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1 , No. 172,

2 1996 Oncorhynchus mykiss Salmo trutta aTaniguchi et al Morita et al. 2004Hasegawa et al Taniguchi et al (Aoyama et al Arai et al. 2002) Salvelinus leucomaenis S. malma , O. keta O. gorbuscha O. masou O. nerka a a -2-

3 , No. 172, 2006 Hucho perryi , , , (Baglinière 1991Elliott 1994) (MacCrimmon et al. 1970) a -3-

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5 , No. 172, ,,

6 100 m Kolmogorov-Smirnov 2 4 Kolmogorov-Smirnov 22 n 1 n n i 40 Kolmogorov-Smirnov n i (n i < 40) 153 Kolmogorv-Smirnov 2 (D max) 1,000 n i D max D max 153 (n i ) D max D max % -6-

7 , No. 172, % %

8 S. leucomaenis 9 O. kisutch

9 , No. 172, d, f, g, n f, g, n a, b, g, n a, b, g, n a, d, f, g, n, s a, b, n a, d, f, i a, b, d, i, n a, n n a a, b d d o n k, p a, b, c, d, n, p n d, e, l, n, r c, d a, d a, b, c, d a, b, c, d, o c 1 1 a, b, c, d, o, p a, b, c, d, e, n, o, p, q d, n 0 0 a a, d d 1 0 a, b, c, d 0 0 a, b, d d a, b, c, d, m d d -9-

10 d a, c, d, e, n, q a, o 0 0 a 1 0 a a, b a, b, d, n 0 0 a a, b 1 0 a, b, d 1 0 a, e 0 0 a, b, d a, b, c, d, p a, c, d 1 0 a, b, c, d p a 1 0 a, b, p 0 0 a, b, c, d a, b f, g f, g, n f, g d, f, g, n f, g f, g g g g d, f, g d, n a, d, g, n d, n d d a, b, d, n, o, r d, n, o d c, d, e, n, o, d, h, n, o a, b, d, e, j, n o a, b, c, d, n a, b, d a, b, l, o l a, l a, b, d, l a, d, l, n l a, n o -10-

11 , No. 172, s a, d a a a, c, o 0 0 a c, d, n a, c, d d a, b, c, d, p a, b, c, d b, e, p a a a a b c d e f, g h i j, k, l m n o 4, p q, r, s

12

13 , No. 172, 2006 (Kolmogorov-Smirnov 2 D max 0.29, n 1 = 153, n 2 = 48, p < 0.01D max 0.28, n 1 = 153, n 2 = 22, p < 0.05) (Kolmogorov-Smirnov 2 p > 0.05) ,

14 Micropterus salmoides M. dolomieu Lepomis macrochirus Channa argus S. fontinalis 2002b (Aoyama et al ) / (Arai et al. 2002)

15 , No. 172, B

16 (Fausch et al. 2001) Fausch et al

17 , No. 172, Heard

18 , b 2002b -18-

19 , No. 172, , 33: , 34: , 53: Aoyama, T., K. Naito, and T. Takami Occurrence of sea-run migrant brown trout (Salmo trutta) in Hokkaido, Japan. Sci. Rep. Hokkaido Fish Hatchery, 53: a., 38: b.., 56: Arai, T., A. Kotake, T. Aoyama, H. Hayano, and N. Miyazaki Identifying sea-run brown trout, Salmo trutta, using Sr:Ca ratios of otolith. Ichthyol. Res., 49: Baglinière, J. L La truite commune (Salmo trutta L. ), son origine, son aire de répartition, ses intérêts économique et scientifique. In La truite biologie et écologie (eds. Baglinière, J. L. and Maisse), INRA, Paris. pp , 7: , 10: , 10: , 11: p. Elliott, J. M Quantitative ecology and the brown trout. Oxford University Press, Oxford. 286p. Fausch, K. D., Y. Taniguchi, S. Nakano, G. D. Grossman, and C. R. Townsend Flood disturbance regimes influence rainbow trout invasion success among five holarctic regions. Ecol. Appl., 11: , 29: Hasegawa, H., T. Yamamoto, M. Murakami and K. Maekawa Comparison of competitive -19-

20 ability between native and introduced salmonids: evidence from pairwise contests. Ichthyol. Res., Heard, W. R Life history of pink salmon (Oncorhynchus gorbuscha). Pacific salmon life histories (eds. C. Groot, and L. Margolis), UBC Press, Vancouver. pp , 60: , 52: p. Hokkaido Fish Hatchery Scientific Reports of the Hokkaido Fish Hatchery, 50: Hokkaido Fish Hatchery Scientific Reports of the Hokkaido Fish Hatchery, 50: Hokkaido Fish Hatchery Scientific Reports of the Hokakido Fish Hatchery, 50: Hokkaido Fish Hatchery Scientific Reports of the Hokkaido Fish Hatchery, 51: Hokkaido Fish Hatchery Scientific Reports of the Hokkaido Fish Hatchery, 52: 1-10 Hokkaido Fish Hatchery Scientific Reports of the Hokkaido Fish Hatchery, 52: Hokkaido Fish Hatchery Scientific Reports of the Hokkaido Fish Hatchery, 56: : 10 p : 12 p pp , 45: , 45: , 45: , 28: , 46: , 46: , 29: , 30: , 31: 288 p , 49: , 49: , 32: , 32: , 50: , 33: , 33:

21 , No. 172, , 33: , 33: , 51: , 34: , 34: , 52: , 35: 418 p , 53: , 53: , 53: , 53: , 53: , 54: , 54: , 56: , 56: , 38: , 38: , 38: , 57: , 57: , 57: , 39: , 58: , 58: , 40: p , 8: ,. 247 p ,. 290 p a....,. p

22 . 2002b..,. 128 p , 17: p ,,. pp , 5: , 59: , 19: ,,. pp , 336: , 20: , 21: , 22: , 27: MacCrimmon, H. R., T. L. Marshall, and B. L. Gots World distribution of brown trout, Salmo trutta: further observations. J. Fish. Res. Bd. Canada, 27: , 2: , 36: , 37: , 3: , 167: , 24: Morita, K., J.Tsuboi, and H. Matsuda The impact of exotic trout on native charr in a Japanese stream. J. Appl. Ecol., 41: , 43: , 49: , 50:

23 , No. 172, ,. 390 p , 13: :., 3: , 67: , 68: , 70: pp , 21: , 23: , 23: , 24: , 25: , 25: , 29: pp , 160: , 164: , 166: , 1: , 4: , 68: Taniguchi, Y., Y. Miyake, T. Saito, H. Urabe, and S. Nakano Redd superimposition by introduced rainbow trout, Oncorhynchus mykiss, on native charrs in a Japanese stream. Ichthyol. Res., 47: Taniguch, Y., K. D. Fausch, and S. Nakano Size-structured interactions between native and introduced species: can intraguild predation facilitate invasion by stream salmonids? Biol. Invasions, 4: The ichthyological Society of Japan Ichthyological Research, 46: The Japanese Society of Fisheries Science Fisheries Science, 67: The Japanese Society of Fisheries Science Fisheries Science, 70: , 3: p

24 ,. 270 p , 131: ,., 19:

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