ｨｨｨｪ06ｨｦ 1,5ｨC2,0 ｨｬ, ｨｺ06ｨｰ060804ｨｦ ｨｪ020902ｨｰ ｨｺ06ｨｪｨｰ02ｨｦｨｪ ｨｨｨｬｨ ｨｪｨｺ06ｨｦ (0,5ｨC1,0 ｨｺ00 ｨｨ04ｨｬ0205ｨｹ ﾂ02ｨｪｨｪ06ｨｦ 09ｨ 0801ｨｨｨｪ04 Sardinops neopilchardus) ｨ ｨｬ02 183

4 13ｨｺ080207ｨｨｨｰ0907 ｨｺ080207ｨｺｨｨｨｦ ｨｰ ｨｮｨｬ07 ｨ ｨｮ07ｨｬｨｨ ｨｨ 0005ｨ 0006ｨｬ ｨｮ ﾂ ｨｰ0409ｨｺｨ ｨｪｨｨ07 ｨｨ ｨｲ02ｨｬｨ ｨｪｨ ｨｪ0609ｨｰｨｹ ｨ ｨｪｨｨ07 09ｨｨ ｨｲ02ｨｬｨｺｨｨ. 04ｨｨ ｨ 00ｨｨ ﾂ0209ｨｺ ｨｪｨｨ02 09ｨｰｨ ｨｪ02ｨｨｨｦ 09 ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ, ｨｨ ｨｹ040609ｨ 05ｨ 09ｨｹ , 01ｨ 02ｨｨ06ｨｪｨ 05ｨｹｨｪ04ｨｦ 07ｨ 08ｨｺ 01ｨ 08ｨｨ00 01ｨｮｨｪｨ ｨｺ 09ｨ 0805ｨｮ, ｨ ｨｮ0800, ｨｪｨ ｨｨｨｰ0608ｨｨ07, ｨｰ08ｨ 05ｨｨ ｨｪ02ｨｪｨｨ07 09 ｨｰ02ｨｺ09ｨｰ02 Fig. 1. Geographical locations of baited remote underwater video system (BRUVS) study sites in Port Essington, Garig Gunak Barlu National Park, Cobourg Peninsula, Northern Territory, Australia. Comments are presented in the text ｨｨ ｨ 07ｨｨ09ｨｨ ｨｨ ｨｹ040609ｨ 05ｨｨ09ｨｹ 09ｨｨ010206ｨｺｨ ｨｬ Sony6ｾ4 Digital 8 Handycam, ｨｬ ｨｹ TRV250E, 0609ｨｪｨ ｨｴ02ｨｪｨｪ ｨｨ0806ｨｺ060006ｨｺｨｮ09ｨｪ04ｨｬｨｨ 06ｨ ｨｲ02ｨｺｨｰｨｨ09ｨ ｨｬｨｨ VCL-0637H. 02ｨｪｨｨ 08ｨ ｨ 06ｨｰｨ 05ｨｨ 09 ｨ 09ｨｰ06ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｬ ｨｨｨｬ02 05ｨｺ ｨｨ02ｨｨｨｨ, 0006ｨｺｨｮ09ｨｪ ｨ 0909ｨｰ0607ｨｪｨｨ02 ｨ ｨｮ09ｨｰｨ ｨｪ ｨｪ06 ｨｪｨ ｨｨｨｬ "ｨ 0209ｨｺ06ｨｪ02 ﾂｨｪ0609ｨｰｨｹ ｪ 08ｨｮ ﾂｨｪ0602 ｨｮ0708ｨ ｨｪｨｨ02" 09 04ｨ 07ｨｨ09ｨｹ06 ﾂｨｨ0905ｨ ｨｨ ｨｬ02ｨｪｨｨ 09ｨｮｨｰ06ｨｺ ｨｪｨ ｨｪｨｺｨｮ ｨｨ ｨｲ02ｨｬｨｺｨｨ ｨｨ05ｨｨ09ｨｹ ｨｰ0605ｨｹｨｺ ｨｪ0209ｨｪ ｨｬ07 (ｨｬ020301ｨｮ 8 ｨｨ 18 ﾂｨ 09) ｨｬ07 ｨｺ09ｨ 0108ｨ ｨｰｨｮ08ｨｪ ｨｨ05ｨｨ090609, ﾂｨｰ06ｨ 04 ｨｨ04ｨ 0203ｨ ｨｰｨｹ 09ｨｨ05ｨｹｨｪ ｨｨ05ｨｨ09ｨｪ0401 ｨｰ02 ﾂ02ｨｪｨｨｨｦ ｨｨ 0706ｨｪｨｨ0302ｨｪｨｪ06ｨｦ 09ｨｨ01ｨｨｨｬ0609ｨｰｨｨ ｨｦ ｨｰｨ ｨｰｨｨ09ｨｪ04ｨｦ 0907ｨｮｨｰｨｪｨｨｨｺ060904ｨｦ ｨｪｨ 09ｨｨ00ｨ ｨｰ0608 ｰGPSmap76 ｱ 0708ｨｨｨｬ02ｨｪ ｨ ｨｪｨ 0601ｨｨｨｪ ｨｨ ｨｰ06ｨｰ 0302 ｨｮ ﾂｨ 09ｨｰ06ｨｺ ｨｨ ｨ ｨｪｨｨｨｦ 09 ｨｺｨ ｨｬ ｨ ｨｨ06ｨｰ ｨ 09ｨｨ00ｨ 02ｨｨ06ｨｪｨｪｨ ｨｪ0609ｨｰｨｹ ｨｰｨ 09ｨｨ05ｨ ﾀ 5ｨC7 ｨｬ ｨｨｨ 04ｨｰｨｨ07 ｨｪｨ ｨｰ06 ﾂｨｺｨｮ 0601ｨｨｨｪ ｨｨ ｨｰｨ 0905ｨｨ09ｨ ｨｺ 09ｨｲ02ｨｬｨｺ02: 09 09ｨｨ010206ｨｺｨ ｨｬ0208ｨｮ 0909ｨｰｨ ｨ 09ｨｹ ﾂｨｨ09ｨｰｨ 07 ｨｺｨ ｨｰｨ ｨｪｨｺ06ｨｦ, ｨｺ06ｨｪｨｰ02ｨｦｨｪ ｨ ｨ ｨｨｨｬｨ ｨｪｨｺ06ｨｦ ｨｬ06ｨｴｨｹ ｨｺｨ ｨｰ0608ｨ Humminbird 323x sounder ｨｨ09ｨｹ 0005ｨｮｨ ｨｨｨｪｨ ｨｨ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨ ｨｮ ｨｪ0609ｨｰｨｨ, 09 01ｨｪ0209ｨｪｨｨｨｺ 04ｨ 07ｨｨ090409ｨ ｨｹ ｨｬ0209ｨｰ06 ｨｨ ｨｬ07 ｨｪｨ ﾂｨ 05ｨ 09ｨｨ ｨｲ02ｨｬｨｺｨｨ

5 ｨｮ09ｨｰｨ ｨｪ0609ｨｺｨｨ 09ｨｨ010206ｨｺｨ ｨｬ ｨｨｨｬ ｰ04ｨ 07ｨｨ09ｨｹ-ｨｪｨ 0006ｨｰ ｱ ｨｺ080403ｨｺｨ ｨ 06ｨｺ09ｨ ｨｺｨ ｨｬ ｨｬ02ｨｰｨｨ ﾂｨｪ06 04ｨ ｨｺ080409ｨ 05ｨ 09ｨｹ, ｨｪ ｨｰ0902ｨｪｨｪ ｨ ｨｬ ｨｪｨ 01ｨｪ06 09ｨｨ010206ｨｺｨ ｨｬ0208ｨｮ ｨｨ05ｨｨ ｨｨｨｬ 09ｨｨ ｨ 07ｨｨ09ｨｨ ｨｬ06ｨｴｨｹ06 07ｨｮ05ｨｹｨｰｨ 01ｨｨ09ｨｰｨ ｨｪ02ｨｨ06ｨｪｨｪ ｨｺ06ｨｪｨｰ ｨｰｨ 09ｨ ｨｪｨ 01ｨｪ02 09 ｨｰ02 ﾂ02ｨｪｨｨ02 60 ｨｬｨｨｨｪ ｨｺｨ ｨｮ010601ｨｨ05ｨ ｨｪｨ 0108ｨｮ0006ｨｦ ｨｨ0006ｨｪ, ｨｮ09ｨｰｨ ｨｪｨ 0905ｨｨ09ｨ ｨｮ0006ｨｦ , ｨ 04ｨ ｨｰ02ｨｬ ｨ ｨｴｨ 05ｨ 09ｨｹ, ﾂｨｰ06ｨ ｨｪ07ｨｰｨｹ ｨｦ ｨ 0707ｨ 08ｨ ｨｰ, ｨｮ ｨ ｨｺ06ｨｪ ﾂｨｨ0903ｨｨｨｦ 09ｨｨ ｨ 07ｨｨ09ｨｹ. 0905ｨ ｨ ｨｨｨｬ02ｨｪ02ｨｪｨｨ ｨｺ06ｨｰ06 ﾂｨｪ ｨｪｨ 09ｨｨ00ｨ ｨｰ0608ｨ, ｨｪ02ｨ 0605ｨｹ0306ｨｦ 0005ｨｮｨ ｨｨｨｪ02 ｨｨ 06ｨｰ09ｨｮｨｰ09ｨｰ09ｨｨ06 ｨｰ02 ﾂ02ｨｪｨｨｨｦ ｨｨ ｨｲ02ｨｬｨｺｨｨ 09 ｨｺｨ ｨｬ ｨｨ04 ｨ ｨｨ06ｨｰ ｨ 0405ｨｨ ｨｨ ｨｪ ｨ 01 06ｨｺ08ｨｮ03ｨｪ0609ｨｰｨｨ 09 08ｨ 01ｨｨｨｮ0906ｨｬ, ｨｪ ｨ 06ｨｴｨｨｨｬ 10 ｨｬ ｨｬ06ｨｰ0807 ｨｪｨ ｨｮ09ｨｨ05ｨｨ07 ｨｨ04ｨ 0203ｨ ｨｰｨｹ ｨｪｨｨ07 09ｨｨ ｨｲ02ｨｬｨｺｨｨ 0708ｨｨ ｨｦ 09ｨｨ01ｨｨｨｬ0609ｨｰｨｨ, ｨｪｨｨｨｰｨｹ 09ｨｨ01ｨｨｨｬ0609ｨｰｨｹ ｨｮ 01ｨｪｨ ｨｪ ｨ ｨ ｨｬ0603ｨｪ06, ｨｨ 0708ｨｨｨｬ0208ｨｪ06 25 % 06ｨｰ09ｨｪ07ｨｰ ｨｬｨ ｨｰ0208ｨｨｨ 05ｨ ｨ ｨｪ020708ｨｨ000601ｨｪ ｨ ｨｪｨ 05ｨｨ04ｨ ｨｮ05ｨｹｨｰｨ ｨｰ02 06ｨｰｨ 0608ｨ ｨｬ04 08ｨ ｨ 00ｨ 05ｨｨ 9 60-ｨｬｨｨｨｪｨｮｨｰｨｪ04ｨｬｨｨ 09ｨｨ010206ｨｺｨ ｨｰｨ ｨｬｨｨ, 06ｨｰ09ｨｪ07ｨｰ04ｨｬｨｨ ｨｪｨ ｨ ｨｨ06ｨｰ , 15 ｨｺｨ ｨｰｨ ｨｬｨｨ ｪ ｨｪｨ ｨ ｨｨ06ｨｰ ｨｨ 14 ｪ ｨｪｨ ｨ ｨｨ06ｨｰ ｨｨ ｨ 07ｨｨ09ｨｨ 06ｨ 08ｨ ｨ ｨ ｨｰ0409ｨ 05ｨｨ09ｨｹ ｨ ｨｪ ｨｨ09ｨ ｨｪｨｪ06ｨｦ ｨｬ02ｨｰ0601ｨｨｨｺ02 (Willis, Babcock, 2000; Cappo et al., 2004; Watson et al., 2005) ｨｬｨｨｨｪ 09ｨｨ ｨ 07ｨｨ09ｨｨ ｨｬｨ ｨｰ08ｨｨ09ｨ 05ｨｨ09ｨｹ, ﾂｨｰ06ｨ 04 06ｨｰｨｬ02ｨｰｨｨｨｰｨｹ ﾂｨｨ ｨ 02ｨｦ ｨｺｨ ｨｨ01ｨ, ｨｨ0903ｨｨ ｨｪ ｨｰ ﾂｨｰ06ｨ 04 ｨｨ04ｨ 0203ｨ ｨｰｨｹ ｨｦｨｪ ｨｮ ﾂ02ｨｰｨ 0601ｨｪｨｨ01 ｨｨ ｨｰ ｨ 02ｨｦ, 0708ｨｨ ｨｮ06ｨｴ02ｨｬ ｨ ｨｪｨ 05ｨｨ0402 ｨｨ ｨｹ040609ｨ 05ｨｨ09ｨｹ ｨｰ0605ｨｹｨｺ06 01ｨ ｨｪｨｪ ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪ06ｨｬ ﾂｨｨ ｨ 02ｨｦ ｨｺｨ ｨｨ01ｨ, ｨｪｨ ｨｨ0903ｨｨ ｨｪ ｨｬ02ｨｪｨｪ ｨｪｨｨ07 ｨｺｨ ｨｬ020804, ｪ ｰMaxN ｱ, 0706ｨｺｨ 04ｨ ｨｰ020507, 03ｨｨ0806ｨｺ ｨｨｨｬ02ｨｪ0706ｨｴ ｨｨ ｨｨ ｨ ｨｪｨｨ ｨｬ06ｨｴｨｹ (Willis et al., 2000; Cappo et al., 2004). 09ｨ 08ｨ ｨｺｨｰ0208ｨｨ09ｨｰｨｨｨｺｨｨ 0106ｨｪｨｪ ｨｺ0804ｨｰｨｨ07 ｨ ｨｪｨ 05ｨｨ04ｨｨ080609ｨ 05ｨｨ09ｨｹ 07ｨｮｨｰ02ｨｬ ｨｨ04ｨｮ ﾂ02ｨｪｨｨ07 09ｨｪｨｨｨｬｨｺ ｨｪｨ, ｨｮ ﾂ02ｨｪｨｪ ｨｨ ｨｬ06ｨｰ ｨｨ ｨ 07ｨｨ0902ｨｦ ﾂｨｨ090502ｨｪｨｨ ｨｨ 06ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ06ｨｦ ｨｴｨ 01ｨｨ 0706ｨｺ0804ｨｰｨｨ07 ｨｰｨ ｨｺｨｨ ｨｬ02ｨｪｨｰ ｨｪｨｪ ｨｮｨ 09ｨｰ08ｨ ｨｰｨ, ｨｺｨ ｨｺ 09ｨｺｨ 0504, 09ｨ 05ｨｮｨｪ04, ｨｺ08ｨｮ07ｨｪ0402 (> 25 09ｨｬ) 06ｨ 0506ｨｬｨｺｨｨ ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ ｨｬｨ ｨｺ ｨｨｨ 02ｨｪｨｰ0609 (00ｨｮｨ ｨｺｨｨ, ｨｺ060506ｨｪｨｨｨｨ ｨｺ0608ｨ , 00ｨｨ010806ｨｨ0104), ｨｺ06ｨｰ ｨｮ090205ｨｨ ﾂｨｨ09ｨ 06ｨｰ ｨｪ020601ｨｪ ｨｪ0609ｨｰｨｹ ｨｹ0200ｨ 01ｨｪｨ, ｨ 0405ｨ ｨｨ ｨｹ040609ｨ ｨｪｨ ｨ ｨｬｨｬｨ Image Pro Express. 0301ｨｨｨｪｨｨ0204 ｨｨ04ｨｬ020802ｨｪｨｨ ｨｴｨ 01ｨｨ 0706ｨｺ0804ｨｰｨｨ07 (%) ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨｨｬ ｨ ｨｪｨ 05ｨｨ0406ｨｬ ｨ 0405ｨｨ ｨ 08ｨ ｨ ｨｪ04 09 ｨｺ09ｨ 0108ｨ ｨｰｨｪ0402 ｨｺ0608ｨｪｨｨ ｨ 08ｨｺ09ｨｨｨｪｨｮ ｨｮ ﾂ02ｨｪｨｪ ｨ ｨｪｨｪ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0802 ｨｪｨ ｨｪ0609ｨｰｨｨ , 0005ｨｮｨ ｨｨｨｪ02 ｨｨ ｨｪ020601ｨｪ ｨｪ0609ｨｰｨｨ 0106ｨｪｨｪ ｨ ｨｪ0103ｨ 00ｨｰｨ 0908ｨ 09ｨｪｨｨ09ｨ 05ｨｨ09ｨｹ ｨｬ06ｨｴｨｹ ｨｪ0600ｨ ｨｺｨｰ0608ｨｪ ｨｨ ｨｨ06ｨｪｨｪ ｨ ｨｪｨ 05ｨｨ04ｨ (ANOVA) 0708ｨｨ 0706ｨｬ06ｨｴｨｨ 07ｨ ｨｺ02ｨｰｨ 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ ｨ ｨｬｨｬ SYSTAT ｨ ｨｪｨｪ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ ｨｮ ﾂｨｰ02ｨｪｨｪ ｨｬ07 09ｨｲ02ｨｬ06ｨｺ 09ｨｨ ｨ 0405ｨｨ ｨｨ ｨｹ040609ｨ ｨｪ ｨｰｨ ｨｪｨｨ ｨｨ09ｨｺｨ 09ｨｨ ｨ 09 ｨｬ0209ｨｰｨ ｨｪｨｨ07 ｨｨ ｨ ｨｪｨｨｨｦ ｨｨ ｨ 09ｨｪ02ｨｪｨｨ ｨ ｨｴ0209ｨｰ ｨ, ｨｪｨ ｨｴｨｨ01 08ｨ 04ｨｪ0402 ｨ ｨｨ06ｨｰ ｨｰ0902ｨｰ09ｨｰ09ｨｨｨｨ 09 ｨｨ ｨｬｨｨ ﾂｨｰ02ｨｪｨｨ07ｨｬｨｨ. 06ｨｰｨｨ ｨ ｨｪｨｪ ｨｨｨｬ02ｨｪ0705ｨｨ09ｨｹ 0708ｨｨ 0904 ﾂｨｨ090502ｨｪｨｨｨｨ ｨｰｨ ｨｬ06ｨｴｨｪ0609ｨｰｨｨ 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺ ｨｺ08ｨｨｨｰ0208ｨｨ ｨｬ ｨｮ090205ｨｨ ﾂ02ｨｪｨｨ07 ﾂｨｨ0905ｨ 0904ｨ ｨｺ, ﾂｨｰ06ｨ ｨｨｨｰｨｹ ﾂｨｨ ｨｨ ｨ ｨｺ, ｨｰｨ ｨｰ06 ﾂｨｪ ｨｨ09ｨｰ08ｨ 02ｨｨｨｨ 25 % ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨｨｦ 09 06ｨ ｨｴ02ｨｬ ﾂｨｨ ｨ (05) ｨｪｨ ｨｮ080609ｨｪ02 04ｨｪｨ ﾂｨｨｨｬ0609ｨｰｨｨ ｦﾁ = 0,05 (Harvey et al., 2001) ｨｺ0605ｨｹｨｺｨｮ ｨｪｨ 05ｨｨ ﾂｨｨ ｨｺｨｨ01, ｨｬｨ 0506 ﾂｨｨ090502ｨｪｨｪ ｨｨ ｨ ｨｬ060302ｨｰ ｨｨ07ｨｰｨｹ ｨｪｨ ｨｮ05ｨｹｨｰｨ ｨｰ04 ｨ ｨｪｨ 05ｨｨ ｨｪｨｺ ｨｰ09ｨ 08ｨ 0405ｨｨ ﾂｨｪ ｨ ｨｴ0209ｨｰ09, 09 ｨ ｨｪｨ 05ｨｨ04ｨ 01 DIVERSE, SIMPER ｨｨ ANOSIM, 0802ｨ 05ｨｨ040609ｨ ｨｪｨｪ0401 ｨｪｨ 07ｨ ｨｺ02ｨｰ ｨｨｨｺ05ｨ 01ｨｪ ｨ ｨｬｨｬ PRIMER 6.0 (Clarke, Warwick, 2001), ｨ 0405ｨｨ ｨｨ ｨｹ040609ｨ ｨｪ04 ｨｰ0605ｨｹｨｺ06 01ｨ ｨｪｨｪ ｨｪｨ ｨｨｨ ｨ 04 ﾂｨｪ0401, ｨｬｨｪ ﾂｨｨ090502ｨｪｨｪ ｨｨ01ｨ 01 (ｨｰｨ ｨ 05. 1). 02ｨ ｨｪｨｪ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ 0804ｨ ｨｨ01 ｨｪ0608ｨｬｨ 05ｨｨ04ｨ 02ｨｨｨｨ ｨ 0405ｨｨ ｨｰ08ｨ ｨｪ ｨｬｨｨ080609ｨ ｨｪ ｨｨ 0706ｨｬ06ｨｴｨｨ ｨｰｨｨ07 ｨｺ0608ｨｪ07 ﾂ02ｨｰ090208ｨｰ06ｨｦ 09ｨｰ020702ｨｪｨｨ ｨ 09ｨｪ02ｨｪｨｨ07 ｨｨ ｨｮ02ｨｬ ｨ ｨｴ0209ｨｰ ｨ ｨ 0405 ｨｰｨ ｨｺ0302 ｨｨ ｨｹ040609ｨ ｨｪ ｨｺ ｨｨ02ｨｨ02ｨｪｨｰ ｨｰ09ｨ 04ｨ ｨｺｨｺｨ 08ｨ (Sj). 0308ｨｨ ﾂｨｨ090502ｨｪｨｨｨｨ ｨ 08ｨｪ ｨ 09ｨｪｨｨ09ｨ 05ｨｨ09ｨｹ ｨｮ05ｨｹｨｰｨ ｨｰ04 ｨｮ ﾂ02ｨｰ ｨ ｨｨ06ｨｰ0607ｨ ｨｨ 09, 08 ｨｨ 00, 09 ｨｨ ｨｬｨｮ0502: Sj = c*(a + b + c) ｨC1, a ｨｨ b ｪ ﾂｨｨ ｨｨ010609, 06ｨ ｨｪｨ 08ｨｮ0302ｨｪｨｪ0401 ｨｰ0605ｨｹｨｺ06 09 ｨ ｨｨ06ｨｰ ｨｨ05ｨｨ 09, c ｪ ﾂｨｨ ｨ ｨｴｨｨ ｨ 06ｨｨ01 ｨ ｨｨ06ｨｰ ｨｨ (Ludwig, Reynolds, 1988). 185

6 1306ｨ ｨ 05ｨｨ02ｨ ｨｬ02ｨｦ09ｨｰ09ｨ ｨｨ 09ｨｨ ｨ, ｨｮ ﾂｨｰ02ｨｪｨｪ ｨｬ07 09ｨｨ ｨｲ02ｨｬｨｺｨｨ 09 ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ, ｨｬ ｨ 00ｨｮ08ｨ, ｨｪｨ ｨｨｨｰ0608ｨｨ07, ｨｰ08ｨ 05ｨｨ ｨｪ04 ｨ ｨｬｨ , ｨｬｨｪ ﾂｨｨ090502ｨｪｨｪ ｨｨ0104 Table 1 Families, species and fish numbers recorded during BRUVS surveys at three study sites in Port Essington, Arafura Sea, the Northern Territory, Australia. More abundant and common species are given in bold 0502ｨｬ02ｨｦ09ｨｰ0906, 09ｨｨ01 ﾁｨｨ ｨｮ ﾂｨｰ02ｨｪｨｪ ｨ, 0209ｨｪ0609ｨｪ06ｨｦ 09ｨｨ01 ｨｺ0608ｨｬｨ 05ｨｺ04./08ｨ ｨｪ00 Carcharhinidae Carcharhinus melanopterus 8/ ｨ ｨ Carcharinhus sealai 6/ ｨ ｨ Negaprion acutidens 1/ ｨ ｨ Stegostomidae Stegostoma fasciatum 2/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Gingglymostomatidae Nebris ferrugineus 14/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Hemigaleidae Triaenodon obesus 1/ ｨ ｨ Dasyatidae Taeniura meyeni 5/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Taeniura sp. 2/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Dasyatis sp. 1/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Pastinachus sephen 1/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Myliobatidae Aetobatus narinari 1/ ｨｺｨｨ Rhinobatidae Rhinobatus typus 2/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Elopidae Elops hawaiensis 2/ ｨ ｨ Ariidae Arius thallasinus 7/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Serranidae Epinephelus coioides 12/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Plectropomus maculatus 2/ ｨ ｨ Cephalopholis boenak 1/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Apogonidae Apogon sp. 39/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Carangidae Gnathanodon speciosus 128/1 0902ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Caranx sexfasciatus 58/6 0404ｨ ｨ Pantolabus radiatus 56/7 0404ｨ ｨ Scomberoides commersonnianus 42/8 0404ｨ ｨ Alepes sp. 34/ ｨｺｨｰ06ｨｪｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Selar sp. 33/ ｨｺｨｰ06ｨｪｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Caranx ignobilis 18/ ｨ ｨ Alectis indicus 10/ ｨｪ06 ﾂｨｪ0402 Scomberoides tol 5/ ｨｺｨｰ06ｨｪｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Trachinotus blochii 3/ ｨｺｨｨ Caranx tille 1/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Ulua aurochs 1/ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Carangoides chrysophrys 1/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Eceneidae Remora sp. 11/20 01ｨ 08ｨｮ03ｨｪ ｨ 08ｨ 04ｨｨｨｰ04 186

7 1302ｨｺ06ｨｪ ﾂｨ ｨｪｨｨ02 ｨｰｨ ｨ Table 1 finished 0502ｨｬ02ｨｦ09ｨｰ0906, 09ｨｨ01 ﾁｨｨ ｨｮ ﾂｨｰ02ｨｪｨｪ ｨ, 0209ｨｪ0609ｨｪ06ｨｦ 09ｨｨ01 ｨｺ0608ｨｬｨ 05ｨｺ04./08ｨ ｨｪ00 Leiognathidae Leiognathus equulus 69/5 0902ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Caesionidae Caesio cunning 3/ ｨ ｨｪｨｺｨｰ06ｨｪ Lutjanidae Lutjanus erythropterus 41/9 0404ｨ ｨ, ｨ 02ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Lutjanus carponatus 31/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Lutjanus malabaricus 4/ ｨ ｨ, ｨ 02ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Lutianus argentomaculatus 3/ ｨ ｨ, ｨ 02ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Lutianus gibbus 1/ ｨ ｨ, ｨ ｨｪ06 ﾂｨｪ0402 Lutjanus russeli 1/ ｨ ｨ, ｨ ｨｪ06 ﾂｨｪ0402 Nemipteridae Pentapodus porosus 104/2 0902ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Scolopsis monogramma 26/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ Scolopsis sp. 8/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Haemulidae Plectorhinchus labiosum 19/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Plectorhinchus gibbosus 5/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Lethrinidae Lethrinus ornatus 72/4 0902ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Lethrinus laticaudis 34/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0401 Lethrinus nebulosus 7/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Lethrinus sp. 4/4 0902ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Mullidae Upeneus vittatus 2/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Ephippidae Platax teira 26/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Chaetodontidae Chelmon marginalis 11/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Parachaetodon ocellatus 7/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Heniochus acuminatus 5/27 05ｨｨ0107 ﾂｨｨ02 ｨ ｨｪ06 ﾂｨｪ0402 Pomacanthidae Chaetodontoplus duboulayi 2/30 05ｨｨ0107 ﾂｨｨ02 ｨ ｨｪ06 ﾂｨｪ0402 Sphyraenidae Sphyraena jello 30/ ｨ ｨ Sphyraena obtusata 2/ ｨ ｨ Labridae Choerodon cyanodus 7/ ｨｺｨｨ Acanthuridae Acanthurus grammoptilus 79/ ｨｨ Monocanthidae Monacanthus chinensis 2/ ｨｪ06 ﾂｨｪ0402 ｨｨ ｨｨ Tetraodontidae Lagocephalus lunaris 1/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402 Feroxodon multistriatus 1/ ｨｪｨｰ0609ｨｪ0402 ｨ ｨｪ06 ﾂｨｪ0402, 0804ｨ ｨ ｨｬ02ｨｦ09ｨｰ09 ｪ 28 00ｨｨ ｪ ｨ ｨｪｨｪ ｨ ｨｴ0209ｨｰ09ｨ ｨｬ 0804ｨ 09 ｨｨ ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ 01 04ｨ ｨｰ02ｨｬ 0908ｨ 09ｨｪｨｨ09ｨ 05ｨｨ09ｨｹ 09 06ｨ 03ｨｨ08ｨｪ04ｨｬｨｨ 05ｨｨｨｰ0208ｨ ｨｰｨｮ08ｨｪ04ｨｬｨｨ ｨｬｨ ｨｰ0208ｨｨｨ 05ｨ ｨｬｨｨ 06 08ｨ ｨｪｨｨｨｨ 0804ｨ ｨｪ06ｨｦ ｨｰ08ｨ 05ｨｨｨｨ ｨｨ 0708ｨｨ050200ｨ 06ｨｴｨｨ01 08ｨ ｨｦ06ｨｪｨ 01 06ｨｪ ｨ 02ｨｨ00ｨｨｨｺｨｨ (Allen, 1997; Blaber et al., 1985, 187

8 131989, 1990, 1992, 1994; Blaber, 1986; Robertson, Duke, 1987; Sainsbury, 1987; Russel, Houston, 1989; Russel, 1990; Kailola et al., 1993; Brewer et al., 1995; Sheaves, 1995; Larson, 1999) ｨｪｨｨ07 ｨｪｨ ｨｦ0102ｨｪｨｪ Fish Base (Froese, Pauly, 2002), ｨｰｨｮ07ｨｪ06ｨｦ 09 ｨｨｨｪｨｰ0208ｨｪ02ｨｰ02, ｨｨ ｨｮｨ 05ｨｨｨｺｨ 02ｨｨｨｦ (Parrish et al., 1986; Robertson, Duke, 1987; Parrish, 1989; Blaber et al., 1990; Kailola et al., 1993), ｨｨ ｨｹ040609ｨ 05ｨｨ09ｨｹ ｨｰｨｪ020902ｨｪｨｨ07 06ｨ ｨｪｨ 08ｨｮ0302ｨｪｨｪ ｨ ｨｪｨｪ06ｨｬ ｨｨ ｨ ｨｪｨｨｨｨ 09ｨｨ ｨ ｨｺ ｨｰ02ｨｬ ｨｨ05ｨｨ ｨｨｨｪ04ｨｬ 07ｨｨｨｴ020904ｨｬ 0008ｨｮ0707ｨ ｨｬ. 0805ｨ 0909ｨｨ00ｨｨｨｺｨ 02ｨｨ07 07ｨｨｨｴ ｨｮ ｨ 01ｨ 02ｨｰ ｨ ｨｪ01ｨ 05ｨｮ (Randal, 1967) ｨｮ05ｨｹｨｰｨ ｨｰ04 ｨｨ ｨｨ01 06ｨ 09ｨｮ030102ｨｪｨｨ ｨｪ0600ｨ ｨｺｨｰ0608ｨｪ04ｨｦ 01ｨｨ ｨｨ06ｨｪｨｪ04ｨｦ ｨ ｨｪｨ 05ｨｨ04 ANOVA ｨｨ ｨｨｨｰｨｹ ｨｪ02ｨｺ06ｨｰ ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ ｨｺ ｨｰ ｨｪ ｨ 0405ｨｨ ﾂｨｨ ｨｪ02ｨｦ 0005ｨｮｨ ｨｨｨｪ02 ｨｪｨ ｨｨ ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ 01 (F = 13,869, p < 0,001). 0105ｨｮｨ ｨｨｨｪ04 09 ｨ ｨｨ06ｨｰ ｨ 0405ｨｨ ｨｬ02ｨｪｨｹ ｨ 09ｨｪ02ｨｪｨｨ06 09 ｨ ｨｨ06ｨｰ0607ｨ ｨｬｨｨ 08 ｨｨ ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ 04ｨｪｨ ﾂｨｨｨｬ ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｬ020301ｨｮ ｨ ｨｨ06ｨｰ0607ｨ ｨｬｨｨ 08 ｨｨ 00 ｨｪ ｨｪ06, p = 0,541 (08ｨｨ09. 2). 0508ｨ 09ｨｪ02ｨｪｨｨ02 01ｨｪｨ ｨｰ ｨｨ ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ06ｨｦ ｨｴｨ 01ｨｨ ｨｨｨ ﾂ0209ｨｺｨｨ01 06ｨ ｨｲ02ｨｺｨｰ0609 (09ｨｺｨ 0504, 09ｨ 05ｨｮｨｪ04) ｨｨ ｨｬｨ ｨｺ ｨｨｨ 02ｨｪｨｰ0609ｨ, ｨｮ090205ｨｨ ﾂｨｨ09ｨ 06ｨｴｨｨ01 ｨｪ020601ｨｪ ｨｪ0609ｨｰｨｹ ｨｹ0200ｨ 01ｨｪｨ (habitat rugosity), ｨｪｨｪ ｨｨ ｨｹ040609ｨ ｨｪｨｨ02ｨｬ ｨ ｨｪｨ 05ｨｨ04ｨ ANOVA, 0706ｨｺｨ 04ｨ 0506, ﾂｨｰ06 ｨ ｨｨ06ｨｰ ｨｨｨｬ ｨ ｨｬｨｮ06 ｨｪｨｨ04ｨｺｨｮ ｨｪ0609ｨｰｨｹ ｨｹ0200ｨ, 09 ｨｰ ｨｬ07 ｨｺｨ ｨｺ 04ｨｪｨ ﾂ02ｨｪｨｨ07 05ｨｰ ｨｺｨ 04ｨ ｨｰ ｨ ｨｨ06ｨｰ0607ｨ 01 09, ｨｺ08ｨ ｨｦ 0406ｨｪ02 ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ00ｨ, ｨｨ 00, 09ｨｺｨ 05ｨｹｨｪ06ｨｬ 08ｨｨ0002, ｨ 0405ｨｨ ｨｪ04ｨｬｨｨ (08ｨｨ09. 2). 0105ｨｮｨ ｨｨｨｪｨ, ｨｬ 00 05ｨｰ020702ｨｪｨｹ ﾂ02ｨｪｨｪ0609ｨｰｨｨ 0706ｨｺ0804ｨｰｨｨ07, arcsin 0602ｨｬ070208ｨ ｨｰｨｮ08ｨ, = ｨ 09 04ｨｨ ｨｪ ｨｮｨ ｨｨｨｪｨ 09 ｨｨ ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ 01 (ｨ, 0708ｨｨ ｨｪ04 09ｨｰｨ ｨｪ01ｨ 08ｨｰｨｪ ｨｰｨｺ0506ｨｪ02ｨｪｨｨ ｨｪｨｨ01 ｨ 08ｨｨ00ｨｬ02ｨｰｨｨ ﾂ0209ｨｺｨｨ01); ｨｪ ｨｰ020702ｨｪｨｹ ﾂ02ｨｪｨｪ0609ｨｰｨｨ ｨｹ0200ｨ 01ｨｪｨ (ｨ, ｨ 0302ｨｪｨ 09 arcsin ｨｰ08ｨ ｨｪ ｨｬｨｨ080609ｨ ｨｪｨｪ ｨｪｨｰ ｨｴｨ 01ｨｨ 0706ｨｺ0804ｨｰｨｨ07 01ｨｪｨ ｨ ｨｪｨｨ04ｨｬｨ ｨｬｨｨ ｨｨ ｨｨ ﾂ0209ｨｺｨｨｨｬｨｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨ ｨｬｨｨ (ｨｬｨ ｨｺ ｨｬｨｨ, 00ｨｮｨ ｨｺｨ ｨｬｨｨ, 09ｨｺｨ 05ｨ ｨｬｨｨ, 09ｨ 05ｨｮｨｪｨ ｨｬｨｨ, 06ｨ 0506ｨｬｨｺｨ ｨｬｨｨ ｨｺ0608ｨ ), 0708ｨｨ ｨｪ04 09ｨｰｨ ｨｪ01ｨ 08ｨｰｨｪ ｨｰｨｺ0506ｨｪ02ｨｪｨｨ ｨｪｨｨ01 ｨ 08ｨｨ00ｨｬ02ｨｰｨｨ ﾂ0209ｨｺｨｨ01); ｨｪ ｨｪ0609ｨｰｨｪｨ 07 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨ (09, 0708ｨｨ ｨｪ04 04ｨｪｨ ﾂ02ｨｪｨｨ07 09ｨｰｨ ｨｪ01ｨ 08ｨｰｨｪ ｨｨｨ 06ｨｺ ｨｪｨｨ01 ｨ 08ｨｨ00ｨｬ02ｨｰｨｨ ﾂ0209ｨｺｨｨ01) 09 06ｨ ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ 01 ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ ｨｮ ﾂｨ ｨｰ ｨｪ0609ｨｰｨｨ 08ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｬ020301ｨｮ 09ｨｰ0605ｨ ｨｨｨｺｨ ｨｬｨｨ 0708ｨｨ ｨｰ ｨｪｨ ﾂ02ｨｪｨｨ07 ｨｮ080609ｨｪ02ｨｦ ｨｰ ｨｪ0609ｨｰｨｨ ｨｮ05ｨｹｨｰｨ ｨｰｨ ｨｬ MSDｨｰ0209ｨｰｨ Fig. 2. ｨ : mean depth values at three BRUVS survey sites at Port Essington, Cobourg Peninsula. Error bars are standard deviations. Significance level (one-way ANOVA, Tukey HSD test) is given for significantly different values; ｨ : bottom habitat rugosity expressed as percent bottom cover (arcsin transformed) of physical structures (rocks, boulders, coral rubble) and macroepibenthos. Error bars are standard errors; 09: mean surface water temperature. Error bars are standart errors. Significance level (Tukey post-hoc MSD test) is given for statistically different values. Vertical bars are standard errors 188

9 130602ｨｬ070208ｨ ｨｰｨｮ08ｨ ｨｪ0609ｨｰｨｪ ｨｨ0601 ｨｨ ｨ ｨｪｨｨｨｦ ｨｺ060502ｨ ｨ 05ｨ 09ｨｹ 06ｨｰ 27, , ｨｨ ｨｨ06ｨｪｨｪ04ｨｦ ｨ ｨｪｨ 05ｨｨ04 ANOVA ｨｪ02 06ｨ ｨｪｨ 08ｨｮ03ｨｨ05 04ｨｪｨ ﾂｨｨｨｬ ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｬ020301ｨｮ ｨｪ02ｨｦ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0806ｨｦ ｨｪｨ ｨ ｨｨ06ｨｰ0607ｨ ｨｨ ｨｰ ｨｬ07 09 ｨ ｨｨ06ｨｰ , 08ｨ ｨｪｨｪ06ｨｬ ｨ 05ｨｨ0302 ｨｺ ｨｮ ｨｨ04 ｨ ｨｮ01ｨｰ04, ｨｪｨｨ02 04ｨｪｨ ﾂ02ｨｪｨｨ07 ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0804 ｨ 0405ｨｨ 09ｨｮｨｴ0209ｨｰ0902ｨｪｨｪ06 ｨｪｨｨ0302 (08ｨｨ09. 2) ｨｬ07 09ｨｨ010206ｨｮ ﾂ02ｨｰ0609 ｨ ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪ ｨｨ01ｨ 0804ｨ, 06ｨｰｨｪ060907ｨｴｨｨ ｨｺ ｨｬ02ｨｦ09ｨｰ09ｨ ｨｬ ｨｰｨ 09ｨｨｨｰ0205ｨｨ 0902ｨｬ02ｨｦ09ｨｰ09ｨ 09ｨｰｨ 0908ｨｨ (Carangidae) 0106ｨｬｨｨｨｪｨｨ080609ｨ 05ｨｨ, ｨｰｨ ,0 %, 04ｨ ｨｰ02ｨｬ ｨ 05ｨｨ ｨｪｨｨｨｰ (Nemipteridae) ｪ 12,4 %, 0502ｨｰ08ｨｨｨｪｨｨ0104 (Lethrinidae) ｪ 10,5, 01ｨｨ08ｨｮ (Acanthuridae) ｪ 7,1, 05ｨｨ0600ｨｪｨ ｨｰ (Leiognathidae) ｪ 6,2, ｨ 0609ｨｰｨ 0903ｨｨ ｨｬ02ｨｦ09ｨｰ09ｨ ｨｪｨ 09 ﾂｨｨｨｰ0409ｨ 05ｨｨ 28,8 % ｨｮ ﾂｨｰ02ｨｪｨｪ ｨ (ｨｰｨ ｨ 05. 1) ｨｮ05ｨｹｨｰｨ ｨｰｨ ｨｬ ｨ ｨｪｨ 05ｨｨ04ｨ DIVERSE, 09 ｨ ｨｨ06ｨｰ , ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺ02, ｨ ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪ06 ｨｪｨ ｨｨｨｬ02ｨｪｨｹ ﾂｨｨ ｨｨ ｨ ｨ 09ｨｪ02ｨｪｨｨ06 09 ｨ ｨｨ06ｨｰ0607ｨ ｨｬｨｨ 09 ｨｨ 00, ｨｬ0208ｨｰ0904ｨｬ ｨｺ0608ｨ ｨｬ 08ｨｨ0006ｨｬ ｨｨ 09ｨｺｨ 05ｨｹｨｪ04ｨｬ 08ｨｨ0006ｨｬ (ｨｰｨ ｨ 05. 2). 0308ｨｨ 05ｨｰ06ｨｬ ｨｬ020301ｨｮ ｨ ｨｨ06ｨｰ0607ｨ ｨｬｨｨ ｨｪ02 ｨ ｨｪｨ ｨｦ0102ｨｪ06 09ｨｮｨｴ0209ｨｰ0902ｨｪｨｪ ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ01 08ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｪ02ｨｬｨｮ ﾂｨｨ0905ｨｮ 09ｨｨ ｨ, 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪｨｪ ｨｬ ｨｪ ｨｨ010206ｨｮ ﾂ02ｨｰｨ. 00 ｨｰ ｨｬ ｨｪ0202 ﾂｨｨ ｨ 02ｨｦ 08ｨ 04ｨｪ ｨｨ010609, 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪｨｪ ｨｬ ｨｪ ｨｨ010206ｨｮ ﾂ02ｨｰｨ, ｨ ｨｮｨｴ0209ｨｰ0902ｨｪｨｪ06 ｨｨ 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ ｨｰ ｨｪ06 ｨｪｨｨ ｨ ｨｨ06ｨｰ ｨ 09ｨｪ02ｨｪｨｨ06 09 ｨ ｨｨ06ｨｰ060706ｨｬ 09 (ｨｰｨ ｨ 05. 2). 02ｨ ｨｴ0202 ｨｨ ｨｪ0202 ﾂｨｨ ｨｨ ｨ ｨｨ ｨｪ0202 ﾂｨｨ ｨ 02ｨｦ, 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪｨｪ ｨｬ07 09ｨｨ010206ｨｮ ﾂ02ｨｰ ｨｰ ｨ ｨｨ06ｨｰ0607ｨ ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ, ｨｪｨ ｨｨｨｰ0608ｨｨ07, ｨｰ08ｨ 05ｨｨ07 Total and mean number of fish species and mean number of fish in one video survey in three habitats in Port Essington, Arafura Sea, Northern Territory, Australia ｨ ｨ 05ｨｨ02ｨ 2 Table 2 07 ﾂｨｰ02ｨｪ ｨｪ ｨｪ0202 ﾂｨｨ ｨｨ06- ﾁｨｨ ﾂｨｰ02ｨｪ ｨ, ﾂｨｨ ｨｨ p 0804ｨ ｨｪ06ｨｬ p ｨｰ0607 ｨｮ ﾂ02ｨｰ ｨｨ ｨｺ ｨｪ06ｨｬ ｨｮ ﾂ02ｨｰ02 ｨｮ ﾂ02ｨｰ02, 05ｨｺ04. A ,33 ﾀ 1,18 31,44 ﾀ 4,74 0,106* ,07 ﾀ 0,91 34,64 ﾀ 4,98** ns 0,027** ,21 ﾀ 0,94 19,79 ﾀ 3,48** 0308ｨｨｨｬ02 ﾂｨ ｨｪｨｨ ｨｨ ｨｰ ｨｪ ｨｪｨ ﾂ02ｨｪｨｨ02 ﾀ 09ｨｰｨ ｨｪ01ｨ 08ｨｰｨｪｨ ｨｨｨ ｨｺｨ. * 05ｨｮｨｴ0209ｨｰ0902ｨｪｨｪ ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ01 08ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｬ020301y ｨ ｨｨ06ｨｰ0607aｨｬｨｨ ｨｰ06ｨｬｨｮ 07oｨｺｨ 04ｨ ｨｰ ｨｪ02 06ｨ ｨｪｨ 08ｨｮ0302ｨｪ06 (F = = 2,393, p = 0,106, ANOVA). ** ｨｪ0202 ﾂｨｨ ｨ, 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪｨｪ ｨｪ06ｨｬ 09ｨｨ010206ｨｮ ﾂ02ｨｰ02 09 ｨ ｨｨ06ｨｰ , 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ 04ｨｪｨ ﾂｨｨｨｬ06 ｨｪｨｨ0302, ﾂ02ｨｬ 09 ｨ ｨｨ06ｨｰ (F = 4,024, p = 0,027). 01ｨｨ04ｨｺｨｨ02 04ｨｪｨ ﾂ02ｨｪｨｨ07 ｨｺ ｨｨ02ｨｨ02ｨｪｨｰｨ ｨｰ09ｨ 04ｨ ｨｺｨｺｨ 08ｨ 0708ｨｨ 0908ｨ 09ｨｪ02ｨｪｨｨｨｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺｨｨ ｨｨ ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ00ｨ (Sj = 0,10 ﾀ 0,02) 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ06ｨｰ 06 ｨｰ06ｨｬ, ﾂｨｰ06 ﾂｨｨ ｨ ｨｴｨｨ ｨｰｨｨ01 ｨ ｨｨ06ｨｰ ｨｨ ｨ ｨｪ ｨｨｨｺ ｨｪ0602 ｨｪｨｨ04ｨｺ ｨｪｨ ﾂ02ｨｪｨｨ02 05ｨｰ ｨｺ ｨｨ02ｨｨ02ｨｪｨｰｨ ｨ ｨｮ ﾂ02ｨｪ ｨｨ 0908ｨ 09ｨｪ02ｨｪｨｨｨｨ ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ00ｨ ｨｨ 09ｨｺｨ 05ｨｨ09ｨｰ ｨｨ00ｨ (Sj = 0,15 ﾀ 0,01). 00 ｨｰ ｨｬ ｨ 09ｨｪ02ｨｪｨｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺｨｨ ｨｨ 09ｨｺｨ 05ｨｨ09ｨｰ ｨｨ00ｨ ｨｪ020603ｨｨ01ｨ ｨｪｨｪ06 01ｨ ｨ ｨｬ ｨｺ ｨｪｨ ﾂ02ｨｪｨｨ02 ｨｺ ｨｨ02ｨｨ02ｨｪｨｰｨ (Sj = 0,20 ﾀ 0,06). 0308ｨｨ 05ｨｰ06ｨｬ 08ｨ 0405ｨｨ ﾂｨｨ ｨｨ ﾂｨｨｨｪｨ 01 ｨｺ ｨｨ02ｨｨ02ｨｪｨｰ ｨｰｨ ｨｰｨｨ09ｨｰｨｨ- ﾂ0209ｨｺｨｨ ｨｪ ｨｰ ｨｪ04 (ｨ ｨｪｨ 05ｨｨ04 ANOVA df = 2; F = 2,289; 08 = 0,123) ｨｨ ﾂｨｨｨｪｨ 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺ ｨｺｨ 04ｨ ｨｰ R 09 ｨ ｨｪｨ 05ｨｨ0402 ANOSIM 06ｨｰ08ｨ 03ｨ 02ｨｰ 09ｨｰ020702ｨｪｨｹ ｨｰ09ｨ ｨｨ 08ｨ 0405ｨｨ ﾂｨｨ ｨ ｨｴ0209ｨｰ ｨ ｨｨ ｨ 09ｨｪｨｨ09ｨ 02ｨｬ0401 ｨ ｨｨ06ｨｰ R = ｨｮ ﾂｨ 02, ｨｨ ｨ 0608ｨｺｨｨ ｨｨ ｨｪ ｨ ｨｨ06ｨｰ0607ｨ ｨｪ04 ｨｬ020301ｨｮ 0906ｨ 06ｨｦ ｨ , ﾂ02ｨｬ ｨ 06ｨｦ ｨｨ ｨ ｨｺ, ｨｮ ﾂ02ｨｪｨｪ ｨｮ0006ｨｬ ｨ ｨｨ06ｨｰ (070605ｨｪ0602 ｨｪ ｨｰ0906). 01ｨ ｨｰｨｨ09, ｨｨ ﾂｨｨｨｪｨ R ｨ 05ｨｨ04ｨｺｨ ｨｺ ｨｪｨｮ0506, ｨｨ 0904ｨ 0608ｨｺｨｨ ｨｨ ｨｪ ｨ ｨｨ06ｨｰ0607ｨ ｨｬ ｨｪ04 09 ｨｰｨ ｨｺ060904ｨｬｨｨ ｨｨ ｨｮ ｨ ｨｨ06ｨｰ0607ｨ (070605ｨｪ ｨｰ0906, Clarke, Warwick, 2001) ｨｮ- ﾂ02ｨｪｨｪｨ ｨｮ05ｨｹｨｰｨ ｨｰ02 ｨ ｨｪｨ 05ｨｨ04ｨ ｨｨ ﾂｨｨｨｪｨ R-0706ｨｺｨ 04ｨ ｨｰ ｨ 09ｨｪｨ 0,394 ｨｪｨ ｨｺ06ｨｬ ｨｮ080609ｨｪ02 04ｨｪｨ ﾂｨｨｨｬ0609ｨｰｨｨ (ｨｰｨ ｨ 05. 3). 06ｨｰ ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ02ｨｰ 06 ﾂ ﾂｨ ｨｦｨｪ06 ｨｪｨｨ04ｨｺ06ｨｦ ｨｰｨｪ0609ｨｰｨｨ ｨｰ060006, ﾂｨｰ06 08ｨ 0405ｨｨ ﾂｨｨ07 ｨｬ020301ｨｮ ｨｬｨｨ ｨｰ0802ｨｬ07 ｨ ｨｪｨ 05ｨｨ04ｨｨ08ｨｮ02ｨｬ04ｨｬｨｨ 0008ｨｮ0707ｨｨ080609ｨｺｨ ｨｬｨｨ

10 130804ｨ ｨｪ060907ｨｰ 0905ｨｮ ﾂｨ ｨｦｨｪ04ｨｦ 01ｨ 08ｨ ｨｺｨｰ0208, ｨｨ ｨｰ06ｨｬｨｮ ｨｪｨｮ05ｨｹ-00ｨｨ0706ｨｰ0204ｨ ("ｨｪ02ｨｰ 08ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｬ020301ｨｮ 0008ｨｮ0707ｨｨ080609ｨｺｨ ｨｬｨｨ 0804ｨ 09 ｨ ｨｨ06ｨｰ0607ｨ 01") ｨｬ060302ｨｰ ｨ 04ｨｰｨｹ 06ｨｰ ｨｪｨｮｨｰｨ. 0508ｨ 09ｨｪ02ｨｪｨｨ ｨｺｨ 04ｨ ｨｰ020502ｨｦ R 0708ｨｨ ｨ 08ｨｪ ｨ 09ｨｪ02ｨｪｨｨ ｨｺｨ ｨ 02ｨｰ, ﾂｨｰ ｨ ｨｴ0209ｨｰ09ｨ 0804ｨ 0709ｨｪ06 08ｨ 0405ｨｨ ﾂｨｪ04, 0106ｨｰ07 ｨｨ ｨｪ0209ｨｺ0605ｨｹｨｺ ｨｺ080409ｨ 06ｨｰ0907 (ｨｰｨ ｨ 05. 3). 04ｨ 0405ｨｨ ﾂｨｨ ｨｰｨ ｨｮ0707ｨｨ ｨｺ 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ ｨｰ ｨｪ04 ｨｪｨ ｨｺ06ｨｬ ｨｮ080609ｨｪ02 04ｨｪｨ ﾂｨｨｨｬ0609ｨｰｨｨ. 0306ｨｺｨ 04ｨ ｨｰ0205ｨｹ R ｨｨｨｬ0202ｨｰ ｨｪｨ ｨｨｨｬ02ｨｪｨｹ03ｨｮ ｨｨ ﾂｨｨｨｪｨｮ 0708ｨｨ ｨ 08ｨｪ06ｨｬ 0908ｨ 09ｨｪ02ｨｪｨｨｨｨ ｨ ｨｴ0209ｨｰ09 ｨｨ04 ｨ ｨｨ06ｨｰ ｨｨ 00, ﾂｨｰ ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ02ｨｰ 06 03ｨｨ0806ｨｺ06ｨｬ ｨｺ080409ｨ ｨｪｨｨｨｨ 05ｨｰｨｨ ｨ ｨｴ0209ｨｰ09, 09 ｨｰ ｨｬ ｨｨ- ﾂｨｨｨｪｨ R 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｨ ｨｰ ｨｪ06 06ｨｰ05ｨｨ ﾂｨ 02ｨｰ ｨｰ ｨｪｨｮ0507 (ｨｰｨ ｨ 05. 3). 06ｨ ｨ 05ｨｨ02ｨ ｨｮ05ｨｹｨｰｨ ｨｰ04 ｨ ｨｪｨ 05ｨｨ04ｨ ANOSIM 0708ｨｨ 0908ｨ 09ｨｪ02ｨｪｨｨｨｨ ｨｰｨ 09ｨ ｨｨ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ ｨ ｨｴ0209ｨｰ ｨ 09 ｨｰ ｨ ｨｨ06ｨｰ0607ｨ 01 ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ, ｨｪｨ ｨｨｨｰ0608ｨｨ07, ｨｰ08ｨ 05ｨｨ07 Table 3 ANOSIM test results of fish species composition and abundance at three studied habitats in Port Essington Arafura Sea, Northern Territory, Australia 0508ｨ 09ｨｪｨｨ09ｨ 02ｨｬ0402 ｨ ｨｨ06ｨｰ ｨｨ ﾂｨｨｨｪｨ R ｨｪｨｹ 04ｨｪｨ ﾂｨｨｨｬ0609ｨｰｨｨ A ｨｨ 09 0,618 p = 0,001 A ｨｨ 00 0,260 p = 0, ｨｨ 00 0,317 p = 0, ｨｨｨｬ02 ﾂｨ ｨｪｨｨ02. R-09ｨｰｨ ｨｰｨｨ09ｨｰｨｨｨｺｨ = 0,394, ｨｮ ｨｪｨｹ 04ｨｪｨ ﾂｨｨｨｬ0609ｨｰｨｨ R-09ｨｰｨ ｨｰｨｨ09ｨｰｨｨｨｺｨｨ 08 = 0,001. Aｨｪｨ 05ｨｨ04 SIMPER ｨｪｨｨ09ｨ 02ｨｰ 09ｨｪｨｮｨｰ0802ｨｪｨｪ ｨｰ ｨ ｨｴ0209ｨｰ ｨ 0601ｨｪ ｨ ｨｨ06ｨｰ0607ｨ, ｨｰ "ｨｰｨｨ07ｨｨ ﾂｨｪ0402" 09ｨｨ ｨｰｨｨ ｨ ｨｴ0209ｨｰ09ｨ ｨｨ04ｨｪｨ ｨｺｨ ｨｬ ｨｰ0607ｨｪｨｪ ｨｰ ｨｴ02ｨｦ ｨｺ06ｨｦ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ ｨ 0608ｨｺｨ 01 ｨｨ ｨｨ ｨｨｨｰ 0908ｨ 09ｨｪ02ｨｪｨｨ ｨ ｨｴ0209ｨｰ ｨ ｨｬ020301ｨｮ 0906ｨ 06ｨｦ (Clarke, Warwick, 2001). 05ｨ ｨｬ ｨｺｨｨ02 08ｨ 0405ｨｨ ﾂｨｨ07, ｨ 03ｨ 02ｨｬ0402 ｨｮ ｨｪ02ｨｪｨｪ04ｨｬｨｨ 04ｨｪｨ ﾂ02ｨｪｨｨ07ｨｬｨｨ 0706ｨｺｨ 04ｨ ｨｰ ｨ 0405ｨｨ ﾂｨｨ ｨｺ ｨｨ02ｨｨ02ｨｪｨｰｨｮ ｨｦ- 08ｨｮ08ｨｰｨｨ09ｨ (Clarke, Warwick, 2001), ｨ 0405ｨｨ ｨｪｨ ｨｦ0102ｨｪ04 ｨｬ020301ｨｮ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ 0804ｨ ｨｨ04 ｨ ｨｨ06ｨｰ ｨｨ ｨｮ070706ｨｦ ｨｨ ｨｬｨｨｨｪｨｨ08ｨｮ06ｨｴｨｨ01 09ｨｨ ｨ ｨ ｨｪ06 52,3 % ｨｪｨ ｨｺ ｨｪｨｪ ｨ 0405ｨｨ ﾂｨｨｨｦ (ｨｰｨ ｨ 05. 4) ｨｪ02ｨｪｨｪ0602 ｨｪ ｨｰ0906 ｨｬ020301ｨｮ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ 0804ｨ ｨｨ04 ｨ ｨｨ06ｨｰ ｨｨ 00 ｨ ｨｪ0204ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ06 ｨｪｨｨ ｨｨ01ｨ ｨｬｨｨ 0804ｨ ｨ ｨｪ06 52,9 % ｨｪｨ ｨｺ ｨｪｨｪ ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｬ020301ｨｮ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ ｨｨ04 05ｨｰｨｨ01 ｨ ｨｨ06ｨｰ ｨｰ06ｨｰ ｨ ｨｪｨ 05ｨｨ04, ｨｺｨ ｨｺ ｨｨ ｨｮｨｴｨｨｨｦ (ANOSIM), 0706ｨｺｨ 04ｨ 05, ﾂｨｰ06 09ｨ ｨｬ0402 ｨｪｨｨ04ｨｺｨｨ02 08ｨ 0405ｨｨ ﾂｨｨ07 (0706ｨｺｨ 04ｨ ｨｰ0205ｨｹ ｨｪ ｨｰ09ｨ ｨｦ-08ｨｮ08ｨｰｨｨ09ｨ 79,6 %) ｨ 0405ｨｨ ｨｬ020301ｨｮ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ 0804ｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺｨｨ, ｨ ｨｨ06ｨｰ , ｨｨ 09ｨｺｨ 05ｨｹｨｪ ｨｨ00ｨ, ｨ ｨｨ06ｨｰ (ｨｰｨ ｨ 05. 4) ｨｮ05ｨｹｨｰｨ ｨｰ ｨｨｨｪｨ 02ｨｨ06ｨｪｨｪ ｨ ｨｪｨ 05ｨｨ04ｨ, 0802ｨ 05ｨｨ040609ｨ ｨｪｨｪ ｨｨ 0706ｨｬ06ｨｴｨｨ ｨｬ02ｨｰ0601ｨ ｨｬｨｪ060006ｨｬ0208ｨｪ ｨｺｨ 05ｨｨ080609ｨ ｨｪｨｨ07 (nmds, Clarke, Warwick, 2001), 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ06ｨｰ 06 ﾂ02ｨｰｨｺ06ｨｬ 08ｨ ｨｪｨｨｨｨ ｨｬ020301ｨｮ 0904ｨ 0608ｨｺｨ ｨｬｨｨ 0804ｨ ｨｨ04 ｨ ｨｨ06ｨｰ ｨｨ 09, ｨｰｨ ｨｴｨｨ ｨ 06ｨｦ ｨｺ06ｨｬ07ｨ ｨｺｨｰｨｪ ｨｮ (08ｨｨ09. 3). 00 ｨｰ ｨｬ ｨ 0608ｨｺｨｨ ｨｨ04 ｨ ｨｨ06ｨｰ0607ｨ 00 ｨｨｨｬ0206ｨｰ ｨｪｨ 0008ｨ 00ｨｨｨｺ ｨｮ06, 08ｨ 04ｨｬｨ 04ｨ ｨｪｨｪｨｮ06 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｮ, 04ｨ ｨｪｨｨｨｬｨ 06ｨｰ ｨｬ0203ｨｮｨｰ06 ﾂｨｪ ｨｪｨｨ02 ｨｨ ｨｺ080409ｨ 06ｨｰ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ 0804ｨ ﾂｨ ｨｪ06ｨｦ ｨ ｨ ｨｪｨｺｨｨ ｨｨ ｨｬ0208ｨｰ ｨｺ0608ｨ ｨｨ00ｨ (08ｨｨ09. 3). 02ｨ ｨｴｨ 07 ｨｺｨ 08ｨｰｨｨｨｪｨ 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ02ｨｰ 06 ｨｪ0209ｨｺ0605ｨｹｨｺ06 ｨ 0605ｨｹ0302ｨｬ ｨｰ ｨｨ ｨｨ ｨｺ0605ｨｨ ﾂ0209ｨｰ0902ｨｪｨｪ ｨｰｨ 09ｨ ｨ ｨｴ0209ｨｰ ｨ 09 ｨ ｨｨ06ｨｰ0607ｨ ｨｨ 00 ｨｨ ｨｪ0209ｨｺ0605ｨｹｨｺ06 ｨ 0605ｨｹ0302ｨｦ 08ｨ 0406ｨ ｨｴ02ｨｪｨｪ0609ｨｰｨｨ 09 ｨｪｨｨｨｬｨｨ ｨ ｨｴ0209ｨｰ09ｨ 0804ｨ ｨｨ04 ｨ ｨｨ06ｨｰ0607ｨ 09, ﾂｨｰ ｨ 01ｨ 02ｨｰ ｨｮ05ｨｹｨｰｨ ｨｰｨ ｨｬｨｨ ｨ ｨｪｨ 05ｨｨ ANOSIM ｨｨ SIMPER ｨｨ 0908ｨ 09ｨｪ02ｨｪｨｨ02ｨｬ 04ｨｪｨ ﾂ02ｨｪｨｨｨｦ ｨｺ ｨｨ02ｨｨ02ｨｪｨｰ ｨ ｨｺｨｺｨ 08ｨ. 01ｨ 08ｨｨ ｨｰｨ ｨｪ04 ｨｺ08ｨｨ ｨｪｨ ｨｺ ｨｪｨｨ07 ﾂｨ 09ｨｰ06ｨｰ04 08ｨ 0405ｨｨ ﾂｨｪ ｨｨ ｨ, ｨｮ090205ｨｨ ﾂｨｨ09ｨ 06ｨｴｨｨ ｨｰ06ｨｬ ﾂｨｨ0905ｨ 0904ｨ ｨｺ. 02ｨ ｨｪｨｪ0402 ｨ 0405ｨｨ 06ｨ 06ｨ ｨｴ02ｨｪ ｨｮ05ｨｹｨｰｨ ｨｰｨ ｨｬ ｨｰｨ ｨｪ060906ｨｺ (Clarke, Warwick, 2001), ﾂｨｰ ｨｨ ｨｮ ﾂｨｨｨｰｨｹ ｨ ｨ 09ｨｪ0402 ｨｺ08ｨｨ ｨｪｨ 03ｨｨｨｬ 01ｨ ｨｪｨｪ04ｨｬ, ｨｪ0206ｨ ｨｨｨｬ06 8ｨC9 09ｨｨ ｨ ｨｺ 09 ｨ ｨｨ06ｨｰ0607ｨ ｨｨ 09, ﾂｨｰ06ｨ 04 ｨｪｨ ｨｺ ｨｪｨｪ0402 ｨｺ08ｨｨ ｨｰｨｨ0005ｨｨ ｨ 09ｨｨｨｬ07ｨｰ06ｨｰ04, 09 ｨｰ ｨｬ07 ｨｺｨ ｨｺ ｨｺ08ｨｨ09ｨ ｨ ｨｪｨｪ04ｨｬ ｨ ｨｺ 09 ｨ ｨｨ06ｨｰ ｨｴ02 ｨｪ ｨｰｨｨ0005ｨ ｨ 09ｨｨｨｬ07ｨｰ06ｨｰ04 ｨｨ ｨｪ02ｨｺ06ｨｰ ﾂｨｨ ｨｪ ｨｨ ｨｴ02 ｨ ｨ 04 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪ06, ｨｨ ｨ 04 09ｨｨ ｨｲ02ｨｬｨｺｨ ｨ 05ｨ 09ｨｹ (08ｨｨ09. 4). 190

11 1306ｨ ｨ 05ｨｨ02ｨ ｨ 09ｨｪ02ｨｪｨｨ ｨｰｨ 09ｨ ｨ ｨｴ0209ｨｰ ｨ 09 06ｨ ｨ ｨｪｨｪ0401 ｨ ｨｨ06ｨｰ0607ｨ ｨｬｨ ｨｰ0208ｨｨｨ 05ｨ ｨｬ 09ｨｨ010206ｨｮ ﾂ02ｨｰ0609, ｨｪｨｪ ｨ ｨｮ01ｨｰ ｨｰ ｨｨｨｪ00ｨｰ06ｨｪ, ｨｮ05ｨｹｨｰｨ ｨｰｨ ｨｬ ｨ ｨｪｨ 05ｨｨ04ｨ SIMPER Table 4 Comparison of fish assemblages at three survey sites in Port Essington using SIMPER analysis 00ｨｨ01 ﾁｨｨ090502ｨｪｨｪ0609ｨｰｨｹ ﾁｨｨ090502ｨｪｨｪ0609ｨｰｨｹ ｦﾄ j, % ｦｲｦﾄ, % j ｨｨ02ｨｨ02ｨｪｨｰ ｨｪ ｨｰ09ｨ ｨ ｨｴ0209ｨｰ ｨ ｨｨ04 ｨ ｨｨ06ｨｰ ｨｨ 09, ｦﾄ = 84,3 % Pantolabus radiatus 5,78 ﾀ 1,71 0 6,47 6,47 Lutjanus erythropterus 4,56 ﾀ 1,60 0 5,71 12,18 Leiognathus equulus 4,33 ﾀ 1,13 1,33 ﾀ 0,13 5,66 17,84 Caranx sexfasciatus 3,22 ﾀ 0,76 0,73 ﾀ 0,52 5,44 23,27 Acanthurus grammoptilus 0 4,80 ﾀ 2,04 5,22 28,50 Lethrinus ornatus 3,00 ﾀ 1,07 0,60 ﾀ 0,25 4,41 32,91 Pentapodus porosus 2,78 ﾀ 0,98 2,67 ﾀ 0,95 4,38 37,29 Gnathanodon speciosus 1,56 ﾀ 1,56 6,27 ﾀ 2,37 4,02 41,52 Plectorhinchus labiosum 0,11 ﾀ 0,11 1,13 ﾀ 0,22 3,97 45,50 Scolopsis monogramma 0,22 ﾀ 0,15 0,93 ﾀ 0,21 3,47 48,97 Scomberoides commersonnianus 0,67 ﾀ 0,33 1,67 ﾀ 1,04 3,30 52, ｨｨ02ｨｨ02ｨｪｨｰ ｨｪ ｨｰ09ｨ ｨ ｨｴ0209ｨｰ ｨ ｨｨ04 ｨ ｨｨ06ｨｰ ｨｨ 00, ｦﾄ = 79,6 % Pantolabus radiatus 5,78 ﾀ 1,71 0,29 ﾀ 0,19 7,38 7,38 Lutjanus erythropterus 4,56 ﾀ 1,60 0 7,09 14,46 Leiognathus equulus 4,33 ﾀ 1,13 0,71 ﾀ 0,07 6,80 21,27 Caranx sexfasciatus 3,22 ﾀ 0,76 1,29 ﾀ 0,71 5,65 26,91 Lethrinus ornatus 3,00 ﾀ 1,07 2,57 ﾀ 0,92 5,51 32,43 Pentapodus porosus 2,78 ﾀ 0,98 2,79 ﾀ 1,77 5,48 37,91 Sphyraena jello 0 0,79 ﾀ 0,19 4,26 42,16 Scomberoides commersonnianus 0,67 ﾀ 0,33 0,79 ﾀ 0,24 3,94 46,11 Carcharhinus melanopterus 0,67 ﾀ 0,24 0,07 ﾀ 0,07 3,52 49,63 Lutjanus carponatus 0,11 ﾀ 0,11 0,79 ﾀ 0,32 3,24 52, ｨｨ02ｨｨ02ｨｪｨｰ ｨｪ ｨｰ09ｨ ｨ ｨｴ0209ｨｰ ｨ ｨｨ04 ｨ ｨｨ06ｨｰ ｨｨ 00, ｦﾄ = 80,7 % Acanthurus grammoptilus 4,80 ﾀ 2,04 0,50 ﾀ 0,25 5,90 5,90 Pentapodus porosus 2,67 ﾀ 0,95 2,79 ﾀ 1,77 5,23 11,14 Plectorhinchus labiosum 1,13 ﾀ 0,22 0,07 ﾀ 0,07 4,78 15,92 Gnathanodon speciosus 6,27 ﾀ 2,37 1,43 ﾀ 1,10 4,76 20,68 Lethrinus ornatus 0,60 ﾀ 0,25 2,57 ﾀ 0,92 4,62 25,30 Scolopsis monogramma 0,93 ﾀ 0,21 0,71 ﾀ 0,46 4,39 29,69 Lutjanus carponatus 1,27 ﾀ 0,53 0,79 ﾀ 0,32 4,21 33,89 Sphyraena jello 1,27 ﾀ 0,99 0,79 ﾀ 0,19 4,12 38,01 Scomberoides commersonnianus 1,67 ﾀ 1,04 0,79 ﾀ 0,24 4,03 42,04 Nebris ferrugineus 0,80 ﾀ 0,20 0,07 ﾀ 0,07 3,86 45,91 Apogon sp. 2,13 ﾀ 0,98 0,50 ﾀ 0,34 3,58 49,49 Caranx sexfasciatus 0,73 ﾀ 0,52 1,29 ﾀ 0,71 3,54 53, ｨｨｨｬ02 ﾂｨ ｨｪｨｨ02. ﾁｨｨ090502ｨｪｨｪ0609ｨｰｨｹ ｪ ｨｪ ｨｪｨ ﾂ02ｨｪｨｨ02 ﾀ 09ｨｰｨ ｨｪ01ｨ 08ｨｰｨｪｨ ｨｨｨ ｨｺｨ ﾂｨｨ0905ｨ 0804ｨ, 04ｨ ｨｨ09ｨｰ08ｨｨ080609ｨ ｨｪｨｪ ｨｬ ｨｪ ｨｨ010206ｨｮ ﾂ02ｨｰｨ. 02ｨ ｨｪｨｪ ﾂｨｨ090502ｨｪｨｪ0609ｨｰｨｨ 09ｨｨ ｨ 0405ｨｨ ｨ 08ｨ ｨ ｨｪ ｨｬ07 ｨ ｨｪｨ 05ｨｨ04ｨ SIMPER ｪ ｨｨ ﾂ02ｨｪ ｨｺ060802ｨｪｨｹ 4-ｨｦ 09ｨｰ020702ｨｪｨｨ. ｦﾄ j ｪ 06ｨｰｨｪ0609ｨｨｨｰ0205ｨｹｨｪ04ｨｦ 09ｨｺ05ｨ 01 i ｨｨ01ｨ ｨｪ0202 ｨｪ ｨｰ ｨｦ-08ｨｮ08ｨｰｨｨ09ｨ (ｦﾄ) ｨｬ020301ｨｮ ｨ ｨｴ0209ｨｰ09ｨ ｨｬｨｨ; ｦｲｦﾄ j ｪ ｨｪｨ ｨｺ ｨｪｨｪ0402 ｨｺ ｨｨ02ｨｨ02ｨｪｨｰ04 ｨｪ ｨｰ09ｨ. 0507ｨｨ0906ｨｺ 09ｨｨ ｨｨ090601ｨｨｨｰ ｨｬ ｨｮｨ 0409ｨ ｨｪｨｨ07 ｨｨ01 09ｨｺ05ｨ 01ｨ ｨｪ0202 ｨｪ ｨｰ0906 (ｦﾄ). 0308ｨｨ ｨｪ04 01ｨ ｨｪｨｪ ｨｨ01ｨ ｨｬ c ｨｺｨｨｨｬ 09ｨｺ05ｨ 0106ｨｬ (6ﾕ5 50 % ｨｨ ). 0006ｨｴｨｪ0609ｨｰｨｹ 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺ ｨｺ08ｨｨｨｰ0208ｨｨ07 ｨ 0409ｨｰ ｨ 09ｨｰｨ 05ｨ 0706 ｨｬ ｨｮ090205ｨｨ ﾂ02ｨｪｨｨ07 ﾂｨｨ0905ｨ 0904ｨ ｨｺ, ｨｨ ｨ ｨｨ06ｨｰ0607ｨ 01 ｨｮ ｨ ｨｺ ｨ ｨｰｨ ｨｰ06 ﾂｨｪ06, ﾂｨｰ06ｨ ｨｮ ﾂｨｨｨｰｨｹ 0708ｨｨ02ｨｬ0502ｨｬｨｮ06 ｨｬ06ｨｴｨｪ0609ｨｰｨｹ, 08ｨ 09ｨｪｨｮ06 80 %. 0308ｨｨ ｨｪｨ 05ｨｨ ﾂｨｨｨｨ ｨ ｨｺ ｨｬ06ｨｴｨｪ0609ｨｰｨｹ 08ｨ 09ｨｪ0705ｨ 09ｨｹ 100 % (08ｨｨ09. 4). 0508ｨ 09ｨｪ02ｨｪｨｨ02 ｨｰ080600ｨｨ ﾂ0209ｨｺ06ｨｦ 09ｨｰ08ｨｮｨｺｨｰｨｮ ｨ ｨｴ0209ｨｰ ｨ ｨｨ04 08ｨ 04ｨｪ0401 ｨ ｨｨ06ｨｰ ｨｺｨ 04ｨ 0506, ﾂｨｰ06 ｨ 0605ｨｹ03ｨｨｨｪ09ｨｰ0906 ｨｮ ﾂｨｰ02ｨｪｨｪ ｨ 0708ｨｨｨｪｨ ｨｨｨｰ ｨｺ 0109ｨｮｨｬ ｨｺｨ ｨｰ ｨｨ07ｨｬ ｪ