130503090505080209020508070506000301010808 0906020902010608, 2013, 01 5, 09. 61-68 050609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｹ 0708ｨｨ 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨｨｨ 070208 633.18:631.522/.524:575.222.7 doi: 10.15389/agrobiology.2013.5.61rus 050603040609050102050605 030406 00030403020200060202000207 010609040602060508000606 04060508 Oryza sativa L. 00 0500080506 05 03020605080200 0103010200 01060402080207 05020000030506060002050606 06 02060103050301060300 0209040805000200 08 0302020006020800 indica 06 japonica 6ﾓ5 03.00. 090804060602010200, 07.08. 010201 ﾁ0804020008 00020301ｨｮ 07060109ｨｨ01ｨ ｨｬｨｨ 08ｨｨ09ｨ (Oryza sativa L.) indica ｨｨ japonica 09ｨｮｨｴ0209ｨｰ09ｨｮ02ｨｰ 080701 07020802010601ｨｪ0401 000608ｨｬ ｨｺｨ ｨｺ 0706 0002ｨｪ06ｨｰｨｨ07ｨｮ, ｨｰｨ ｨｺ ｨｨ 0706 00ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨｨｬ 0708ｨｨ04ｨｪｨ ｨｺｨ ｨｬ, 06ｨ ｨｮ0905060905ｨｨ09ｨ 06ｨｴｨｨｨｬｨｨ 060906ｨ 02ｨｪｨｪ0609ｨｰｨｨ 08ｨ 0409ｨｨｨｰｨｨ07 ｨｨ ｨ 01ｨ 07ｨｰｨ 02ｨｨｨｨ ｨｺ 09ｨｰ08020909ｨ ｨｬ. 0102ｨｰ02080604ｨｨ09ｨｪ04ｨｦ 05000002ｨｺｨｰ ｨｮ ｨｬ020307060109ｨｨ0106090401 00ｨｨｨ 08ｨｨ010609 (japonica/indica) ｨｬ060302ｨｰ 090609ｨｰｨ 090507ｨｰｨｹ 100 % ｨｨ 09040302. 0609070605ｨｹ040609ｨ ｨｰｨｹ 090902080109040906ｨｺｨｨｨｦ 0002ｨｰ02080604ｨｨ09 ｨｰｨ ｨｺｨｨ01 00ｨｨｨ 08ｨｨ010609 ｨｬ0603ｨｪ06, 020905ｨｨ 0906090502ｨｺｨ ｨｰｨｹ 09 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨ06 06ｨ 08ｨ 040204 09 0002ｨｪｨ ｨｬｨｨ 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ (WC), ｨｺ06ｨｰ06080402 06ｨ 02090702 ﾂｨｨ09ｨ 06ｨｰ 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｹ 00ｨｨｨ 08ｨｨ010609 ｨ 06050202 70 %. 0004 070806090205ｨｨ 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨ06 ｨｬ020301ｨｮ 06ｨ 08ｨ 0402ｨ ｨｬｨｨ 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ06ｨｦ ｨｨ 04ｨ 08ｨｮｨ 0203ｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 06ｨｰｨｪ060907ｨｴｨｨｨｬｨｨ0907 ｨｺ 07060109ｨｨ01ｨ ｨｬ indica ｨｨ japonica. 08ｨｪｨ 05ｨｨ04 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｨ 070605ｨｮ ﾂ02ｨｪｨｪ0401 ｨｬ020307060109ｨｨ0106090401 00ｨｨｨ 08ｨｨ010609 ｨｪ02 070604090605ｨｨ05 0904010205ｨｨｨｰｨｹ 09080201ｨｨ 06ｨ 08ｨ 04020609 08060909ｨｨｨｦ09ｨｺ06ｨｦ 09020502ｨｺ02ｨｨｨｨ 090608ｨｰｨ 09 0002ｨｪｨ ｨｬｨｨ 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ. 020507 ｨｺ06080802ｨｺｨｰｨｪ060006 06ｨｰｨｪ020902ｨｪｨｨ07 090608ｨｰ0609 08ｨｨ09ｨ ｨｺ 07060109ｨｨ01ｨ ｨｬ 0904070605ｨｪ02ｨｪ06 ｨｬ060502ｨｺｨｮ050708ｨｪ0602 ｨｬｨ 08ｨｺｨｨ080609ｨ ｨｪｨｨ02 09 ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ02ｨｬ 11 070605ｨｨｨｬ060800ｨｪ0401 SSR-ｨｬｨ 08ｨｺ02080609, ｨｰｨｨ07ｨｨ ﾂｨｪ0401 010507 07060109ｨｨ010609. 02ｨ 09ｨｮ0301ｨ 06ｨｰ0907 09060708060904 070605ｨｮ ﾂ02ｨｪｨｨ07 000208ｨｰｨｨ05ｨｹｨｪ0401 ｨｬ020307060109ｨｨ0106090401 00ｨｨｨ 08ｨｨ010609 08ｨｨ09ｨ ｨｨ 07020802 ﾂ02ｨｪｨｹ 0708ｨｨ04ｨｪｨ ｨｺ0609, ｨｨ09070605ｨｹ04ｨｮ02ｨｬ0401 010507 08ｨ 0401020502ｨｪｨｨ07 06ｨ 08ｨ 04020609 ｨｪｨ 07060109ｨｨ0104. 0709ｨｰｨ ｨｪ06090502ｨｪｨ ｨｪ0206ｨ 010601ｨｨｨｬ0609ｨｰｨｹ 09ｨｺ0506 ﾂ02ｨｪｨｨ07 0106070605ｨｪｨｨｨｰ0205ｨｹｨｪ0401 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ09ｨｰｨｨｨｺ 09 0907ｨｨ0906ｨｺ 0708ｨｨ04ｨｪｨ ｨｺ0609, ｨｪｨ 0609ｨｪ0609ｨ ｨｪｨｨｨｨ ｨｺ06ｨｰ06080401 01060503ｨｪ06 070806090601ｨｨｨｰｨｹ0907 08ｨ 0401020502ｨｪｨｨ02 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ0401 06ｨ 08ｨ 04020609 ｨｪｨ 07060109ｨｨ0104 indica ｨｨ japonica. 080506 ﾂ02090402 09050609ｨ : 08ｨｨ09, 0002ｨｰ02080604ｨｨ09, 00ｨｨｨ 08ｨｨ0104, 0002ｨｪ04 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ, 07060109ｨｨ0104 08ｨｨ09ｨ 070706ｨｪ09ｨｺｨｨｨｦ ｨｨ ｨｨｨｪ01ｨｨｨｦ09ｨｺｨｨｨｦ, japonica, indica. 04ｨ 0409ｨｨｨｰｨｨ02 080207080601ｨｮｨｺｨｰｨｨ09ｨｪ0401 ｨ ｨ 08ｨｹ02080609 ｨｨ0008ｨ 02ｨｰ ｨｺ0506 ﾂ0209ｨｮ06 050906050602ｨｨ06ｨｪｨｪｨｮ06 080605ｨｹ, 07060101020803ｨｨ09ｨ 07 0002ｨｪ02ｨｰｨｨ ﾂ0209ｨｺｨｮ06 01ｨｨ0902080002ｨｪ02ｨｨ06 ｨｨ 0906ｨｺ08ｨ ｨｴｨ 07 06ｨ ｨｬ02ｨｪ 0002ｨｪｨ ｨｬｨｨ ｨｬ020301ｨｮ 08ｨ 0405ｨｨ ﾂｨｪ04ｨｬｨｨ 070607ｨｮ050702ｨｨ07ｨｬｨｨ. 0306-09ｨｨ01ｨｨｨｬ06ｨｬｨｮ, ｨｪｨ 05ｨｨ ﾂｨｨ02 ｨｰｨ ｨｺｨｨ01 ｨ ｨ 08ｨｹ02080609 ｨｨｨｬ020506 0609ｨｪ0609ｨｪ0602 04ｨｪｨ ﾂ02ｨｪｨｨ02 09 08ｨ 0401020502ｨｪｨｨｨｨ 07060109ｨｨ010609 08ｨｨ09ｨ japonica ｨｨ indica (1). 050608ｨｰｨ 07060109ｨｨ01ｨ japonica 08ｨ 09ｨｰｨｮｨｰ 09 0406ｨｪｨ 01 09 ｨｮｨｬ020802ｨｪｨｪ04ｨｬ ｨｺ05ｨｨｨｬｨ ｨｰ06ｨｬ, 090608ｨｰｨ ｨｨｨｪ01ｨｨｨｦ09ｨｺ060006 07060109ｨｨ01ｨ 08ｨ 0907080609ｨｰ08ｨ ｨｪ02ｨｪ04 09 ｨｰ080607ｨｨ ﾂ0209ｨｺｨｨ01 ｨｨ 09ｨｮｨ ｨｰ080607ｨｨ ﾂ0209ｨｺｨｨ01 080200ｨｨ06ｨｪｨ 01 (2). 00020301ｨｮ 07060109ｨｨ01ｨ ｨｬｨｨ 09ｨｮｨｴ0209ｨｰ09ｨｮ02ｨｰ 080701 07020802010601ｨｪ0401 000608ｨｬ ｨｺｨ ｨｺ 0706 0708ｨｨ04ｨｪｨ ｨｺｨ ｨｬ, 060708020102050706ｨｴｨｨｨｬ 0002ｨｪ06ｨｰｨｨ07, ｨｰｨ ｨｺ ｨｨ 0706 00ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨｨｬ 090906ｨｦ09ｨｰ09ｨ ｨｬ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ, 06ｨ ｨｮ0905060905ｨｨ09ｨ 06ｨｴｨｨｨｬ 060906ｨ 02ｨｪｨｪ0609ｨｰｨｨ 08ｨ 0409ｨｨｨｰｨｨ07 ｨｨ ｨ 01ｨ 07ｨｰｨ 02ｨｨｨｨ ｨｺ 09ｨｰ08020909ｨ ｨｬ (3). 05ｨｮｨｴ0209ｨｰ09ｨｮ06ｨｰ 010902 00ｨｨ0706ｨｰ020404 070806ｨｨ090106030102ｨｪｨｨ07 07060109ｨｨ010609. 05060005ｨ 09ｨｪ06 0702080906ｨｦ, 070706ｨｪ09ｨｺｨｨｨｦ 07060109ｨｨ01 070806ｨｨ0406030205 ｨｨ04 06ｨ 08ｨ 04020609 ｨｨｨｪ01ｨｨｨｦ09ｨｺ060006 07060109ｨｨ01ｨ 0708ｨｨ ｨｨ01 ｨ 01ｨ 07ｨｰｨ 02ｨｨｨｨ ｨｺ ｨｮ09050609ｨｨ07ｨｬ ｨ 06050202 0902090208ｨｪ0401 080200ｨｨ06ｨｪ0609 (4). 0208ｨｮ00ｨ 07 ｨｰ020608ｨｨ07 0006090608ｨｨｨｰ 06 0902080607ｨｰｨｪ06ｨｦ 07ｨ 08ｨ 05050205ｨｹｨｪ06ｨｦ 0106ｨｬ0209ｨｰｨｨ00ｨｨｨｺｨ 02ｨｨｨｨ 07060109ｨｨ010609, 070806ｨｨ0901060107ｨｴｨｨ01 06ｨｰ 08ｨ 0405ｨｨ ﾂｨｪ0401 01ｨｨｨｺｨｨ01 08060109ｨｰ0902ｨｪｨｪｨｨｨｺ0609 (5). 05ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ0402 0002ｨｪ02ｨｰｨｨ ﾂ0209ｨｺｨｨ02 08ｨ 0405ｨｨ ﾂｨｨ07 ｨｬ020301ｨｮ 07060109ｨｨ01ｨ ｨｬｨｨ 0708ｨｨ09060107ｨｰ ｨｺ 09040906ｨｺ06ｨｬｨｮ 0002ｨｰ02080604ｨｨ09ｨｮ 0708ｨｨ ｨｨ01 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨｨｨ, ﾂｨｰ06 0706040906050702ｨｰ ｨｪｨ 010207ｨｰｨｹ0907 ｨｪｨ 090604ｨｬ0603ｨｪ0609ｨｰｨｹ ｨｮ090205ｨｨ ﾂ02ｨｪｨｨ07 0706ｨｰ02ｨｪ02ｨｨｨ 05ｨ 07080601ｨｮｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｨ ｨｺｨｮ05ｨｹｨｰｨｮ0804 04ｨ 09 ﾂ02ｨｰ ｨｬ020307060109ｨｨ01060906ｨｦ 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨｨｨ (6, 7). 0609090502010609ｨ ｨｪｨｨ07 0706ｨｺｨ 04ｨ 05ｨｨ, ﾂｨｰ06 ｨｺ 01ｨｨ0000020802ｨｪ02ｨｨｨ 02ｨｨｨｨ 07060109ｨｨ010609 0708ｨｨ090205ｨ ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨ07 ｨｰｨ ｨｺｨｨ01 0002ｨｪ02ｨｰｨｨ ﾂ0209ｨｺｨｨ01 0906ｨ 04ｨｰｨｨｨｦ, ｨｺｨ ｨｺ 0909ｨｰｨ 09ｨｺｨｨ, 0102050202ｨｨｨｨ, 04ｨ ｨｬ02ｨｪ04 ｨｪｨｮｨｺ050206ｨｰｨｨ010609. 01ｨ ｨｨｨ 0605ｨｹ030202 ﾂｨｨ090506 08ｨ 0405ｨｨ ﾂｨｨｨｦ ｨｬ020301ｨｮ 07060109ｨｨ01ｨ ｨｬｨｨ, 09090704ｨ ｨｪｨｪ0401 09 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ02ｨｬ ｨｪｨｮｨｺ050206ｨｰｨｨ01ｨｪ0401 0706090502010609ｨ ｨｰ0205ｨｹｨｪ0609ｨｰ02ｨｦ, 04ｨ 00ｨｨｨｺ09ｨｨ080609ｨ ｨｪ04 ｨｪｨ 11-ｨｦ ｨｨ 12-ｨｦ 010806ｨｬ060906ｨｬｨ 01 (8-10). 01ｨ ｨｰ020808ｨｨｨｰ0608ｨｨｨｨ 04060909ｨｨｨｦ09ｨｺ06ｨｦ 0802010208ｨ 02ｨｨｨｨ 09 0609ｨｪ0609ｨｪ06ｨｬ ｨｨ09070605ｨｹ04ｨｮ06ｨｰ0907 090608ｨｰｨ 08ｨｨ09ｨ 07060109ｨｨ01ｨ japonica. 01ｨｨ04ｨｺｨ 07 0002ｨｪ02ｨｰｨｨ ﾂ0209ｨｺｨ 07 0002ｨｰ0208060002ｨｪｨｪ0609ｨｰｨｹ 090608ｨｰ0609 05ｨｰ060006 07060109ｨｨ01ｨ 0708ｨｨ090601ｨｨｨｰ ｨｺ ｨｪ0209040906ｨｺ06ｨｬｨｮ 0002ｨｰ02080604ｨｨ09ｨｮ 0708ｨｨ ｨｨ01 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨｨｨ 6ﾓ5 04ｨ ｨ 06ｨｰｨ 07060101020803ｨ ｨｪｨ 0008ｨ ｨｪｨｰ06ｨｬ 04080806 01 11-04-96534-08_0600_02. 61
13(11, 12). 07 ｨｬ020307060109ｨｨ0106090401 00ｨｨｨ 08ｨｨ010609 (japonica/indica) 0002ｨｰ02080604ｨｨ09ｨｪ04ｨｦ 05000002ｨｺｨｰ ｨｬ060302ｨｰ 090609ｨｰｨ 090507ｨｰｨｹ 100 % ｨｨ ｨ 06050202. 0201ｨｪｨ ｨｺ06 090905020109ｨｰ09ｨｨ02 09040906ｨｺ06ｨｦ 07ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｨ ｨｬ020307060109ｨｨ0106090402 00ｨｨｨ 08ｨｨ0104 08ｨｨ09ｨ 0106 09ｨｨ01 070608 070806ｨｬ04030502ｨｪｨｪ06 ｨｪ02 070806ｨｨ0409060107ｨｰ0907, 00ｨｨｨ 08ｨｨ0104 07060109ｨｨ01ｨ japonica ｨｰｨ ｨｺ0302 ｨｪ02 070605ｨｮ ﾂｨｨ05ｨｨ 03ｨｨ0806ｨｺ060006 08ｨ 0907080609ｨｰ08ｨ ｨｪ02ｨｪｨｨ07 (0706090209ｨｪ0402 070506ｨｴｨ 01ｨｨ ｨｪ02 070802090403ｨ 06ｨｰ 5 %) ｨｨ04-04ｨ ｨｪ0204ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ060006 0002ｨｰ02080604ｨｨ09ｨｪ060006 05000002ｨｺｨｰｨ (13). 00 ｨｰ06 0302 090802ｨｬ07 0706090209ｨｪ0402 070506ｨｴｨ 01ｨｨ 070601 00ｨｨｨ 08ｨｨ01ｨ ｨｬｨｨ ｨｨｨｪ01ｨｨｨｦ09ｨｺ060006 07060109ｨｨ01ｨ 09 08ｨｨｨｰｨ 02 09 4 08ｨ 04ｨ ｨ 0605ｨｹ0302 04ｨ ｨｪｨｨｨｬｨ 02ｨｬ0401 090608ｨｰｨ ｨｬｨｨ ｨｰ060006 0302 07060109ｨｨ01ｨ (14). 0609070605ｨｹ040609ｨ ｨｰｨｹ 090902080109040906ｨｺｨｨｨｦ 0002ｨｰ02080604ｨｨ09 ｨｬ020307060109ｨｨ0106090401 00ｨｨｨ 08ｨｨ010609, 0601ｨｪｨ ｨｺ06, ｨｬ0603ｨｪ06, 020905ｨｨ 0906090502ｨｺｨ ｨｰｨｹ 09 09020502ｨｺ02ｨｨ06 090608ｨｰｨ 09 0002ｨｪｨ ｨｬｨｨ 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ (WC), ｨｺ06ｨｰ06080402 06ｨ 02090702 ﾂｨｨ09ｨ 06ｨｰ 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｹ 00ｨｨｨ 08ｨｨ010609 ｨ 06050202 70 % 01ｨ 0302 0708ｨｨ ｨｬ020307060109ｨｨ01060906ｨｦ 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨｨｨ (15). 03060109ｨｨ01 japonica ｨｪ020902ｨｰ ｨ 05050205ｨｹ S5 J, indica ｪ S5 i, 06ｨ 08ｨ 040204 09 0002ｨｪ06ｨｬ 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ ｪ ｨ 05050205ｨｹ S5 n. 0102ｨｰ02080604ｨｨ0006ｨｰｨｪ0402 000608ｨｬ04 S5 J /S5 i 09ｨｰ0208ｨｨ05ｨｹｨｪ04, 06ｨ 08ｨ 040204, ｨｪ0209ｨｮｨｴｨｨ02 0002ｨｪ04 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ, 000208ｨｰｨｨ05ｨｹｨｪ04 (S5 J /S5 n, S5 i /S5 n ) (16, 17). 00ｨｪ02010802ｨｪｨｨ02 0002ｨｪ0609 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ 09 000608ｨｬ04 09 09040906ｨｺ06ｨｦ ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨ06ｨｪｨｪ06ｨｦ 0907060906ｨ ｨｪ0609ｨｰｨｹ06 01ｨ 09ｨｰ 090604ｨｬ0603ｨｪ0609ｨｰｨｹ 070605ｨｮ ﾂｨｨｨｰｨｹ 000208ｨｰｨｨ05ｨｹｨｪ0402 ｨｬ020307060109ｨｨ0106090402 00ｨｨｨ 08ｨｨ0104 09 090205ｨｨ ﾂｨｨｨｪ06ｨｦ 0002ｨｰ02080604ｨｨ09ｨ 0106 100 % (18). 080806ｨｬ02 ｨｰ060006, ｨｬ020307060109ｨｨ010609ｨ 07 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨ07 070604090605ｨｨｨｰ 08ｨ 0903ｨｨ08ｨｨｨｰｨｹ ｨｪ0608ｨｬｨｮ 0802ｨ ｨｺ02ｨｨｨｨ 09060401ｨ 09ｨ 02ｨｬ060006 06ｨ 08ｨ 0402ｨ 04ｨ 09 ﾂ02ｨｰ ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ07 0002ｨｪ02ｨｰｨｨ ﾂ0209ｨｺ060006 0706ｨｰ02ｨｪ02ｨｨｨ 05ｨ 06ｨ 06ｨｨ01 07060109ｨｨ010609 ｨｨ, 090502010609ｨ ｨｰ0205ｨｹｨｪ06, 06ｨ 02090702 ﾂｨｨｨｰ ｨ 0605ｨｹ03ｨｮ06 09ｨｰｨ ｨ ｨｨ05ｨｹｨｪ0609ｨｰｨｹ 070605ｨｮ ﾂｨ 02ｨｬ0401 ｨｮ080603ｨ 0209 08ｨｨ09ｨ. 0102ｨｪ04, 06ｨ ｨｮ0905060905ｨｨ09ｨ 06ｨｴｨｨ02 03ｨｨ0806ｨｺｨｮ06 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｹ, ｨｺｨ 08ｨｰｨｨ080609ｨ ｨｪ04, 0607080201020502ｨｪ 0907ｨｨ0906ｨｺ ｨｬｨｨｨｺ080609ｨ ｨｰ020505ｨｨｨｰｨｪ0401 SSR-ｨｬｨ 08ｨｺ02080609 (simple sequence repeat), 0706040906050706ｨｴｨｨ01 0609ｨｮｨｴ0209ｨｰ090507ｨｰｨｹ ｨｨｨｪｨｰ08060008020909ｨｨ06 05ｨｰｨｨ01 0002ｨｪ0609 09 070208090702ｨｺｨｰｨｨ09ｨｪ0402 090608ｨｰｨ (19-22). 0201ｨｪｨ ｨｺ06 090407090502ｨｪｨｪ0402 0106ｨｪ060804 0002ｨｪ0609 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ 09 ｨ 0605ｨｹ03ｨｨｨｪ09ｨｰ0902 0905ｨｮ ﾂｨ 0209 0708020109ｨｰｨ 090502ｨｪ04 06ｨ 08ｨ 0402ｨ ｨｬｨｨ ｨｨ04 ｨｰ080607ｨｨ ﾂ0209ｨｺ060006 080200ｨｨ06ｨｪｨ (ｨｪｨ 0708ｨｨｨｬ0208, 02ｨｮ05ｨ 08, 01ｨ 00ｨｨｨｪｨ 22) ｨｨ ｨｪ0209ｨｮｨｰ 080701 0708ｨｨ04ｨｪｨ ｨｺ0609, 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ0401 010507 01ｨｨｨｺｨｨ01 09ｨｨ010609 (06090407ｨ 02ｨｬ0609ｨｰｨｹ, ｨｪｨｨ04ｨｺｨ 07 07080601ｨｮｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ, 09ｨｺ0506ｨｪｨｪ0609ｨｰｨｹ ｨｺ 0706050200ｨ ｨｪｨｨ06, 09040906ｨｺ06080609050609ｨｰｨｹ ｨｨ ｨｰ.01.) 00 ｨｰ06 0302 090802ｨｬ07 ｨｨｨｬ0206ｨｰ0907 090606ｨ ｨｴ02ｨｪｨｨ07 06 090407090502ｨｪｨｨｨｨ 0002ｨｪ0609 WC ｨｮ 0906090802ｨｬ02ｨｪｨｪ0401 090608ｨｰ0609 (23). 000205ｨｹ ｨｪｨ 09ｨｰ0607ｨｴ02ｨｦ 08ｨ ｨ 06ｨｰ04 04ｨ ｨｺ0506 ﾂｨ 05ｨ 09ｨｹ 09 ｨｮｨｰ06 ﾂｨｪ02ｨｪｨｨｨｨ ｨｰｨ ｨｺ0906ｨｪ06ｨｬｨｨ ﾂ0209ｨｺ06ｨｦ 0708ｨｨｨｪｨ 01050203ｨｪ0609ｨｰｨｨ ｨｨ04ｨｮ ﾂｨ 02ｨｬ0401 06ｨ 08ｨ 04020609 08ｨｨ09ｨ ｨｺ 0109ｨｮｨｬ 07060109ｨｨ01ｨ ｨｬ, 090407090502ｨｪｨｨｨｨ ｨｮ ｨｪｨｨ01 0002ｨｪ0609 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ ｨｨ 0908ｨ 09ｨｪ02ｨｪｨｨｨｨ 090906ｨｦ09ｨｰ09 070605ｨｮ ﾂｨ 02ｨｬ0401 00ｨｨｨ 08ｨｨ010609 ｨｨ ｨｨ01 080601ｨｨｨｰ020502ｨｦ 09 09090704ｨｨ 09 070208090702ｨｺｨｰｨｨ0906ｨｦ 09060401ｨ ｨｪｨｨ07 000208ｨｰｨｨ05ｨｹｨｪ0401 ｨｬ020307060109ｨｨ0106090401 0002ｨｰ02080604ｨｨ09ｨｪ0401 00ｨｨｨ 08ｨｨ010609. 0002ｨｰ0601ｨｨｨｺｨ. 0004 070806090205ｨｨ 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨ06 ｨｬ020301ｨｮ 06ｨ 08ｨ 0402ｨ ｨｬｨｨ 08ｨｨ09ｨ Oryza sativa L. 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 06ｨｰｨｪ060907ｨｴｨｨｨｬｨｨ0907 ｨｺ 07060109ｨｨ01ｨｮ japonica (09ｨ 04ｨ 08, 040200ｨｮ05, 09ｨｨｨｬｨ ｨｪ, 0505ｨ 0907ｨｪ0202, 0208ｨｮ03ｨｪ04ｨｦ, 0001060604 7679, 0001060604 7653 ｨｨ ｨｰ.01.), ｨ ｨｰｨ ｨｺ0302 06ｨ 08ｨ 0402ｨ ｨｬｨｨ 07060109ｨｨ01ｨ indica 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ06ｨｦ ｨｨ 04ｨ 08ｨｮｨ 0203ｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ. 020202ｨｪｨｺｨｮ 00ｨｨｨ 08ｨｨ010609 070806090601ｨｨ05ｨｨ 0708ｨｨ 090408ｨ ｨｴｨｨ09ｨ ｨｪｨｨｨｨ 09 090609ｨｮ01ｨ 01 ｨｪｨ 09020002ｨｰｨ 02ｨｨ06ｨｪｨｪ06ｨｦ 070506ｨｴｨ 01ｨｺ02 (10 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨｪｨ 090609ｨｮ01) ｨｪｨ 0607ｨｰｨｨｨｬｨ 05ｨｹｨｪ06ｨｬ 0006ｨｪ02 ｨｬｨｨｨｪ0208ｨ 05ｨｹｨｪ060006 07ｨｨｨｰｨ ｨｪｨｨ07 N 120 P 60 K 60. 00 0904ｨ 0608ｨｺｨ 01 (20 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨｪｨ 090608ｨｰ ｨｨ 00ｨｨｨ 08ｨｨ01ｨｪｨｮ06 ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨ06) 09 00ｨ 04ｨｮ 090604080209ｨ ｨｪｨｨ07 070806090601ｨｨ05ｨｨ ｨ ｨｨ06ｨｬ02ｨｰ08ｨｨ ﾂ0209ｨｺｨｨｨｦ ｨ ｨｪｨ 05ｨｨ04. 040601ｨｨｨｰ0205ｨｹ09ｨｺｨｨ02 000608ｨｬ04, 00ｨｨｨ 08ｨｨ01ｨｪ0402 ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨｨｨ ｨｺ06ｨｰ06080401 0706ｨｺｨ 040409ｨ 05ｨｨ 09ｨｰ0208ｨｨ05ｨｹｨｪ0609ｨｰｨｹ ｨ 06050202 30 %, 06ｨｰｨｪ0609ｨｨ05ｨｨ ｨｺ ｨｪ02 06ｨ 05ｨ 01ｨ 06ｨｴｨｨｨｬ 0002ｨｪｨ ｨｬｨｨ 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ. 03ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ 060708020102050705ｨｨ ｨｺｨ ｨｺ 0708060202ｨｪｨｰｨｪ0602 06ｨｰｨｪ060302ｨｪｨｨ02 ﾂｨｨ0905ｨ 09ｨｰ0208ｨｨ05ｨｹｨｪ0401 ｨｺ06050609ｨｺ0609 ｨｺ ｨｨ01 06ｨ ｨｴ02ｨｬｨｮ ﾂｨｨ0905ｨｮ. 00060502ｨｺｨｮ050708ｨｪ04ｨｦ ｨ ｨｪｨ 05ｨｨ04 0904070605ｨｪ0705ｨｨ ｨｪｨ 20 08ｨ 09ｨｰ02ｨｪｨｨ0701 ｨｺｨ 0301060006 06ｨ 08ｨ 0402ｨ 09 ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ02ｨｬ SSR-ｨｬｨ 08ｨｺ02080609, 070605ｨｨｨｬ060800ｨｪ0401 010507 ｨｰｨｨ07ｨｨ ﾂｨｪ0401 06ｨ 08ｨ 04020609 07060109ｨｨ010609. 0308ｨｨ ｨｬ060502ｨｺｨｮ050708ｨｪ06ｨｬ ｨｬｨ 08ｨｺｨｨ080609ｨ ｨｪｨｨｨｨ ｨｨ09070605ｨｹ040609ｨ 05ｨｨ 090608ｨｰｨ 09ｨ ｨｪｨｺｨ ｨｦ09ｨｺｨｨｨｦ, 05ｨ 01ｨｺ06, 0308ｨｨｨｬ060809ｨｺｨｨｨｦ, 09ｨｨｨｬｨ ｨｪ, 01ｨ 08ｨ ｨｪｨｰ, 03ｨ 0905060909ｨｺｨｨｨｦ, 04ｨ 07ｨ ｨｪ, 010609ｨ ｨｰ0608, 05020807ｨ ｨｪｨｰｨｨｨｪ, 09060708ｨｨｨｪ, 040200ｨｮ05, 08ｨｪｨｰｨ 08ｨｹ, 0402ｨｬ ﾂｨｮ00, 09ｨｨ010208, 09ｨ 04ｨ 08, 08ｨｬ02ｨｰｨｨ09ｨｰ, 01ｨ 0802ｨｨ0909, 0208ｨｮ03ｨｪ04ｨｦ, 050708ｨｨｨｪｨｰ, 00ｨｨ0605ｨ, 02ｨ 05ｨｹｨｪ02090609ｨｰ06 ﾂｨｪ04ｨｦ, 0806ｨｪｨｰｨ ｨｪ, 08ｨ 09ｨｮｨｪ, 0707ｨｨ- 62
13ｨｰ0208, 08ｨｰ05ｨ ｨｪｨｰ, 08ｨｮ08 ﾂｨ ｨｪｨｺｨ, 08ｨ ｨｺ0205, 05ｨｪ0203ｨｨｨｪｨｺｨ, 01ｨ 08ｨｬ, 08ｨｪｨ ｨｨｨｰ, 0805ｨ 00ｨｬｨ ｨｪ, 0604ｨｮｨｬ08ｨｮ01, IR 66, NSIC RC 158, IR 68897B (070609050201ｨｪｨｨ02 ｨｰ08ｨｨ ｪ ｨｰｨｨ07ｨｨ ﾂｨｪ0402 06ｨ 08ｨ 040204 07060109ｨｨ01ｨ indica 09020502ｨｺ02ｨｨｨｨ IRRI (IRRI ｪ International Rice Research Institute, 08ｨｨ05ｨｨ0707ｨｨｨｪ04). 05080201ｨｨ 090608ｨｰ0609 08060909ｨｨｨｦ09ｨｺ06ｨｦ 09020502ｨｺ02ｨｨｨｨ 07060109ｨｨ01 indica ｨ 0405 0708020109ｨｰｨ 090502ｨｪ 090608ｨｰｨ ｨｬｨｨ 05020807ｨ ｨｪｨｰｨｨｨｪ, 01ｨ 0802ｨｨ0909 05ｨｪ0203ｨｨｨｪｨｺｨ, 01ｨ 08ｨｬ, 0604ｨｮｨｬ08ｨｮ01. 020108 09040102050705ｨｨ ｨｨ04 05ｨｰｨｨ0605ｨｨ080609ｨ ｨｪｨｪ0401 070806080609ｨｰｨｺ0609 ｨｨ 05ｨｨ09ｨｰｨｹ0209 09 0706ｨｬ06ｨｴｨｹ06 STAB-ｨｬ02ｨｰ0601ｨ 09 08ｨ 0405ｨｨ ﾂｨｪ0401 ｨｬ0601ｨｨ00ｨｨｨｺｨ 02ｨｨ0701, 070609ｨｰｨ ｨｪ0609ｨｺｨｮ 070605ｨｨｨｬ0208ｨ 04ｨｪ06ｨｦ 020207ｨｪ06ｨｦ 0802ｨ ｨｺ02ｨｨｨｨ (030004) ｨｨ 09ｨｨ04ｨｮｨ 05ｨｨ04ｨ 02ｨｨ06 07080601ｨｮｨｺｨｰ0609 ｨ ｨｬ0705ｨｨ00ｨｨｨｺｨ 02ｨｨｨｨ 070806090601ｨｨ05ｨｨ 0706 ｨｬ02ｨｰ0601ｨｨｨｺ02 IRRI (24). 05ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺｨｮ06 06ｨ 08ｨ ｨ 06ｨｰｨｺｨｮ 01ｨ ｨｪｨｪ0401 0609ｨｮｨｴ0209ｨｰ09050705ｨｨ 09 0708060008ｨ ｨｬｨｬ02 Statistica v. 6.0. 1. 02040208ｨｪ02ｨｪｨｪ0609ｨｰｨｹ 00ｨｨｨ 08ｨｨ010609 07020809060006 0706ｨｺ060502ｨｪｨｨ07 0708ｨｨ 09ｨｺ0802ｨｴｨｨ09ｨ ｨｪｨｨｨｨ 090608ｨｰ0609 08ｨｨ09ｨ (Oryza sa-tiva L.) 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 06ｨｰｨｪ0609ｨｨｨｬ0401 ｨｺ 07060109ｨｨ01ｨ ｨｬ japonica ｨｨ indica 0706 ｨｨｨｪ0102ｨｺ09ｨｮ 040208ｨｪ0609ｨｺｨｨ (09020002ｨｰｨ 02ｨｨ06ｨｪｨｪｨ 07 070506ｨｴｨ 01ｨｺｨ, 2008-2009 00060104) 00ｨ ｨｰ0208ｨｨｨｪ09ｨｺｨ 07 000608ｨｬｨ 05 06070405ｨｨｨｰ0205ｨｹ 03ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ, % japonica 05 indica: 09ｨ 04ｨ 08 05 05ｨｪ0203ｨｨｨｪｨｺｨ 5,76 09ｨ 04ｨ 08 05 0604ｨｮｨｬ08ｨｮ01 18,13 02ｨｨｨ ｨｪｨ 05 00ｨ ｨｦ07 14,04 02ｨｨｨ ｨｪｨ 05 05ｨｪ0203ｨｨｨｪｨｺｨ 9,09 0208ｨｮ03ｨｪ04ｨｦ 05 05ｨｪ0203ｨｨｨｪｨｺｨ 20,70 indica 05 japonica: 05ｨｪ0203ｨｨｨｪｨｺｨ 05 09ｨ 04ｨ 08 12,57 05ｨｪ0203ｨｨｨｪｨｺｨ 05 040200ｨｮ05 9,90 00ｨ ｨｦ07 05 09ｨ 04ｨ 08 6,75 0604ｨｮｨｬ08ｨｮ01 05 09ｨ 04ｨ 08 14,25 japonica 05 japonica: 09ｨ 04ｨ 08 05 09ｨｨｨｬｨ ｨｪ 3,60 040200ｨｮ05 05 0505ｨ 0907ｨｪ0202 30,40 0208ｨｮ03ｨｪ04ｨｦ 05 09ｨｨｨｬｨ ｨｪ 20,00 02ｨｨｨ ｨｪｨ 05 09ｨｨｨｬｨ ｨｪ 19,40 0001060604 7653 05 0208ｨｮ03ｨｪ04ｨｦ 17,20 040204ｨｮ05ｨｹｨｰｨ ｨｰ04. 0004 070806090205ｨｨ 0706ｨｨ09ｨｺ 0002ｨｪ0609 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ (WC) 09080201ｨｨ ｨｺ06050502ｨｺ02ｨｨ06ｨｪｨｪ0401 06ｨ 08ｨ 04020609 08ｨｨ09ｨ. 040601ｨｨｨｰ0205ｨｹ09ｨｺｨｨ02 000608ｨｬ04, 00ｨｨｨ 08ｨｨ01ｨｪ0402 ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨｨｨ ｨｺ06ｨｰ06080401 0708ｨｨ ｨｬ020307060109ｨｨ01060906ｨｦ 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨｨｨ 0706ｨｺｨ 040409ｨ 05ｨｨ 07ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ ｨ 06050202 30 %, 06ｨｰｨｪ0609ｨｨ05ｨｨ ｨｺ ｨｪ02 06ｨ 05ｨ 01ｨ 06ｨｴｨｨｨｬ 0002ｨｪｨ ｨｬｨｨ 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ (ｨｰｨ ｨ 05. 1). 0508ｨ 09ｨｪ02ｨｪｨｨ02 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｨ 00ｨｨｨ 08ｨｨ010609 06ｨｰ 09ｨｺ0802ｨｴｨｨ09ｨ ｨｪｨｨ07 06ｨ 08ｨ 04020609 08060909ｨｨｨｦ09ｨｺ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, ｨｺ06ｨｰ06080402 08ｨ ｨｪ0202 ｨ 0405ｨｨ 06ｨｰｨｪ020902ｨｪ04 ｨｺ 08ｨ 04ｨｪ04ｨｬ 07060109ｨｨ01ｨ ｨｬ ｨｪｨ 0609ｨｪ0609ｨ ｨｪｨｨｨｨ 08ｨ 0405ｨｨ ﾂｨｨｨｦ 0706 ｨｨｨｪ0102ｨｺ09ｨｮ 040208ｨｪ0609ｨｺｨｨ (06ｨｰｨｪ060302ｨｪｨｨ02 0202 0105ｨｨｨｪ04 ｨｺ 03ｨｨ08ｨｨｨｪ02), 0706ｨｺｨ 04ｨ 0506 ｨｪｨｨ04ｨｺｨｮ06 07ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ, ｨｪ02 06ｨｰ05ｨｨ ﾂｨ 06ｨｴｨｮ060907 010609ｨｰ06090208ｨｪ06 06ｨｰ ｨｰｨ ｨｺ060906ｨｦ ｨｮ 00ｨｨｨ 08ｨｨ010609 0601ｨｪ060006 07060109ｨｨ01ｨ (09ｨｬ. ｨｰｨ ｨ 05. 1.). 0701ｨｨ09ｨｨ0506 ｨｰ06, ﾂｨｰ06 0708ｨｨ 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨｨｨ 08060909ｨｨｨｦ09ｨｺｨｨ01 06ｨ 08ｨ 04020609 0601ｨｪ060006 07060109ｨｨ01ｨ 07ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ 0906 ｨｬｨｪ0600ｨｨ01 0905ｨｮ ﾂｨ 0701 070802090403ｨ 05ｨ ｨｰｨ ｨｺ0609ｨｮ06 ｨｮ ｨｬ020307060109ｨｨ0106090401 00ｨｨｨ 08ｨｨ010609. 0004090601 ｨｨ04 05ｨｰ060006 ｨｬ060302ｨｰ ｨ 04ｨｰｨｹ ｨｰ0605ｨｹｨｺ06 0601ｨｨｨｪ: ｨｨ05ｨｨ ｨｬｨｪ0600ｨｨ02 08060909ｨｨｨｦ09ｨｺｨｨ02 090608ｨｰｨ ｨｪ0209ｨｮｨｰ 0002ｨｪ04 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ, ｨｨ05ｨｨ 0708ｨｨｨｪ07ｨｰｨ 07 09 04060909ｨｨｨｨ ｨｺ05ｨ 0909ｨｨ00ｨｨｨｺｨ 02ｨｨ07 ｨｪ02 0706040906050702ｨｰ ｨｺ06080802ｨｺｨｰｨｪ06 06ｨｰｨｪ0209ｨｰｨｨ 000608ｨｬ04 ｨｺ 07060109ｨｨ01ｨ ｨｬ. 08ｨｪｨ 05ｨｨ04 070806ｨｨ090106030102ｨｪｨｨ07 09020502ｨｺ02ｨｨ06ｨｪｨｪ060006 ｨｬｨ ｨｰ0208ｨｨｨ 05ｨ 09040709ｨｨ05 010609ｨｰｨ ｨｰ06 ﾂｨｪ06 ｨ 05ｨｨ04ｨｺ0602 08060109ｨｰ0906 06ｨ 08ｨ 04020609 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 09 ｨｰ06ｨｬ ﾂｨｨ090502 ｨｪ06ｨｬｨｨｨｪｨｨ08ｨｮ02ｨｬ0401 ｨｺｨ ｨｺ 08ｨ 04ｨｪ0402 07060109ｨｨ0104. 04ｨ ｨｪ0202 06ｨ 08ｨ 040204 08ｨｨ09ｨ ｨｬ0603ｨｪ06 ｨ 040506 08ｨ 0401020507ｨｰｨｹ ｨｪｨ 07060109ｨｨ0104 09060005ｨ 09ｨｪ06 070806ｨｨ090106030102ｨｪｨｨ06 ｨｨ 000608ｨｬ02 040208ｨｪ0609ｨｺｨｨ, 0601ｨｪｨ ｨｺ06 03ｨｨ0806ｨｺｨ 07 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨ07 07060109ｨｨ010609 09 08070102 ｨｨ09090502010609ｨ ｨｰ0205ｨｹ09ｨｺｨｨ01 0202ｨｪｨｰ080609 0708ｨｨ090205ｨ ｨｺ 070607090502ｨｪｨｨ06 ｨ 0605ｨｹ03060006 ﾂｨｨ0905ｨ 000608ｨｬ, ｨｺ06ｨｰ06080402 04ｨ 09 ﾂ02ｨｰ 0802ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨｨｨ ｨｪ0209ｨｮｨｰ 0002ｨｪ04, 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ0402 010507 08ｨ 04ｨｪ0401 07060109ｨｨ010609 (25-27). 0306 ｨｪｨ ｨ 05060102ｨｪｨｨ07ｨｬ ｨｬｨｪ0600ｨｨ01 ｨｮ ﾂ02ｨｪ0401, 09 ｨｪｨ 09ｨｰ0607ｨｴ0202 090802ｨｬ07 ｨｪ02090604ｨｬ0603ｨｪ06 ｨｪｨ 0609ｨｪ0609ｨ ｨｪｨｨｨｨ 08ｨ 0405ｨｨ ﾂｨｨｨｦ 0706 0601ｨｪ06ｨｬｨｮ 0708ｨｨ04ｨｪｨ ｨｺｨｮ ｨｨ05ｨｨ 0002ｨｪｨｮ 010609ｨｰ06090208ｨｪ06 01ｨｨ09ｨｺ08ｨｨｨｬｨｨｨｪｨｨ080609ｨ ｨｰｨｹ 07060109ｨｨ0104 indica ｨｨ japonica (28), ｨｨ01 08ｨ 0401020502ｨｪｨｨ02 ｨｰ0605ｨｹｨｺ06 0706 ｨｨｨｪ0102ｨｺ09ｨｮ 040208ｨｪ0609ｨｺｨｨ ｨ 06050202 ﾂ02ｨｬ 09 39 % 0905ｨｮ ﾂｨ 0209 0708ｨｨ090601ｨｨｨｰ ｨｺ ｨｪ020708ｨ 09ｨｨ05ｨｹｨｪ06ｨｦ ｨｺ05ｨ 0909ｨｨ00ｨｨｨｺｨ 02ｨｨｨｨ. 08ｨｰｨｰ08ｨｨｨ ｨｮｨｰｨｨ080609ｨ ｨｪｨｨ02 0706 0105ｨｨｨｪ02 0607ｨｮ0302ｨｪｨｨ07 020902ｨｰｨｺ06090401 ﾂ0203ｨｮｨｦ ｨｰｨ ｨｺ0302 01ｨ 02ｨｰ 0603ｨｨｨ ｨｺｨｮ 09 20-40 % 09ｨ 08ｨｨｨ ｨｪｨｰ0609. 020507 ｨ 06050202 ｨｰ06 ﾂｨｪ06ｨｦ 01ｨｨ0000020802ｨｪ02ｨｨｨ 02ｨｨｨｨ 0708020105060302ｨｪ06 ｨｮ ﾂｨｨｨｰ0409ｨ ｨｰｨｹ ｨｪ0209ｨｺ0605ｨｹｨｺ06 0708ｨｨ04ｨｪｨ ｨｺ0609: 06ｨｺ08ｨ 09ｨｺｨｮ 05ｨｨ09ｨｰｨｹ0209, 0002ｨｪ0605ｨｹｨｪｨｮ06 0802ｨ ｨｺ02ｨｨ06, 0105ｨｨｨｪｨｮ 0607ｨｮ0302ｨｪｨｨ07 020902ｨｰｨｺ06090401 ﾂ0203ｨｮｨｦ (ｨｨ05ｨｨ 0105ｨｨｨｪｨｮ 0906050609ｨｺ0609 ｨ 07ｨｨｨｺｨｮ050609ｨ ), ﾂｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｹ ｨｺ ｨｪｨｨ04ｨｺｨｨｨｬ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ08ｨ ｨｬ, 01050608ｨ ｨｰｨｮ ｨｺｨ 05ｨｨ07 (ｨｰｨ ｨ 05. 2). 0308ｨｨ 63
13ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨｨｨ 0904030207020802 ﾂｨｨ090502ｨｪｨｪ06ｨｦ ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨｨｨ 0708ｨｨ04ｨｪｨ ｨｺ0609 070600080203ｨｪ0609ｨｰｨｹ ｨｺ05ｨ 0909ｨｨ00ｨｨｨｺｨ 02ｨｨｨｨ ｨｪ02 070802090403ｨ 02ｨｰ 0,5 % (29-31). 07ｨｺｨ 04ｨ ｨｪｨｪ0402 09 05ｨｰ06ｨｦ ｨｰｨ ｨ 05ｨｨ0202 0708ｨｨ04ｨｪｨ ｨｺｨｨ 08ｨ 09070605060302ｨｪ04 09 0706080701ｨｺ02 ｨｨ01 0708ｨｨｨｬ02ｨｪｨｨｨｬ0609ｨｰｨｨ 010507 01ｨｨ09ｨｺ08ｨｨｨｬｨｨｨｪｨ 02ｨｨｨｨ 06ｨ 08ｨ 04020609. 06ｨ ｨｺ, ｨｮ09ｨｰ06ｨｦ ﾂｨｨ090609ｨｰｨｹ ｨｺ 01050608ｨ ｨｰｨｮ ｨｺｨ 05ｨｨ07 0706040906050702ｨｰ ｨ 06050202 ｨｰ06 ﾂｨｪ06 01ｨｨ0000020802ｨｪ02ｨｨ080609ｨ ｨｰｨｹ 06ｨ 08ｨ 040204 ｨｪｨ 07060109ｨｨ0104. 020507 090608ｨｰ0609 ｨｨｨｪ01ｨｨｨｦ09ｨｺ060006 07060109ｨｨ01ｨ 09 ｨ 0605ｨｹ03ｨｨｨｪ09ｨｰ0902 0905ｨｮ ﾂｨ 0209 01ｨ 08ｨ ｨｺｨｰ020802ｨｪ 070604ｨｨｨｰｨｨ09ｨｪ04ｨｦ 06ｨｰ0902ｨｰ ｨｪｨ 0002ｨｪ0605ｨｹｨｪｨｮ06 0802ｨ ｨｺ02ｨｨ06 (32, 33). 0906ｨｰ07 090702ｨｺｨｰ08 09ｨ 08ｨｨｨ 02ｨｨｨｨ 0706 ｨｺｨ 030106ｨｬｨｮ 0708ｨｨ04ｨｪｨ ｨｺｨｮ 010507 07060109ｨｨ010609 ﾂｨ 09ｨｰｨｨ ﾂｨｪ06 07020802ｨｺ080409ｨ 02ｨｰ0907, 090604ｨｬ0603ｨｪ0609ｨｰｨｹ 01ｨｨ0000020802ｨｪ02ｨｨｨ 02ｨｨｨｨ 090902 0302 ｨｮｨｺｨ 040409ｨ 02ｨｰ ｨｪｨ ｨｪｨ 05ｨｨ- ﾂｨｨ02 ｨｮ ｨｪｨｨ01 04ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ0401 ｨｨ 08ｨ 04ｨｪ06ｨｪｨ 0708ｨ 090502ｨｪｨｪ0401 ｨｬ0601ｨｨ00ｨｨｨｺｨ 02ｨｨｨｦ. 64 2. 0308ｨｨ04ｨｪｨ ｨｺｨｨ, 01ｨｨ0000020802ｨｪ02ｨｨ08ｨｮ06ｨｴｨｨ02 07060109ｨｨ0104 indica ｨｨ japonica ｨｮ 08ｨｨ09ｨ (Oryza sativa L.) (33) 0308ｨｨ04ｨｪｨ ｨｺ indica (n = 42) japonica (n = 47) 0709ｨｰ06ｨｦ ﾂｨｨ090609ｨｰｨｹ ｨｺ 01050608ｨ ｨｰｨｮ ｨｺｨ 05ｨｨ07 (log 07060908020301ｨ 06ｨｴ02ｨｦ ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨｨｨ) 01ｨｨ04ｨｺｨ 07 (3,34 ﾀ0,60) 00040906ｨｺｨ 07 (6ﾓ11,49 ﾀ0,35) 0709ｨｰ06ｨｦ ﾂｨｨ090609ｨｰｨｹ ｨｺ 04ｨ 09ｨｮ0102 (090403ｨｨ09ｨ ｨｪｨｨ02, %) 00040906ｨｺｨ 07 (92,4 ﾀ8,75) 01ｨｨ04ｨｺｨ 07 (59,3 ﾀ20,67) 0205ｨｨｨｪｨ 0906050609ｨｺ0609 ｨ 07ｨｨｨｺｨｮ050609ｨ (ｨｬｨｬ) 08060806ｨｰｨｺｨｨ02 (0,26 ﾀ0,06) 0205ｨｨｨｪｨｪ0402 (0,44 ﾀ0,22) 0709ｨｰ06ｨｦ ﾂｨｨ090609ｨｰｨｹ ｨｺ 0106050601ｨｮ (%) ﾁｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ04ｨｦ (0,58 ﾀ0,18) 0709ｨｰ06ｨｦ ﾂｨｨ0904ｨｦ (0,18 ﾀ0,22) 0802ｨｪ0605ｨｹｨｪｨ 07 0802ｨ ｨｺ02ｨｨ07 030604ｨｨｨｰｨｨ09ｨｪｨ 07 010200ｨ ｨｰｨｨ09ｨｪｨ 07 03ｨｴ02 0601ｨｨｨｪ 09ｨ 08ｨｨｨ ｨｪｨｰ ｨ ｨｰｨｰ08ｨｨｨ ｨｮｨｰｨｨ080609ｨ ｨｪｨｨ07 0708020105060302ｨｪ ｨｺｨｨｨｰｨ ｨｦ09ｨｺｨｨｨｬｨｨ ｨｮ ﾂ02ｨｪ04ｨｬｨｨ, ｨｺ06ｨｰ06080402 09 ﾂｨｨｨｰｨ 06ｨｰ 02020502090606ｨ 08ｨ 04ｨｪ04ｨｬ 09 07020802 ﾂ02ｨｪｨｹ 0706ｨｺｨ 04ｨ ｨｰ020502ｨｦ, 0706040906050706ｨｴｨｨ01 010609ｨｰ06090208ｨｪ06 08ｨ 04010205ｨｨｨｰｨｹ 06ｨ 08ｨ 040204 08ｨｨ09ｨ ｨｪｨ 07060109ｨｨ0104, 09ｨｺ0506 ﾂｨｨｨｰｨｹ (09 0106070605ｨｪ02ｨｪｨｨ02 ｨｺ ｨｮ0302 07020802 ﾂｨｨ090502ｨｪｨｪ04ｨｬ) 06ｨｰｨｪ060302ｨｪｨｨ02 ﾂｨｨ0905ｨ ｨ 0605ｨｹ03ｨｨ01 090609ｨｮ01ｨｨ09ｨｰ0401 07ｨｮ ﾂｨｺ0609 ｨｺ ﾂｨｨ0905ｨｮ ｨｬｨ 050401 09 ｨｪ0603ｨｺ02 ｨｬ02ｨｰ0205ｨｺｨｨ. 07 07060109ｨｨ01ｨ indica ｨｮｨｺｨ 04ｨ ｨｪｨｪ04ｨｦ ｨｨｨｪ0102ｨｺ09 0708ｨｨｨ 05ｨｨ04ｨｨｨｰ0205ｨｹｨｪ06 08ｨ 0902ｨｪ 1, ｨｮ japonica ｪ 0,5 (18). 05050201ｨｮ02ｨｰ 06ｨｰｨｬ02ｨｰｨｨｨｰｨｹ, ﾂｨｰ06 01ｨ 0302 ｨｰ0209ｨｰ ｨｪｨ 09ｨｰ0208ｨｨ05ｨｹｨｪ0609ｨｰｨｹ 00ｨｨｨ 08ｨｨ010609 ｨｪ02 0909020001ｨ 0706040906050702ｨｰ 0708ｨ 09ｨｨ05ｨｹｨｪ06 ｨｺ05ｨ 0909ｨｨ00ｨｨ02ｨｨ080609ｨ ｨｰｨｹ 06ｨ 08ｨ 040204. 00 ﾂｨ 09ｨｰｨｪ0609ｨｰｨｨ, 09ｨｺ0802ｨｴｨｨ09ｨ ｨｪｨｨ02 06ｨ 08ｨ 04020609, ｨｪ0209ｨｮｨｴｨｨ01 ｨｪ02ｨｺ06ｨｰ06080402 0002ｨｪ04, 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ0402 010507 01ｨｨｨｺｨｨ01 ｨｨ 07080201ｨｺ06090401 000608ｨｬ, 09 0906090802ｨｬ02ｨｪｨｪ04ｨｬｨｨ 090608ｨｰｨ ｨｬｨｨ ｨｰｨ ｨｺ0302 0708ｨｨ090601ｨｨｨｰ ｨｺ ﾂｨ 09ｨｰｨｨ ﾂｨｪ06ｨｦ 09ｨｰ0208ｨｨ05ｨｹｨｪ0609ｨｰｨｨ 00ｨｨｨ 08ｨｨ010609 (2). 3. 02040208ｨｪ02ｨｪｨｪ0609ｨｰｨｹ ｨｮ 00ｨｨｨ 08ｨｨ010609 08ｨｨ09ｨ (Oryza sativa L.) 07020809060006 0706ｨｺ060502ｨｪｨｨ07 0708ｨｨ 09ｨｺ0802ｨｴｨｨ09ｨ ｨｪｨｨｨｨ 080601ｨｨｨｰ0205ｨｹ09ｨｺｨｨ01 000608ｨｬ 07060109ｨｨ010609 japonica (j) ｨｨ indica (i) ｨｨ04 08ｨ 0405ｨｨ ﾂｨｪ0401 05ｨｺ0605060006-0002060008ｨ 00ｨｨ ﾂ0209ｨｺｨｨ01 0008ｨｮ0707 (09020002ｨｰｨ 02ｨｨ06ｨｪｨｪｨ 07 070506ｨｴｨ 01ｨｺｨ, 2008-2009 00060104) 00ｨ ｨｰ0208ｨｨｨｪ09ｨｺｨ 07 000608ｨｬｨ 05 06070405ｨｨｨｰ0205ｨｹ 03ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ, % japonica 05 indica: 09ｨ 04ｨ 08 05 0205ｨｨｨｬ07ｨｨｨ 01ｨ 50,90 0208ｨｮ03ｨｪ04ｨｦ 05 0205ｨｨｨｬ07ｨｨｨ 01ｨ 65,10 0001060604 7679 05 03ｨ 08ｨｨｨｰ02ｨｰ 59,64 0206ｨｪ00 0106ｨｪ00 418 (japonica ｨｰ080607ｨｨ ﾂ0209ｨｺｨ 07) 05 05ｨｰ0802050202 83,98 Vialone Nano 05 08/12 6205 58,27 0402ｨｬ ﾂｨｮ00 05 08/12 6206 54,50 indica/japonica: 08/12 6205 05 0308ｨｨｨｬ060809ｨｺｨｨｨｦ 93,30 08/12 6206 05 08ｨｬ02ｨｰｨｨ09ｨｰ 40,00 08/12 6205 05 Baldo 74,90 08/12 6205 05 08ｨｰ05ｨ ｨｪｨｰ 41,20 08/12 6205 05 0505ｨ 0907ｨｪ0202 20,70 08/12 6205 05 0206ｨｪ ﾂｨ ｨｺ 56,89 08/12 6205 05 09ｨｨｨｬｨ ｨｪ 32,04 Jefferson 05 Arborio 8,02 Jefferson 05 08ｨｬ02ｨｰｨｨ09ｨｰ 6,73 Jefferson 05 00ｨｨ0605ｨ 7,71 Jefferson 05 0806ｨｪｨｰｨ ｨｪ 3,75 00ｨｪｨｮｨｰ08ｨｨ07060109ｨｨ0106090402 00ｨｨｨ 08ｨｨ0104: 0206ｨｪ00 0106ｨｪ00 418 (japonica ｨｰ080607ｨｨ ﾂ0209ｨｺｨ 07) 05 09060708ｨｨｨｪ (j) 46,82 0206ｨｪ00 0106ｨｪ00 418 (japonica ｨｰ080607ｨｨ ﾂ0209ｨｺｨ 07) 05 03ｨ 0905060909ｨｺｨｨｨｦ (j) 32,85 08/12 6205 (i) 05 0604ｨｮｨｬ08ｨｮ01 (i) 89,70 05ｨｪ0203ｨｨｨｪｨｺｨ (i) 05 08CL (i) 30,59 00 07080609020102ｨｪｨｪ06ｨｬ ｨｪｨ ｨｬｨｨ ｨｨ09090502010609ｨ ｨｪｨｨｨｨ 00ｨｨｨ 08ｨｨ010609 09 ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨ0701, 000102 080601ｨｨｨｰ0205ｨｹ09ｨｺｨｨ02 000608ｨｬ04 0708ｨｨｨｪｨ 01050203ｨ 05ｨｨ ｨｺ 08ｨ 0405ｨｨ ﾂｨｪ04ｨｬ 05ｨｺ0605060006-0002060008ｨ 00ｨｨ ﾂ0209ｨｺｨｨｨｬ 0008ｨｮ0707ｨ ｨｬ, ｨ 0405ｨ 06ｨ ｨｪｨ 08ｨｮ0302ｨｪｨ 09040906ｨｺｨ 07 07ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ F 1 (ｨｰｨ ｨ 05. 3). 4. 04ｨ 0907080201020502ｨｪｨｨ02 070605ｨｨｨｬ060800ｨｪ0401 SSR-
13ｨｬｨ 08ｨｺ02080609, 0706040906050706ｨｴｨｨ01 ｨｰｨｨ07ｨｨ080609ｨ ｨｰｨｹ 07060109ｨｨ0104 japonica ｨｨ indica, ｨｪｨ 010806ｨｬ060906ｨｬｨ 01 08ｨｨ09ｨ (Oryza sativa L.) (7) 090806ｨｬ060906ｨｬｨ SSR-ｨｬｨ 08ｨｺ0208 1 RM 226, RM 259, RM 23 2 RM 250, RM 29, RM 211, RM 240 3 RM 231, RM 232, RM 16, RM 251 4 RM 255, RM 335 5 RM 249, RM 267 6 RM 190 7 RM 248, RM 234, RM 214, RM 18 8 RM 25 9 RM 242 10 RM 258, RM 222, RM 228 11 RM 4 12 RM 247, RM 19 030605ｨｮ ﾂ02ｨｪｨｪ0402 01ｨ ｨｪｨｪ0402 0706ｨｺｨ 040409ｨ 06ｨｰ 06ｨｰ09ｨｮｨｰ09ｨｰ09ｨｨ02 0002ｨｪ0609 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ ｨｮ 06ｨ 08ｨ 04020609, 0906090502 ﾂ02ｨｪｨｪ0401 09 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨ06. 060605ｨｹｨｺ06 06ｨ 08ｨ 040202 ｨｨ04 050108 Jefferson 0906 09090201 ｨｺ06ｨｬｨ ｨｨｨｪｨ 02ｨｨ0701 ｨｨｨｬ0205 04ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪｨｮ06 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｹ 00ｨｨｨ 08ｨｨ010609 (09 09ｨ 08ｨｨｨ ｨｪｨｰｨ 01 09 ｨｪｨｨｨｬ 07ｨｮ09ｨｰ06040208ｨｪ0609ｨｰｨｹ 09ｨ 08ｨｹｨｨ080609ｨ 05ｨ 06ｨｰ 3,75 0106 8,00 %). 08ｨｪｨ 05ｨｨ04 0002ｨｪ0204ｨｨ09ｨ 090608ｨｰｨ 070601ｨｰ09020801ｨｨ05, ﾂｨｰ06 0708ｨｨ ﾂｨｨｨｪ06ｨｦ 09040906ｨｺ06ｨｦ 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｨ 00ｨｨｨ 08ｨｨ010609 09 05ｨｰ06ｨｬ 0905ｨｮ ﾂｨ 02 ｨｪ02 ｨｬ060302ｨｰ ｨ 04ｨｰｨｹ ｨ 05ｨｨ04ｨｺ0608060109ｨｰ0902ｨｪｨｪ0602 070806ｨｨ090106030102ｨｪｨｨ02 ｨｨ09010601ｨｪ0401 000608ｨｬ. 02 ｨｰ06ｨｬ 0302 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ090609ｨ 05ｨｨ 04ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ0402 08ｨ 0405ｨｨ ﾂｨｨ07 0706 ｨｬ0608000600ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨｨｬ 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ09ｨｰｨｨｨｺｨ ｨｬ ｨｮ 06ｨ 08ｨ 04020609, 0906090502 ﾂ02ｨｪｨｪ0401 09 00ｨｨｨ 08ｨｨ01ｨｨ04ｨ 02ｨｨ06 09 05ｨｰｨｨｨｬ 090608ｨｰ06ｨｬ. 01ｨ ｨｨｨ 06050202 010609ｨｰ06090208ｨｪ06 08ｨ 04010205ｨｨｨｰｨｹ 06ｨ 08ｨ 040204 ｨｪｨ 07060109ｨｨ0104 ｨｬ0603ｨｪ06 09 0706ｨｬ06ｨｴｨｹ06 ｨｬ060502ｨｺｨｮ050708ｨｪ060006 ｨｬｨ 08ｨｺｨｨ080609ｨ ｨｪｨｨ07. 00 ｨｪｨ 09ｨｰ0607ｨｴ0202 090802ｨｬ07 010507 07060106ｨ ｨｪ06ｨｦ 01ｨｨ0000020802ｨｪ02ｨｨｨ 02ｨｨｨｨ 0708020105ｨ 00ｨ 02ｨｰ0907 ｨｨ09070605ｨｹ040609ｨ ｨｰｨｹ 080701 ｨｰｨｨ07ｨｨ ﾂｨｪ0401 010507 07060109ｨｨ010609 08ｨｨ09ｨ 070605ｨｨｨｬ060800ｨｪ0401 SSR-ｨｬｨ 08ｨｺ02080609, 08ｨ 0907080201020502ｨｪｨｪ0401 0706 090902ｨｬｨｮ 0002ｨｪ06ｨｬｨｮ (ｨｰｨ ｨ 05. 4). 02ｨ 08ｨ 040204 0708ｨｨｨｪ07ｨｰ06 06ｨｰｨｪ0609ｨｨｨｰｨｹ ｨｺ ｨｰ06ｨｬｨｮ 07060109ｨｨ01ｨｮ, ｨ 0605ｨｹ030202 ﾂｨｨ090506 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ0401 ｨ 0505020502ｨｦ ｨｺ06ｨｰ0608060006 06ｨｪｨｨ ｨｪ0209ｨｮｨｰ (7). 020507 ｨｰ060006 ﾂｨｰ06ｨ 04 ｨｨ09ｨｺ0506 ﾂｨｨｨｰｨｹ 0603ｨｨｨ 06 ﾂｨｪ0602 06ｨｰｨｪ020902ｨｪｨｨ02 090608ｨｰ0609 ｨｺ 07060109ｨｨ01ｨｮ indica, ｨｬ04 ｨｰｨ ｨｺ0302 070806090205ｨｨ ｨｨ01 ｨｬ060502ｨｺｨｮ050708ｨｪ0602 ｨｬｨ 08ｨｺｨｨ080609ｨ ｨｪｨｨ02. 04ｨｨ09. 1. 030605ｨｨｨｬ060800ｨｨ04ｨｬ ｨｮ 32 090608ｨｰ0609 08ｨｨ09ｨ (Oryza sativa L.) 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 09040709050702ｨｬ04ｨｦ 09 ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ02ｨｬ SSR-ｨｬｨ 08ｨｺ0208ｨ RM 335: 4 ｪ 01ｨ 08ｨｬ, 5 ｪ 05ｨｪ0203ｨｨｨｪｨｺｨ, 9 ｪ 05020807ｨ ｨｪｨｰｨｨｨｪ, 17 ｪ 01ｨ 0802ｨｨ0909, 32 ｪ 0604ｨｮｨｬ08ｨｮ01 (090608ｨｰｨ 08060909ｨｨｨｦ09ｨｺ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 07060109ｨｨ01 indica); 0609ｨｰｨ 05ｨｹｨｪ0402 0106080603ｨｺｨｨ ｪ 090608ｨｰｨ 08060909ｨｨｨｦ09ｨｺ06ｨｦ 09020502ｨｺ02ｨｨｨｨ 07060109ｨｨ01ｨ japonica. 00ｨ 08ｨｺ0208 ｨｬ060502ｨｺｨｮ050708ｨｪ06ｨｦ ｨｬｨ 090904 GenPak DNA Markers M 50 (020202 0009ｨ ｨ 0608ｨ ｨｰ0608ｨｨ07 0604060002ｨｪ03, 04060909ｨｨ07) (06ｨｰｨｬ02 ﾂ02ｨｪ 09ｨｰ080205ｨｺ06ｨｦ). 0604ｨｮ ﾂ02ｨｪｨｨ02 0002ｨｪ06ｨｬｨ 09 ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ02ｨｬ 11 SSR-ｨｬｨ 08ｨｺ02080609 0706ｨｺｨ 04ｨ 0506, ﾂｨｰ06 08060909ｨｨｨｦ09ｨｺｨｨ02 090608ｨｰｨ, 06ｨｰｨｪ0609ｨｨｨｬ0402 ｨｺ 07060109ｨｨ01ｨｮ indica, 070605ｨｨｨｬ060800ｨｪ04 0706 ｨｪ02ｨｺ06ｨｰ060804ｨｬ ｨｨ04 05ｨｰｨｨ01 ｨｬｨ 08ｨｺ02080609, ｨｪｨ 0708ｨｨｨｬ0208 0706 RM 335 (08ｨｨ09. 1), RM 240, RM 247, RM 25, RM 259, RM 242. 0006ｨｪ06ｨｬ060800ｨｪ04ｨｬｨｨ ｨｮ 09090201 08060909ｨｨｨｦ09ｨｺｨｨ01 090608ｨｰ0609 06ｨｺｨ 04ｨ 05ｨｨ09ｨｹ ｨｬｨ 08ｨｺ020804 RM 255, RM 258, RM 4, RM 29, RM 19 (08ｨｨ09. 2, 08 ｨｨ 09). 0306 ｨｬ06ｨｪ06ｨｬ060800ｨｪ04ｨｬ ｨｬｨ 08ｨｺ0208ｨ ｨｬ 090902 ｨｨ04ｨｮ ﾂ02ｨｪｨｪ0402 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ0402 090608ｨｰｨ ｨｪ020905ｨｨ ｨ 05050205ｨｹ, 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ04ｨｦ 010507 07060109ｨｨ01ｨ japonica, 0706 070605ｨｨｨｬ060800ｨｪ04ｨｬ ｪ ｨｮ ｨ 0605ｨｹ0302ｨｦ ﾂｨ 09ｨｰｨｨ 05ｨｰｨｨ01 090608ｨｰ0609, 06ｨｰｨｪ0609ｨｨｨｬ0401 ｨｺ 07060109ｨｨ01ｨｮ indica, ｨｰｨ ｨｺ0302 ｨｨｨｬ0205ｨｨ09ｨｹ ｨ 05050205ｨｨ, ｨｰｨｨ07ｨｨ ﾂｨｪ0402 010507 07060109ｨｨ01ｨ japonica (ｨｰｨ ｨ 05. 5). 06ｨ ｨｺ, 09 11 ｨｨ04ｨｮ ﾂ02ｨｪｨｪ0401 0506ｨｺｨｮ09ｨ 01 ｨｮ 090608ｨｰ0609 09ｨ ｨｪｨｺｨ ｨｦ09ｨｺｨｨｨｦ, 0308ｨｨｨｬ060809ｨｺｨｨｨｦ, 0604ｨｮｨｬ08ｨｮ01 ｨ 0405ｨｨ 090407090502ｨｪ04 ｨｰ0605ｨｹｨｺ06 ｨｰ08ｨｨ ｨ 0505020507, 09070202ｨｨ00ｨｨ ﾂｨｪ0401 010507 07060109ｨｨ01ｨ indica, ｨｮ 090608ｨｰ0609 01ｨ 0802ｨｨ0909 ｨｨ 01ｨ 08ｨｬ ｪ ﾂ02ｨｰ040802, ｨｮ 090608ｨｰｨ 05ｨｪ0203ｨｨｨｪｨｺｨ ｪ 0601ｨｨｨｪ ｨ 05050205ｨｹ, ｨｰｨｨ07ｨｨ ﾂｨｪ04ｨｦ 010507 05ｨｰ060006 07060109ｨｨ01ｨ. 060605ｨｹｨｺ06 090608ｨｰ 05020807ｨ ｨｪｨｰｨｨｨｪ ｨｬ060302ｨｰ ｨ 04ｨｰｨｹ 06ｨｰｨｪ020902ｨｪ ｨｺ 07060109ｨｨ01ｨｮ indica ｨｪｨ 0609ｨｪ0609ｨ ｨｪｨｨｨｨ ｨ ｨｪｨ 05ｨｨ04ｨ 09 ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ02ｨｬ ｨｨ04ｨｮ ﾂ02ｨｪｨｪ0401 ｨｬｨ 08ｨｺ02080609, ｨｺ06ｨｰ06080402 090202070502ｨｪ04 09 0506ｨｺｨｮ09ｨ ｨｬｨｨ, 070605ｨｨｨｬ060800ｨｪ04ｨｬｨｨ 010507 06ｨ 08ｨ 04020609 07060109ｨｨ010609. 05 ｨｰ06 ﾂｨｺｨｨ 040802ｨｪｨｨ07 08ｨ 0401020502ｨｪｨｨ07 08060909ｨｨｨｦ09ｨｺｨｨ01 06ｨ 08ｨ 04020609 ｨｪｨ 07060109ｨｨ0104 ｨｪｨ ｨｨｨ 06050202 ｨｨｨｪ000608ｨｬｨ ｨｰｨｨ09ｨｪ04ｨｬｨｨ 06ｨｺｨ 04ｨ 05ｨｨ09ｨｹ ｨｬｨ 08ｨｺ020804 RM 335, RM 240, RM 247, RM 25, RM 259 ｨｨ RM 242. 65
1308 09 04ｨｨ09. 2. 030605ｨｨｨｬ060800ｨｨ04ｨｬ ｨｮ 090608ｨｰ0609 08ｨｨ09ｨ (Oryza sativa L.) 08ｨ 04ｨｪ060006 05ｨｺ0605060006-0002060008ｨ 00ｨｨ ﾂ0209ｨｺ060006 070806ｨｨ090106030102ｨｪｨｨ07, 09040709050702ｨｬ04ｨｦ 09 ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ02ｨｬ SSR-ｨｬｨ 08ｨｺ02080609 RM 29 (08) ｨｨ RM 19 (09): ｨｰ08ｨｨ 070609050201ｨｪｨｨ02 0106080603ｨｺｨｨ 090708ｨ 09ｨ ｪ ｨｰｨｨ07ｨｨ ﾂｨｪ0402 090608ｨｰｨ 07060109ｨｨ01ｨ indica 04ｨ 08ｨｮｨ 0203ｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 0609ｨｰｨ 05ｨｹｨｪ0402 0106080603ｨｺｨｨ ｪ 090608ｨｰｨ 08060909ｨｨｨｦ09ｨｺ06ｨｦ 09020502ｨｺ02ｨｨｨｨ. 00ｨ 08ｨｺ0208 ｨｬ060502ｨｺｨｮ050708ｨｪ06ｨｦ ｨｬｨ 090904 GenPak DNA Markers M 50 (020202 0009ｨ ｨ 0608ｨ ｨｰ0608ｨｨ07 0604060002ｨｪ03, 04060909ｨｨ07) (06ｨｰｨｬ02 ﾂ02ｨｪ 09ｨｰ080205ｨｺ06ｨｦ). 5. 030605ｨｨｨｬ060800ｨｨ04ｨｬ ｨｮ 090608ｨｰ0609 08ｨｨ09ｨ (Oryza sativa L.) 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ06ｨｦ 09020502ｨｺ02ｨｨｨｨ, 09040709050702ｨｬ04ｨｦ 09 ｨｨ09070605ｨｹ040609ｨ ｨｪｨｨ02ｨｬ SSR-ｨｬｨ 08ｨｺ02080609, ｨｰｨｨ07ｨｨ ﾂｨｪ0401 010507 07060109ｨｨ010609 indica ｨｨ japonica 02060507 ｨ 0505020502ｨｦ, C0608ｨｰ ｨｪ02 01ｨ 08ｨ ｨｺｨｰ0208ｨｪ0401 010507 07060109ｨｨ01ｨ japonica 09ｨ ｨｪｨｺｨ ｨｦ09ｨｺｨｨｨｦ 1 2 1 2 2 1 2 1 2 2 1 3/11 0308ｨｨｨｬ060809ｨｺｨｨｨｦ 1 1 1 1 1 1 2 1 2 1 1 3/11 05020807ｨ ｨｪｨｰｨｨｨｪ 1 1 1 2 2 1 2 1 1 1 1 6/11 01ｨ 0802ｨｨ0909 1 2 1 2 1 1 2 1 1 1 1 4/11 05ｨｪ0203ｨｨｨｪｨｺｨ 2 2 1 2 1 1 2 1 2 2 1 1/11 01ｨ 08ｨｬ 1 2 1 2 2 1 2 1 1 2 1 4/11 0604ｨｮｨｬ08ｨｮ01 1 2 1 2 1 1 2 1 2 1 1 3/11 japonica 2 2 1 1 1 1 2 1 2 2 1 0 p p m p p m m m 08 p m 03 08 ｨｨ ｨｬ 02 ﾂ ｨ ｨｪ ｨｨ 02. p ｨｨ m ｪ 090606ｨｰ0902ｨｰ09ｨｰ0902ｨｪｨｪ06 070605ｨｨｨｬ060800ｨｪ04ｨｦ ｨｨ ｨｬ06ｨｪ06ｨｬ060800ｨｪ04ｨｦ ｨｬｨ 08ｨｺ0208. RM 335 RM 240 RM 255 RM 247 RM 242 RM 4 06ｨ ｨｺｨｨｨｬ 06ｨ 08ｨ 0406ｨｬ, ｨｪｨ ｨ 050601ｨ 02ｨｬｨｮ06 ｨｮ ｨｬ020307060109ｨｨ0106090401 00ｨｨｨ 08ｨｨ010609 09040906ｨｺｨｮ06 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｹ 0708ｨｨ 09ｨｺ0802ｨｴｨｨ09ｨ ｨｪｨｨｨｨ 08060909ｨｨｨｦ09ｨｺｨｨ01 06ｨ 08ｨ 04020609 ｨｬ0603ｨｪ06 06ｨ ｨｲ0709ｨｪｨｨｨｰｨｹ ｨｨ01 0002ｨｪ02ｨｰｨｨ ﾂ0209ｨｺｨｨｨｬ 08060109ｨｰ0906ｨｬ ｨｨ 0902080607ｨｰｨｪ04ｨｬ ｨｪ020708ｨ 09ｨｨ05ｨｹｨｪ04ｨｬ 06ｨｰｨｪ020902ｨｪｨｨ02ｨｬ ｨｺ 07060109ｨｨ01ｨｮ indica 0706 ｨｨｨｪ0102ｨｺ09ｨｮ 040208ｨｪ0609ｨｺｨｨ. 0306 080204ｨｮ05ｨｹｨｰｨ ｨｰｨ ｨｬ ｨ ｨｪｨ 05ｨｨ04ｨ 06040208ｨｪ02ｨｪｨｪ0609ｨｰｨｨ 00ｨｨｨ 08ｨｨ010609 ｨｮ09ｨｰｨ ｨｪ06090502ｨｪ06, ﾂｨｰ06 0906090502 ﾂ02ｨｪｨｪ0402 09 09ｨｺ0802ｨｴｨｨ09ｨ ｨｪｨｨ07 090608ｨｰｨ 08060909ｨｨｨｦ09ｨｺ06ｨｦ 09020502ｨｺ02ｨｨｨｨ ｨｪ02 ｨｪ0209ｨｮｨｰ 0002ｨｪ04 03ｨｨ0806ｨｺ06ｨｦ 090609ｨｬ0209ｨｰｨｨｨｬ0609ｨｰｨｨ, 060708020102050706ｨｴｨｨ02 09040906ｨｺｨｨ02 04ｨｪｨ ﾂ02ｨｪｨｨ07 05ｨｰ060006 0706ｨｺｨ 04ｨ ｨｰ020507 ｨｮ 00ｨｨｨ 08ｨｨ010609. 0308060102ｨｬ06ｨｪ09ｨｰ08ｨｨ080609ｨ ｨｪｨ ｨｪ0206ｨ 010601ｨｨｨｬ0609ｨｰｨｹ 09ｨｺ0506 ﾂ02ｨｪｨｨ07 0106070605ｨｪｨｨｨｰ0205ｨｹｨｪ0401 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ09ｨｰｨｨｨｺ 09 07020802 ﾂ02ｨｪｨｹ 0708ｨｨ04ｨｪｨ ｨｺ0609, ｨｪｨ 0609ｨｪ0609ｨ ｨｪｨｨｨｨ ｨｺ06ｨｰ06080401 070806090601ｨｨｨｰ0907 08ｨ 0401020502ｨｪｨｨ02 06ｨｰ02 ﾂ0209ｨｰ0902ｨｪｨｪ0401 06ｨ 08ｨ 04020609 ｨｪｨ 07060109ｨｨ0104 indica ｨｨ japonica, 010507 0906ｨｺ08ｨ ｨｴ02ｨｪｨｨ07 ﾂｨｨ0905ｨ ｨｪ02ｨｺ06080802ｨｺｨｰｨｪ0401 ｨｨ0102ｨｪｨｰｨｨ00ｨｨｨｺｨ 02ｨｨｨｦ. RM 29 RM 19 RM 25 RM 259 RM 258 010107 00090208060909ｨｨｨｦ09ｨｺｨｨｨｦ 010606 08ｨｨ09ｨ 0408050901, 350921 04060909ｨｨ07, 00. 0808ｨ 09ｨｪ0601ｨ 08, 070609. 09020506040208ｨｪ04ｨｦ, e-mail: serggontchar@mail.ru 030609ｨｰｨｮ07ｨｨ05ｨ 09 080201ｨ ｨｺ02ｨｨ06 22 06ｨｺｨｰ07ｨ 0807 2012 000601ｨ Sel ｯskokhozyaistvennaya biologiya [Agricultural Biology], 2013, 01 5, pp. 61-68 STERILITY OF INTERSUBSPECIFIC HYBRIDS OF Oryza sativa L. IN CONNECTION WITH SEARCH OF GENES OF WIDE COMPATIBILITY AND CORRECT REFERENCE OF SAMPLES TO SUBSPECIES indica AND japonica 66
13E.M. Kharitonov, Yu.K. Goncharova All-Russian Research Institute of Rice, Russian Academy of Agricultural Sciences, pos. Belozerniy, Krasnodar, 350921 Russia, e-mail serggontchar@mail.ru Received October 22, 2012 doi: 10.15389/agrobiology.2013.5.61eng Acknowledgements: Supported by Russian Foundation for Basic Research. Abstract In rice (Oryza sativa L.) between indica and japonica subspecies a number of transitional forms both on phenotype and on physiological determinants present, which have the characteristic features in their development and adaptation to stress. The heterosis effect in intersubspecific hybrids (japonica/indica) can be of 100 % and more. The superhigh heterosis can be used during intersubspecific hybridization on conditions that the genes of wide compatibility (WC), which ensure a grain content more of 70 %, will be involve in breeding. The authors made the crossings between the samples of domestic and foreign breeding from japonica subspecies. The analysis of grain formation in the hybrid combinations did not allow to reveal WC genes in studied varieties of Russian selection defining high fertility of hybrids. The molecular scanning of hybrids and their parental forms was made with the use of 11 polymorphous SSR-markers, typical for subspecies. A possibility to obtain the fertile intersubspecific hybrids of rice and the list of trait used for differentiating samples to subspecies japonica and indica are discussed. The obtained results show that it is necessary to include additional characteristics into the list of trait for the taxonomic attributing samples to indica and japonica subspecies. indica. Keywords: rice, heterosis, hybrid, wide compatibility genes (WC), rice subspecies, japonica, REFERENCES 1. Ting M., Hai X., Quan X., Yan-hua G., Chunjie Z., Kai C., Jiayu W., Zhengjin X. Comparison of morphological and genetic differentiations in filial generation of cross between indica and japonica rice. Rice Sci., 2010, 17(1): 82-86. 2. Cheng K.S. Discrimination of indica-japonica subspecies in Asia 09ultivated rice. Kunming, Yunnan Science and Technology Press, 1993: 1-23. 3. Oka H.I. Origin of cultivated rice. Tokyo, Scientific Societies Press, 1988: 181-209. 4. Ting Y. The origin and evolution of cultivated rice in China. Acta Agron. Sin., 1957, 8: 243-260. 5. Chang T.T. The origin, evolution, cultivation, dissemination, and diversification of Asian and African rice. Euphytica, 1976, 25: 435-441. 6. Wang C.L., Zhang Z.L., Tang S.Z., Shi J.D. Exploitation of heterosis between indica and japonica by three-line method. I. Differentiation between indica-japonica sterility and cytoplasmic male sterility. Jiangsu J. Agr. Sci., 1992, 8: 1-7. 7. Zhang Q., Liu K.D., Yang G.P., Saghai M.A., Maroof S.A., Xu G., Zhou Z.Q. Molecular marker diversity and hybrid sterility in indica-japonica rice crosses. Theor. Appl. Genet., 1997, 95: 112-118. 8. Yang G.P., Maroof S.A., Xu C.G., Zhang Q., Biyashev R.M. 05omparative analysis of microsatellite DNA polymorphism in landraces and cultivars of rice. Mol. Gen. Genet., 1994, 245: 187-194. 9. Garris A.J., Tai T.H., Coburn J. et al. Genetic structure and diversity in Oryza sativa L. Genetics, 2005, 169: 1631-1638. 10. Mackill D.J. Classifying japonica cultivars with RAPD markers. Crop. Sci., 1995, 35: 889-894. 11. Gontcharova I.K., Gontcharov S.V. Hybrid rice breeding in Russia. Proc. 4th Intern. Symp. on hybrid rice 00Hybrid rice for food security, poverty alleviation, and environmental protection03 S.S. Virmani, S.X. Mao, B. Hardy (eds.). Los Banos (Philippines), Int. Rice Res. Inst., 2003: 321-328. 12. Goncharova J.K. Fixing heterosis effect in rice hybrids by Strunnikov ｯs method. Proc. Int. Conf. on Integration of Science and Technology for Sustainable Development. Bangkok, Thailand, 2007: 393-396. 13. Zhang G.Q., Lu Y.G. Genetics of F1 pollen sterility in Oryza sativa. In: Rice genetics III. Manila, IRRI, 1996: 418-422. 14. Yuan L.P. Strategic hypotheses of breeding for hybrid rice (in Chinese). Hybrid Rice, 1987, 1: 1-3. 15. Ikehashi H., Araki H. Variety screening of compatibility types revealed in F 1 fertility of distant cross in rice. Jpn. J. Breed., 1984, 34: 304-313. 16. Ikehashi H., Araki H. Genetics of F1 sterility in remote crosses of rice. In: Rice genetics: proceedings of the international rice genetics symposium, 27-31 May 1985. Manila, Philippines, IRRI, 1986: 119-130. 67
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