Bull. Kanagawa prefect. Mus. (Nat. Sci.), no. 3, pp. 1-6, Mar. 003 Three New Naturalized Plants of Carex L. (Cyperaceae) in Japan Teruo KATSUYAMA Abstract. Carex aenea Fernard, C. gravida L.H.Bailey var. lunelliana (Mack.) F.J.Herm. and C. fissa Mack. newly naturalized in Japan. C. aenea and C. fissa were collected from Tama-shi, Tokyo Metropolis, and C. gravida from Asao-ku, Kawasaki, Kanagawa prefecture and Tamashi. They all are natives of North America and were probably brought to Japan together with seeds of grasses for slope plantation. Key words: naturalized plant, Carex aenea, Carex gravida, Carex fissa Carex aenea Fernald 1 C. gravida L.H.Bailey var. lunelliana (Mack.) F.J.Herm. C. fissa Mack. 3 3 1-F Gleason & Cronquist 1991 1931 C. aenea C. aenea C. foenea Willd. Mackenzie C. foenea Mackenzie 1931 aenea Carex aenea Fernald, Proc. Amer. Acad. Arts 37: 480 (190) 50cm 1.5-4cm 3-6 4-4.5mm 3-5 1.5-mm 1.7-mm 1-B 3-7mm 1.-1.3mm 1-C, 1990 scoparia Schk. ex Willd 1-1.5mm 1-D,E C. sect. Ovales C. maackii Maxim. 1-.5mm 5-10mm 0.1mm C. 0.7mm : May 1, 000, (KPM-NA0118785); May 8, 000, (KPMNA01067). Carex gravida L.H.Bailey var. lunelliana(mack.) F.J.Herm., Amer. Midl. Naturalist 17: 855 (1936); C. lunelliana Mack., Bull. Torrey Bot. Club 4: 615 (1915)
T. Katsuyama 60-10cm 3-7mm -C 15-30mm 10mm 5-10 1 1 -B 3-4 mm -D 3.5-4 mm.5-3mm 3-5 1/3-1/4 - E,F mm - G 1-1.5mm 3 C. gravida 1/ var. gravida 3/4 var. lunelliana Yatskievych 1999 Gleason & Cronquist 1991 Mohlenbrock 1999 var. lunelliana Var. lunelliana Heavy Sedge Sect. Bracteosae Sect. Bracteosae sect. Multiflorae 1 C. sparganioides Muhl. ex Willd. C. aggregata Mack. C. cephaloidea (Dewey) Dewey Sect. Bracteosae C. leavenworthii Dewey, 000 4, May, 00, (KPM-NA01071, KPM-NA0107); 1, May 7, 00, (KPM- NA01073). Carex fissa Mack., N. Amer. Fl. 18: 64 (1931) 60-100cm 4-6mm 3-6mm 3-C 3-4cm 1-1.5cm 1-3 10-0 3-B 3-3.5mm 1mm 3-D 3.5-4mm.5-3mm 3-5 3 1/3 3-E,F mm 1.5-1.8mm 3-G sect. Multiflorae C. brachyglossa Mack. C. vulpinoidea Michx., 001-4mm -3mm 3 : Jun. 4, 001, (KPM-NA01068, KPM-NA01070); May 4, 001, (KPM-NA01069). Gleason, H.A. & A.Cronquist, 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada, Second Edition. lxxv+910pp. New York Botanical Garden, New York., 000.. : 34-36., 001.., 001. pp.44-48. Mackenzie, K.K., 1931-1935. Cyperaceae-Cariceae. North American Flora : 1-478. New York Botanical Garden, New York., 1990.., (1): 4505-4508. Mohlenbrock, R.H., 1999. The Illustrated Flora of Illinois, Sedges: Carex. 38pp. Southern Illinois University Press, Carbondale and Edwardville. Yatskievych, G., 1999. Steyermark's Flora of Missouri, reviced ed. 1: 1-991. The Missouri Department of Conservation, Jefferson City.
New Naturalized Plants of Carex in Japan 3 1. Carex aenea Fernard A:. B:. C:. D:. E:. F:., A: 10cm, B: 1cm, C-F: 1mm. Fig. 1. Carex aenea Fernard A: Plant. B: Inflorescens. C: Pistillate scale. D: Perigynium (dorsal view). E: Perigynium (ventral view). F: Achen. Scale, A: 10cm, B: 1cm, C-F: 1mm.
4 T. Katsuyama. Carex gravida L.H.Bailey var. lunelliana (Mack.) F.J.Herm. A:. B:. C:. D:. E:. F:. G:., A: 10cm, B: 1cm, C: 1mm, D-G: 1mm. Fig.. Carex gravida L.H.Bailey var. lunelliana (Mack.) F.J.Herm. A: Plant. B: Inflorescens. C: Sheath.(side view) D: Pistillate scale. E: Perigynium (dorsal view). F: Perigynium (ventral view). G: Achen. Scale, A: 10cm, B: 1cm, C: 1mm, D-G: 1mm.
New Naturalized Plants of Carex in Japan 5 3. Carex fissa Mack. A:. B:. C:. D:. E:. F:. G:., A: 10cm, B: 1cm, C: 5mm, D-G: 1mm. Fig. 3. Carex fissa Mack. A: Plant. B: Inflorescens. C: Sheath.(ventral view) D: Pistillate scale. E: Perigynium (dorsal view). F: Perigynium (ventral view). G: Achen. Scale, A: 10cm, B: 1cm, C: 5mm, D-G: 1mm.
6 T. Katsuyama, 00.., (3): 1-6. (T. Katsuyama, 003. Three New Naturalized Plants of Carex L. (Cyperaceae) in Japan. Bull. Kanagawa prefect, Mus. (Nat. Sci.), (3): 1-6.) Carex aenea Fernald C. gravida L.H.Bailey var. lunelliana (Mack.) F.J.Herm. C. fissa Mack. 3