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Limited Plasticity of Rabbit ｯs Visual Cortex and Hippocampal Neurons of Differences V. B. Polyansky, D. V. Evtikhin, E. N. Sokolov, A. V. Kryuchkova Departments of Higher Nervous Activity and Psychophysiology, M.V. Lomonosov State University, Moscow, e-mail: pol@protein.bio.msu.ru The activity of 41 visual cortex and 20 hippocampal neurons from field CA1 was registered in experiments using oddball-stimulation with different color stimuli varied in intensity. 34% cortical and 37% hippocampal neurons demonstrated plasticity reactions. The significant increase of latest phases of neuronal activity (200ｨC500 and 200ｨC1000 ms after stimulation for cortical neurons and 300ｨC550 ms for hippocampal neurons) was shown in responses to rare deviant stimuli, which had a less intensity than frequently standards. The quantity of the earliest neuronal phase of activity (40ｨC120 ms after stimulation) was stabilized in responses to deviants and standards during the experiment. We propose that such increase of the latest phases of neuronal activity (the limited plasticity) may reflect the mechanisms of orienting reaction. Key words: oddball, rabbit, orienting reflex, deviants and standards, visual cortex and hippocampal neurons of differences. ｨｨ07060200 076 56ｦ003000200006ｦ0fl 096ｦ00300006ｦ0 07 00076 707 0ｷ36 50003076 5076 707 0603090304026ｦ0 apple046 70903 0807096 5070209026ｦ0 07 006ｦ0apple0408090300 6 5 09apple0903000200-000706 0107apple00 01apple070209016ｦ0 000006apple07000ｸ8, 006ｦ00ｹ06ｦ00200000ｸ806 06086ｦ00601076 50ｸ806 apple6ｦ009applefl 01070307apple0ｸ803 0307080003 06020408090300 0300apple0706 0ｷ36 50003076 5076 707 apple6ｦ00902090ｹ009fl 06030000fl00030ｸ803 060309030402076 5 [5]. ｨｮ09060207 06086ｦ00601076 5, 060706036ｦ06 502fl0ｸ90ｸ70903 apple6ｦ009applefl apple6ｦ00902090ｹ009fl, 02090000060007 066 5fl096ｦ00007 06 6ｦ0 0607020ｸ903000706 6 50002090ｹ009000706 030003 6 500010307apple076 5 6 50709 0408 000009fl, 01070307apple0ｸ800 0807 6ｦ000000ｸ803 08076 500 00000ｹ00006010903 0ｹ101060800apple0903000003076 5 09 6ｦ0006ｦ00209096ｦ0 026ｦ009 09apple090300020000076 707 6 50ｸ8096 56ｦ00000076 707 08070300000ｷ3096ｦ0-026ｦ0 (ｨ 05ｨｨ) 0107 09apple076 56ｦ00209 0ｹ10309 0ｷ36 50003076 50ｸ800 0603090304020ｸ8 [3, 6]. ｨｨ07 036 500apple08 0000090003 03076 707, 0ｹ00307 apple6ｦ000000900 apple6ｦ009applefl 0ｸ8 (50ｨC90 0306 0703 0307030000036ｦ0 060300000ｸ8 060309030402076 5) 000006apple0700076 5 apple6ｦ00902090ｹ009fl 000606036 509030002000007 0708apple00 0002fl0ｸ90306fl apple6ｦ00900070603000ｸ9 6 500010307apple076 5 6 50709 0408 000009fl 06030000fl00030ｸ803 060309030402076 5, 06020408090207 08070603apple0700000900 0ｷ36 50003076 5076 707 08apple070603apple6ｦ00006036 56ｦ0 0807 036ｦ003apple090ｷ300 06086ｦ006-01076 50ｸ803 apple6ｦ009applefl 076 5 036ｦ001076 707 000006apple07006ｦ0 apple6ｦ00902090ｹ009fl 006ｦ0 apple6ｦ009-000ｸ800 0ｷ36 500036ｦ0. ｨｴ0307 6 500010307apple000700 0ｷ36 50003076 50700 08apple070603apple6ｦ00006036 507, 07 apple6ｦ009040ｸ90ｸ70000 6 7090800apple060200apple04 6 5 0ｹ000030ｸ8apple00030300apple000703 006 5010209-076 50703 08apple070603apple6ｦ00006036 500, 06076 5086ｦ00207 06 0ｷ36 50003076 50ｸ803 08apple070603apple6ｦ00006036 50703, 080702040ｹ00000000703 006ｦ0 070600076 500 0ｷ36 50003076 50ｸ803 040602076 5-0007apple00020200010307apple000ｸ803 09020200apple00000ｷ309apple076 50701 [5]. ｨｺ6ｦ000000900 apple6ｦ009applefl 0ｸ8 000006apple0700076 5 apple6ｦ00902090ｹ009fl 036ｦ0apple6ｦ0010300apple09-09076 56ｦ002090600 6 50ｸ80607010706 06036ｦ0 09020000070603000ｸ9. 0307020000 080709 000900 apple6ｦ009applefl 0ｸ8 036ｦ0010706 06036ｦ0 09020000070603000ｸ9 0000 070302090ｹ06ｦ002090600. ｨｨapple09 360
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13362 ｨｨ0702fl0006010906 09 apple. 02 05006 5096ｦ000030ｸ8 05036ｦ000 6ｦ0apple030ｸ8 N = 40 046ｦ00ｹ06ｦ00200000ｸ800 N = 40 03 18.9 12.6 19.2 14.1 05 N = 24 Oddball N = 216 07 33.9 16.7 21.0 15.9 05 N = 40 010700000ｹ0000ｸ800 N = 40 00 25.9 14.2 20.6 14.0 ｨｺ0906. 1. ｨｦ036 500030ｸ8 000006apple07006ｦ0 6ｧ8 2 09apple0903000200000706 0107apple0ｸ8 01apple070209016ｦ0 006ｦ0 0603090304020ｸ8 060900006 707 0ｷ36 500036ｦ0. ｨ 0003000006096 50007060300 006 5096ｦ00003076 5 ｨC 5 01 /03 2, 06036ｦ000 6ｦ0apple03076 5 ｨC 8 01 /03 2. 046ｦ0 016ｦ008 0703 016ｦ0 apple00 apple09060400016ｦ0 066 500apple0304 04016ｦ0096ｦ00007: N ｨC 0ｹ009060207 060309030402076 5, 6ｦ0020000 08apple096 500 00000ｸ8 apple6ｦ00603apple0ｸ8 09 08070603060309-03040200000ｸ800 6 7090603076 7apple6ｦ003030ｸ8 07036 50003076 5. ｨｮ0906026ｦ0 6 5 0600apple00 090000 016ｦ008 076 707 016ｦ0 apple6ｦ0 ｨC 06apple00 000000 0ｹ009060207 06086ｦ00601076 5 6 5 1 06 6 5 apple6ｦ009000ｸ800 09000300apple6 56ｦ0020ｸ8 6ｦ0006ｦ0-0209096ｦ0: 250ｨC550 0306, 250ｨC1000 0306. 0503apple0002010706 6 56 500apple03 04016ｦ0096ｦ00007 006ｦ00ｹ06ｦ00207 060309030402fl0ｷ30909, 0603apple0002010706 6 5000909 ｨC 010700000ｷ3 000606036 509fl 0603090304026ｦ0. ｨｨ07 070609 07apple 09006ｦ003 070302070800006ｦ0 06apple00 00flfl 0ｹ06ｦ0060307036ｦ0 06086ｦ00601076 5 6 5 1 06, 0807 070609 6ｦ0 060ｷ3090606 ｨC 6 5apple0003fl apple006ｦ0010ｷ30909, 0306. 0502006 56ｦ0 0703 0307030000036ｦ0 0807 6ｦ00ｹ009 0603090304026ｦ0 ｨC 020700076 56ｦ0fl 6ｦ00103096 50007060300. 0509 0007, 0ｹ00307 006ｦ009 070200046ｦ0fl 0ｹ06ｦ0060307036ｦ0 apple6ｦ009applefl 076 5 6 5 09000300apple6 56ｦ0026ｦ003 6ｦ0006ｦ00209096ｦ0 200ｨC500 09 200ｨC1000 0306 006ｦ0 020ｸ9 6ｦ0000306fl 6 5 07036 500036ｦ003 006ｦ0 oddball- 006 5096ｦ000030ｸ8 (0807 06apple6ｦ06 5000000090ｸ9 06 07036 500036ｦ00309 006ｦ0 0ｹ06ｦ006030ｸ800 06036ｦ000 6ｦ0apple030ｸ8 09 07 0900070ｹ0000ｸ800 006 5096ｦ000030ｸ8). 02 ｨC 07 0900070ｹ0000ｸ800 006ｦ00ｹ06ｦ00200000ｸ800 006 5096ｦ000030ｸ8, 03 ｨC 006ｦ00ｹ06ｦ00200000ｸ800 07 0900070ｹ0000ｸ800 06036ｦ000 6ｦ0apple030ｸ8, 05 ｨC 006 5096ｦ000030ｸ8 6 5 08apple076 7apple6ｦ0030300 oddball, 07 ｨC 06036ｦ000 6ｦ0apple030ｸ8 6 5 08apple076 7apple6ｦ0030300 oddball, 05 ｨC 010700000ｹ0000ｸ800 07 0900070ｹ0000ｸ800 006 5096ｦ000030ｸ8, 00 ｨC 010700000ｹ0000ｸ800 07 0900070ｹ0000ｸ800 06036ｦ000 6ｦ0apple030ｸ8. 046ｦ0 apple0906. 1 0807016ｦ0096ｦ0006ｦ0 09006ｦ00309016ｦ0 07036 50003076 5 000006apple07006ｦ0 6ｧ8 2 6 5 08apple070ｷ300 04apple00 oddball. ｨ 000600 6 5 016ｦ00ｹ00006036 500 06036ｦ000 6ｦ0apple- 03076 5 09060807020009076 56ｦ002090600 0609000900 0603090304020ｸ8 090003000006096 5000706-03000ｸ9 8 01 /03 2, 6ｦ0 6 5 016ｦ00ｹ00006036 500 006 5096ｦ00003076 5 ｨC 0300 0800 0609000900 0603090304020ｸ8, 0007 06 03000000040006 090003000006096 500070603000ｸ9 ｨC 5 01 /03 2. ｨｨapple0008 00 6 506006 707 060200 040003 06016ｦ0096ｦ00300, 0ｹ00307 0000 07 006ｦ0apple0408096 56ｦ0-000306fl 08apple0909006ｦ001076 5 08apple096 50ｸ8016ｦ00009fl 07036 500036ｦ0 (0603. apple6ｦ00603apple0ｸ8) 08apple09 020903000200000703 08apple0903000000000909 060309030402076 5: 0009 02fl 00-6 5096ｦ00003076 5, 0009 02fl 06036ｦ000 6ｦ0apple03076 5; 036ｦ0010800 0009 6 5 07 000706 026ｦ00900 07036 50003076 5 0000 6 50ｸ8fl6 502000007 016ｦ001076 707-0209 07 06040ｸ70006036 5000000076 707
13ｨｦ07ｨｺ0204ｨ ｨｮ00040402ｨｹ ｨｨ000205ｨｪｨ ｨｮ04ｨｦ05ｨｪｨｲ 040009ｨｺｨｦ04ｨｦ05 363 ｨｪ6ｦ0 02090ｷ36ｦ0 1. 05apple00 000900 6 50002090ｹ009000ｸ8 0ｹ06ｦ0060307030ｸ8 06086ｦ00601076 5 6 5 1 06 6 5 07036 500036ｦ003 000006apple0700076 5 09apple0903000200000706 0107apple0ｸ8 01apple070209016ｦ0 006ｦ0 006 5096ｦ000030ｸ8 09 06036ｦ000 6ｦ0apple030ｸ8 6 5 07080ｸ8036ｦ003 0607 0603090304026ｦ00309 07 00076 707 0ｷ36 500036ｦ0, 0007 apple6ｦ009000706 090003000006096 50007060309 ｨｴ08070309 6ｦ0006ｦ00209096ｦ0, 0306 0503090304020ｸ8 40ｨC120 200ｨC500 200ｨC1000 0ｨC1000 05006 5096ｦ000030ｸ8 07 0900070ｹ0000ｸ800 6 5 006ｦ00ｹ06ｦ00200 07080ｸ8036ｦ0 78.2 ﾀ 46.8 15.5 ﾀ 9.0** 11.6 ﾀ 6.3** 13.2 ﾀ 5.6* 05006 5096ｦ000030ｸ8 oddball 78.1 ﾀ 46.8 17.6 ﾀ 9.5 12.7 ﾀ 6.0 14.2 ﾀ 6.0 05006 5096ｦ000030ｸ8 07 0900070ｹ0000ｸ800 6 5 0107000ｷ300 07080ｸ8036ｦ0 81.5 ﾀ 44.2 15.7 ﾀ 8.0** 11.9 ﾀ 5.5 13.8 ﾀ 5.8 05036ｦ000 6ｦ0apple030ｸ8 07 0900070ｹ0000ｸ800 6 5 006ｦ00ｹ06ｦ00200 07080ｸ8036ｦ0 75.1 ﾀ 43.9 16.4 ﾀ 10.4* 11.8 ﾀ 6.3* 13.3 ﾀ 6.1* 05036ｦ000 6ｦ0apple030ｸ8 oddball 78.7 ﾀ 46.2 16.3 ﾀ 8.4* 12.1 ﾀ 5.4* 13.9 ﾀ 5.8 05036ｦ000 6ｦ0apple030ｸ8 07 0900070ｹ0000ｸ800 6 5 0107000ｷ300 07080ｸ8036ｦ0 75.8 ﾀ 41.0 17.1 ﾀ 9.5 12.1 ﾀ 5.9 13.8 ﾀ 6.1 ｨｨapple0903000ｹ06ｦ0000900. ｨｮ09060207 000006apple0700076 5 apple6ｦ06 50007 14. ｨｮ09060207 0600apple0906 ｨC 71. ｨ 6 50009 070ｹ0016ｦ00309 070303000ｹ0000007 070603076 500apple000700 070302090ｹ00900 apple006ｦ0010ｷ30909 0703 07036 50003076 5 006ｦ0 oddball- 006 5096ｦ000030ｸ8: * ｨC apple < 0.05, ** ｨC p < 0.01. 0909030000000009fl apple6ｦ009applefl 000706 6ｦ00103096 50007060309 08apple09 0807060200 076 56ｦ0-03000200000703 08apple00 ｨBfl6 50200000909 apple6ｦ009 apple6ｦ008090300020006. ｨｨapple09 6ｦ0006ｦ00209-0900 apple09060400016ｦ0 0307080007 096ｦ0030003090300, 0ｹ00307, 030703fl 006 5096ｦ000030ｸ8 0807 020909090ｹ00006010903 036ｦ0apple6ｦ0010300apple09060309016ｦ003 0603090304020ｸ8 07020000 06026ｦ0-0ｸ800, 0ｹ00003 06036ｦ000 6ｦ0apple030ｸ8, 6ｦ00103096 50007060300 000006apple07006ｦ0 6 5 000001070307apple0ｸ800 0800apple0907 0ｸ8 07036 500036ｦ0 006ｦ0 000903 0807 06apple00 00000304 0ｹ009060204 06086ｦ00601076 5 6 50ｸ80400, 0ｹ00003 6 5 07036 500036ｦ003 006ｦ0 06036ｦ000 6ｦ0apple030ｸ8. ｨ 000600, 6 509 090307, 06016ｦ0090ｸ86 56ｦ0000306fl 000606036 50900 0000070809 6ｦ0000007060309 0807fl6 502000009fl 006 5096ｦ00003076 5, apple046 7090309 0602076 56ｦ00309, 000606036 50900 07apple0900000309apple076 5070ｹ000076 707 apple0002020001066ｦ0. ｨｨapple0008 00 6 506006 707 apple6ｦ006-06030703apple0903, 016ｦ001 0909030000fl000306fl 0800apple6 5090ｹ0000ｸ806 apple6ｦ009applefl, 090209, 016ｦ001 030ｸ8 006 707 006ｦ0096 56ｦ00209 apple6ｦ000000400 [5], ｰapple6ｦ009applefl apple6ｦ00902090ｹ009fl ｱ. 05 07036 50003 006ｦ0 apple00 010900 006 5096ｦ000030ｸ8 6 5 066ｦ0030706 08apple070ｷ300 04apple00 oddball (apple0906. 1, 05) 06apple00 00flfl 0ｹ06ｦ0060307036ｦ0 06086ｦ00601076 5 6 5 000003 ｨC 139.1 0608/06. ｨｴ0307 030000000400, 0ｹ00003 6 5 07036 500036ｦ003 006ｦ0 006ｦ00ｹ06ｦ00200-000ｸ800 07 0900070ｹ0000ｸ800 006 5096ｦ000030ｸ8 (02) 09 06036ｦ000 6ｦ0apple030ｸ8 (03), 0007 0702000400, 0ｹ00003 006ｦ0 06036ｦ000 6ｦ0apple030ｸ8 6 5 08apple070ｷ300 04apple00 oddball (07) 09 6 5 07036 500036ｦ003 006ｦ0 010700000ｹ0000ｸ800 07 0900070ｹ0000ｸ800 006 5096ｦ000030ｸ8 (05) 09 06036ｦ000 6ｦ0apple030ｸ8 (00). ｨｦ 006ｦ00107 036ｦ0010700 0607070300070400000900 036ｦ0apple6ｦ001-0300apple0007 030702000107 02fl 6ｦ00000076 707 000006apple07006ｦ0. 016ｦ001 0307080007 046 509 000300 6ｦ0020000 (036ｦ0 02. 1), 000003 06036ｦ003090603090ｹ000060109 09006ｦ00ｹ009-030706 apple6ｦ00900090ｷ30ｸ8 02fl 0ｹ103076 707 apple6ｦ009applefl 6ｦ0 6 5 07036 500036ｦ003 006ｦ0 oddball- 006 5096ｦ000030ｸ8 09 006ｦ0 6 50600 apple046 70900 6 509 0ｸ8 060309030402076 5. 076ｦ0apple6ｦ0010300apple000ｸ803 066 5070606036 50703 6ｦ00000076 707 000006apple07006ｦ0 fl6 502fl- 000306fl 0307, 0ｹ00307 080709 000900 026ｦ0090ｸ8 07036 500036ｦ0 006ｦ0 apple00 010900 oddball- 006 5096ｦ000030ｸ8 (200ｨC600, 200ｨC1000 0306) 0807016ｦ0090ｸ86 56ｦ00ｸ903 fl6 50007 046 50002090ｹ0000000040ｸ9 06apple00 000ｸ90ｸ9 0ｹ06ｦ00603070304 apple6ｦ009applefl 6ｦ0, 06070703-6 5000306036 500000007 33.9 0608/06 09 16.7 0608/06 0807 06apple6ｦ06 5000000090ｸ9 06 0607-07036 5000306036 5040ｸ90ｸ7090309 026ｦ0096ｦ00309 07036 50003076 5 006ｦ0 6 50600 apple046 70900 6 509-0ｸ8 060309030402076 5. 0407 036ｦ0010900 apple006ｦ0010ｷ30909, 016ｦ001 04 000006apple07006ｦ0 6ｧ8 2, 07 006ｦ0apple04-0800000ｸ8 6ｦ002000107 0000 04 6 5060003 096ｦ0apple006 7090603apple09apple076 56ｦ000000ｸ803 000006apple07-00076 5 0ｹ1030706 0600apple0909. ｨｮ0307 0ｸ8 07 07 0ｸ7090300 apple0009040200036ｦ0030ｸ8, 0ｸ802 08apple076 500 0000 6ｦ0006ｦ0020909 apple006ｦ0010ｷ30906 6 5060003 0906060200 076 56ｦ000000ｸ803 026ｦ009-000ｸ803 000006apple0700076 5 09apple0903000200000706 0107apple0ｸ8. 0506006 707 096ｦ00809066ｦ0000ｸ8 09 08apple076ｦ0006ｦ002090909apple076 56ｦ0000ｸ8 07036 500030ｸ8 6 5 239 0600apple09fl03 07080ｸ803076 5 06 apple6ｦ009000ｸ80309 090003000006096 500070603fl0309 04 41 000006apple07006ｦ0. 02006ｦ0020909 0807016ｦ0096ｦ002, 0ｹ00307 6 5 09apple0903000200000706 0107apple00 060403036ｦ0apple0007 02fl 6 5060003 0600apple0906 07080ｸ803076 5 0000 07 006ｦ0apple0408000007 06040ｸ70006036 50000000706 apple6ｦ0090009-0ｷ30ｸ8 030008 04 07036 500036ｦ00309 006ｦ0 oddball- 006 5096ｦ000030ｸ8 09 07036 500036ｦ0-0309 006ｦ0 6 50600 apple046 70900 apple6ｦ009 apple6ｦ0080903000209 (6 5 030703 0ｹ009060200 006ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8) 0009 02fl 07 00076 707 09000300apple6 56ｦ0026ｦ0 6ｦ0006ｦ00209096ｦ0. ｨｨapple076 500 0000000ｸ806 06036ｦ003090603090ｹ00006010906 6ｦ0006ｦ0020909 0807 086ｦ0apple00070304 01apple090300apple090ｸ9 0509020107010607006ｦ0 0807 036 500apple 0902 0ｹ1030703 6 50ｸ86 507. ｨｦ 006ｦ00107 06apple00 09 07 0ｸ7006 707 036ｦ00606096 56ｦ0 0906060200 076 56ｦ000000ｸ803 000006apple0700076 5 030ｸ8 6 50ｸ8 0002090209 010200030109, 0ｹ00009 apple006ｦ0010ｷ30909 006ｦ0 oddball- 006 5096ｦ000030ｸ8 6 5 07080ｸ80300 030703fl 0ｸ8 6 5 07 000703 09000300apple- 6 56ｦ00200 6ｦ0006ｦ00209096ｦ0 07016ｦ0096ｦ002090600 ｨｮ020004090309 0807 06apple6ｦ06 5000000090ｸ9 06 07036 500036ｦ00309 006ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8. ｨｪ6ｦ0010903 010200030701 07016ｦ0-096ｦ002070600 14, 090209 34% 0703 6 5060003 0906060200 076 56ｦ000000ｸ803 000006apple07-00076 5 (71 0600apple09fl 07080ｸ803076 5). ｨｺ0009040200036ｦ0030ｸ8 6ｦ0006ｦ00209096ｦ0 apple006ｦ001-0ｷ30906 0ｹ1030903 000006apple0700076 5 08apple00 06036ｦ06 50200000ｸ8 6 5 036ｦ0 02. 1. ｨｺ6ｦ006060307-03apple00000900 036ｦ0 02. 1 0807016ｦ0090ｸ86 56ｦ00003, 0ｹ00307 02fl 09000300apple6 56ｦ002076 5 200ｨC500 09 200ｨC1000 0306 apple006ｦ0010ｷ30909 006ｦ0 oddball- 006 5096ｦ000-030ｸ8 0702000400 apple006ｦ0010ｷ30906 0000 030702000107 006ｦ0 oddball-06036ｦ000 6ｦ0apple- 030ｸ8, 0007 09 006ｦ0 07 0900070ｹ0000ｸ800 006ｦ00ｹ06ｦ00200000ｸ800 09 010700000ｹ0000ｸ800 00-6 5096ｦ000030ｸ8 09 006ｦ00ｹ06ｦ00200000ｸ800 06036ｦ000 6ｦ0apple030ｸ8 (09000300apple6 56ｦ002 6ｦ0006ｦ00209096ｦ0 200ｨC500 0306) 09 006ｦ0 006ｦ00ｹ06ｦ00200000ｸ800 07 0900070ｹ0000ｸ800 06036ｦ000 6ｦ0apple030ｸ8 09 006 5096ｦ000030ｸ8 (09000300apple6 56ｦ002 6ｦ0006ｦ00209096ｦ0 200ｨC1000 0306). 050600 0ｹ10307 0807 036 500apple08 6ｦ0000306fl 6ｦ000000ｸ80309 06036ｦ003090603090ｹ0000601076 707 6ｦ0006ｦ00209096ｦ0 apple0009040200036ｦ003076 5 0807 086ｦ0apple00070304 01apple090300apple090ｸ9 050902-0107010607006ｦ0 (06 04apple076 5000003 09006ｦ00ｹ0090307060309 0000 07020000 0.05). 050200 040003 0703030003090300, 0ｹ00307 09 02fl 0ｹ1030903 08026ｦ00603090ｹ0000ｸ803 000006apple0700076 5 006ｦ00ｹ06ｦ00200000ｸ806 apple6ｦ009applefl 07036 50003076 5 006ｦ0 oddball- 006 509-6ｦ000030ｸ8 0706036ｦ0000306fl 0706036ｦ003070ｹ00007 06036ｦ0 090200000ｸ803 09 0000 070303000ｹ06ｦ0000306fl 09006ｦ00ｹ009030ｸ803 070302090ｹ00906 0703 07036 50003076 5 006ｦ0 6 50600 apple046 70900 6 509 0ｸ8 060309030402076 5, 6 5 030703 0ｹ009060200 006ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8. 0502fl 0ｹ009060307030ｸ8 0ｹ101060800apple09030000036ｦ0 0ｸ80209 08apple076 500 00000ｸ8 01070003apple070200000ｸ800 07080ｸ8030ｸ8 006ｦ0 15 000006apple07006ｦ003, 6 5 01070307apple0ｸ803 00-6 5096ｦ000030ｸ8 09 06036ｦ000 6ｦ0apple030ｸ8 0ｸ80209 07 09006ｦ001076 50ｸ8 (016ｦ001 0807 0ｷ36 500-0304, 036ｦ001 09 0807 090003000006096 50007060309). ｨｴ0309 07080ｸ8030ｸ8 0000 6 50ｸ8fl6 509-0209 016ｦ0010903-0209 07 apple6ｦ00902090ｹ00906 6 5 apple006ｦ0010ｷ309fl03 000006apple0700076 5 006ｦ0 apple00 010900 09 0ｹ06ｦ006030ｸ800 0603090304020ｸ8. 0709080807016ｦ00308 0506006 707 0ｸ80207 0906060200 076 56ｦ00007 20 000006apple0700076 5 080702fl CA1. 02006ｦ0020909 07036 50003076 5 000006apple0700076 5 6 709080807016ｦ003086ｦ0 0807016ｦ0096ｦ002, 0ｹ00307 070009 036ｦ0apple6ｦ0010300apple0909040ｸ90306fl 026ｦ009000ｸ803 apple6ｦ009applefl 0703 06 006ｦ0-0ｹ06ｦ00200000ｸ803 6 50ｸ8060701070ｹ06ｦ006030703000ｸ803 0809010703, 0307apple030709000706 086ｦ0-04090706 09 080709 000903 apple6ｦ009applefl 0703. ｨｪ6ｦ0010903 07 apple6ｦ0090703, 0903 07036 500-030ｸ8 0807 0603apple04010304apple00 06076 5086ｦ0 6ｦ00ｸ903 06 apple006ｦ0010ｷ309fl0309 000006apple0700076 5
13364 ｨｨ0702fl0006010906 09 apple. 02 05006 5096ｦ000030ｸ8 05036ｦ000 6ｦ0apple030ｸ8 N = 40 046ｦ00ｹ06ｦ00200000ｸ800 N = 40 03 39.4 25.5 44.9 26.8 05 N = 24 Oddball N = 216 07 49.3 29.5 39.8 27.1 05 N = 40 010700000ｹ0000ｸ800 N = 40 00 41.5 27.7 34.8 22.3 ｨｺ0906. 2. ｨｦ036 500030ｸ8 000006apple07006ｦ0 6ｧ8 14 6 709080807016ｦ003086ｦ0 01apple070209016ｦ0 006ｦ0 0603090304020ｸ8 060900006 707 0ｷ36 500036ｦ0. ｨ 0003000006096 50007060300 006 5096ｦ00003076 5 ｨC 7 01 /03 2, 06036ｦ000 6ｦ0apple- 03076 5 ｨC 8 01 /03 2. 0509 0007, 0ｹ00307 06apple00 00flfl 0ｹ06ｦ0060307036ｦ0 06086ｦ00601076 5 6 5 09000300apple6 56ｦ0026ｦ003 6ｦ0006ｦ00209096ｦ0 250ｨC500 09 250ｨC1000 0306 6 5 07036 500036ｦ003 006ｦ0 oddball- 006 509-6ｦ000030ｸ8 0702000400, 0ｹ00003 006ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8 09 07 0900070ｹ0000ｸ800 006 5096ｦ000030ｸ8. ｨｬ 6ｦ00000076 707 000006apple07006ｦ0 07036 500030ｸ8 006ｦ0 apple00 010900 006 5096ｦ000030ｸ8 6 5 09000300apple- 6 56ｦ00200 6ｦ0006ｦ00209096ｦ0 40ｨC120 0306 0702000400 07036 50003076 5 006ｦ0 0ｹ06ｦ006030ｸ800 06036ｦ000 6ｦ0apple030ｸ8 09 07 0900070ｹ0000ｸ800 006 5096ｦ000030ｸ8, 0007 0ｹ10307 fl6 502fl000306fl 060107apple0000 090601020ｸ90ｹ000-00090003, 0ｹ00003 08apple6ｦ06 509020703. ｨｦ 0709006ｦ00ｹ0000009fl 016ｦ001 006ｦ0 apple0906. 1. 080702fl CA1 6 709080807016ｦ003086ｦ0, 070809066ｦ000000ｸ80309 apple6ｦ0000000 ｨｦ.05. 0509-00076 7apple6ｦ0 076 50706 [23]. 006ｦ003000003000ｸ800 0800apple0907 0ｸ8 0903 apple6ｦ009applefl 076 5 0807 06apple6ｦ06 5000000090ｸ9 06 0107apple01076 50ｸ80309 000006apple07006ｦ00309 0603000ｸ700000ｸ8 6 5 060307apple070004 ｨｮ0200040903 09006ｦ00ｹ000000906 (apple0906. 2). 050600 096ｦ0apple006 7090603apple09apple076 56ｦ000000ｸ800 010200030109 0ｸ80209 070300000600-000ｸ8 01 000006apple07006ｦ003 apple6ｦ00902090ｹ009fl, 07 026ｦ0 6ｦ00ｸ90ｸ70903 08apple090300apple0007 030706 0800 0603apple04010304apple0706 06086ｦ00601076 5076 707 apple6ｦ009applefl 6ｦ0, 0ｹ00307 09 04 000006apple0700076 5 apple6ｦ00902090ｹ009fl 09apple0903000200000706 0107apple0ｸ8. 040006apple07000ｸ8 apple6ｦ00902090ｹ009fl 6 709080807016ｦ003086ｦ0 0000 07 006ｦ0apple0408096 56ｦ00209 0603070601076 707 08apple096 50ｸ8016ｦ00009fl. ｨｪ6ｦ001, 0909070209apple076 56ｦ000000700 08apple0903000000000900 40 06036ｦ000 6ｦ0apple03000ｸ803 09 40 006 5096ｦ00003000ｸ803 060309030402076 5 04 000006apple07-006ｦ0 6ｧ8 14 6 709080807016ｦ003086ｦ0 (apple0906. 2) 0000 08apple096 5000207 01 apple6ｦ0096 50903090ｸ9 08apple096 50ｸ8016ｦ00009fl, 6ｦ0 apple006ｦ0010ｷ30909 006ｦ0 006ｦ00ｹ06ｦ00200000ｸ800 0909070209apple076 56ｦ000-000ｸ800 06036ｦ000 6ｦ0apple03000ｸ806 09 006 5096ｦ00003000ｸ806 0603090304020ｸ8 0000 07030209-
13ｨｦ07ｨｺ0204ｨ ｨｮ00040402ｨｹ ｨｨ000205ｨｪｨ ｨｮ04ｨｦ05ｨｪｨｲ 040009ｨｺｨｦ04ｨｦ05 365 ｨｪ6ｦ0 02090ｷ36ｦ0 2. 05apple00 000900 6 50002090ｹ009000ｸ8 0ｹ06ｦ0060307030ｸ8 06086ｦ00601076 5 6 5 1 c 6 5 07036 500036ｦ003 000006apple0700076 5 080702fl 05A1 6 709080807016ｦ003086ｦ0 01apple070209016ｦ0 006ｦ0 006 509-6ｦ000030ｸ8 09 06036ｦ000 6ｦ0apple030ｸ8 6 5 07080ｸ8036ｦ003 0607 0603090304026ｦ00309 07 00076 707 0ｷ36 500036ｦ0, 0007 apple6ｦ009000706 090003000006096 50007060309 ｨｴ08070309 6ｦ0006ｦ00209096ｦ0, 0306 0503090304020ｸ8 40ｨC120 300ｨC550 200ｨC1000 0ｨC1000 05006 5096ｦ000030ｸ8 07 0900070ｹ0000ｸ800 6 5 006ｦ00ｹ06ｦ00200 07080ｸ8036ｦ0 31.4 ﾀ 17.4 16.3 ﾀ 13.5** 15.0 ﾀ 10.4 13.5 ﾀ 8.4 05006 5096ｦ000030ｸ8 oddball 35.9 ﾀ 19.8 20.4 ﾀ 15.4 16.0 ﾀ 10.7 14.8 ﾀ 9.3 05006 5096ｦ000030ｸ8 07 0900070ｹ0000ｸ800 6 5 0107000ｷ300 07080ｸ8036ｦ0 33.2 ﾀ 18.1 16.1 ﾀ 11.8** 14.3 ﾀ 9.1 13.4 ﾀ 8.3 05036ｦ000 6ｦ0apple030ｸ8 07 0900070ｹ0000ｸ800 6 5 006ｦ00ｹ06ｦ00200 07080ｸ8036ｦ0 32.9 ﾀ 15.5 17.0 ﾀ 15.2** 15.5 ﾀ 11.1 14.2 ﾀ 9.1 05036ｦ000 6ｦ0apple030ｸ8 oddball 36.2 ﾀ 18.0 17.3 ﾀ 12.9* 16.1 ﾀ 10.3 14.9 ﾀ 8.6 C036ｦ000 6ｦ0apple030ｸ8 07 0900070ｹ0000ｸ800 6 5 0107000ｷ300 07080ｸ8036ｦ0 35.9 ﾀ 21.3 18.7 ﾀ 16.5 16.0 ﾀ 10.5 14.5 ﾀ 8.8 ｨｨapple0903000ｹ06ｦ0000900. ｨｮ09060207 000006apple0700076 5 apple6ｦ06 50007 8. ｨｮ09060207 0600apple0906 ｨC 28. ｨ 6 50009 070ｹ0016ｦ00309 070303000ｹ0000007 070603076 500apple000700 070302090ｹ00900 apple006ｦ0010ｷ30909 0703 07036 50003076 5 006ｦ0 oddball- 006 5096ｦ000030ｸ8: * ｨC apple < 0.01, ** ｨC p < 0.001. 0ｹ06ｦ002090600 apple046 7 0703 apple046 76ｦ0 (apple0906. 2, 02, 03). 216 08apple0903000000000906 06036ｦ000 6ｦ0apple0300076 707 0603090304026ｦ0 6 5 040602076 509fl03 08apple070ｷ300 04apple0ｸ8 oddball (apple0906. 2, 07) 036ｦ0010800 0000 0909030000090209 apple006ｦ0010ｷ30906 000006apple07006ｦ0. ｨｦ 006ｦ00107 24 08apple00 ｨBfl6 502000009fl 006 5096ｦ0000300076 707 0603090304026ｦ0 (0307-6 707 0800 060900006 707 0ｷ36 500036ｦ0, 0007 03000000040006 090003000006096 50007060309) 6 5 086ｦ0apple6ｦ0 096 70300 oddball (apple0906.2, 05) 08apple096 5000207 01 6 50709apple6ｦ006036ｦ0-00090ｸ9 apple006ｦ0010ｷ30909, 070607 00000007 096ｦ003000300070304 6 507 6 50307apple090ｹ0000703 apple6ｦ009applefl 00 (250ｨC600 0306). ｨｮ09060207 06086ｦ00601076 5 006ｦ0 006 5096ｦ00003-000ｸ806 060309030402 6 5 086ｦ0apple6ｦ0 096 70300 oddball 6 50709apple07060207 09 0807 0703-0007040000090ｸ9 01 0909070209apple076 56ｦ00000070304 08apple090300000000090ｸ9 006 5096ｦ00003-00076 707 0603090304026ｦ0 (apple0906. 2, 02, 05). ｨ 036ｦ001, 6 50600 026ｦ0090ｸ8 07036 500036ｦ0 000006apple07006ｦ0 006ｦ0 oddball- 006 509-6ｦ000030ｸ8 04 6ｦ00000076 707 000006apple07006ｦ0 (apple0906. 2, 05) 0702000400, 0ｹ00003 0607-07036 5000306036 5040ｸ90ｸ70900 0ｹ06ｦ0060309 07036 50003076 5 006ｦ0 oddball-06036ｦ000 6ｦ0apple- 030ｸ8 09 0909070209apple076 56ｦ000000ｸ800 006ｦ00ｹ06ｦ00200000ｸ800 09 010700000ｹ0000ｸ800 006 509-6ｦ000030ｸ8 09 06036ｦ000 6ｦ0apple030ｸ8. 0407 0ｹ10307 07 09096 5000603000706 06030008000009 040009016ｦ00200000ｸ806 0602040ｹ06ｦ006 (08070ｹ103070304 030ｸ8 006 707 09 000600 09 08apple096 507-0903), 080706010702000104 0807060200 040ｸ90ｸ70906 060403036ｦ0apple000ｸ806 6ｦ0006ｦ0020909 6 5060003 20 0906060200 076 56ｦ000000ｸ803 000006apple0700076 5 (57 0600apple0906 07080ｸ803076 5, 0ｹ10307 0600apple0909 06 apple6ｦ009000ｸ80309 090003000006096 500070603fl0309 060309030402076 5 02fl 006 5096ｦ00003076 5 09 06036ｦ000 6ｦ0apple03076 5) 0000 6 50ｸ8fl6 502fl0003 016ｦ001076 707-0209 07 6 509 0903076 707 apple6ｦ00902090ｹ009fl 030008 04 07036 500036ｦ00309 006ｦ0 oddball- 006 5096ｦ000030ｸ8 09 apple006ｦ0010ｷ309fl0309 006ｦ0 apple046 70900 6 509 0ｸ8 060309030402076 5 0009 02fl 07 00076 707 0909 09000300apple6 56ｦ002076 5 6ｦ0006ｦ00209096ｦ0 (08apple090300apple0007 036ｦ001 0800, 016ｦ001 09 02fl 6 5060006 6 7apple0408080ｸ8 000006apple0700076 5 09apple0903000200000706 0107apple0ｸ8). 05036ｦ003090603090ｹ00006010906 6ｦ0006ｦ0020909 0807 036 500apple08 6ｦ00003 0ｹ10307 096ｦ0-01020ｸ90ｹ000000900. 0507096 5apple6ｦ00ｸ76ｦ0fl0600 01 6ｦ0006ｦ002090904 apple006ｦ0010ｷ30906 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apple006ｦ0010ｷ30909 6 5 07036 500036ｦ003 006ｦ0 oddball- 006 5096ｦ000030ｸ8 (036ｦ0 02. 2). 056ｦ000000ｸ800 036ｦ0 02. 2 fl060007 000307000603apple09apple040ｸ903, 0ｹ00307 080709 000906 apple6ｦ009applefl (300ｨC550 0306) 09006ｦ00ｹ009030002000007 046 50002090ｹ00000 090300000007 6 5 07036 500036ｦ003 006ｦ0 oddball- 006 5096ｦ000030ｸ8 0807 06apple6ｦ06 50000-00090ｸ9 06 apple006ｦ0010ｷ309fl0309 006ｦ0 6 50600 apple046 70900 6 509 0ｸ8 060309030402076 5. 05036ｦ003090603090ｹ00006010906 6ｦ0006ｦ0020909 0807 086ｦ0apple00070304 01apple090300apple090ｸ9 0509020107010607006ｦ0 0807016ｦ0090ｸ86 56ｦ00003, 0ｹ00307 06apple00 000000 0ｹ009060207 06086ｦ006-01076 5 6 5 6ｦ000000703 09000300apple6 56ｦ00200 02fl 07036 50003076 5 006ｦ0 oddball- 00-6 5096ｦ000030ｸ8 09006ｦ00ｹ0090307 070302090ｹ06ｦ0000306fl 016ｦ001 0703 07036 50003076 5 006ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8 (p < 0.01), 036ｦ001 09 0703 07036 50003076 5 006ｦ0 07 09-00070ｹ0000ｸ800 006ｦ00ｹ06ｦ00200000ｸ800 09 010700000ｹ0000ｸ800 006 5096ｦ000030ｸ8 09 006ｦ00ｹ06ｦ00200000ｸ800 07 0900070ｹ0000ｸ800 06036ｦ000 6ｦ0apple030ｸ8 (6 50009 00 p < 0.001). 046ｦ00ｹ06ｦ00200000ｸ806 apple6ｦ009applefl 6ｦ00800 02fl 0ｹ1030903 08026ｦ00603090ｹ0000ｸ803 000006apple0700076 5 0706036ｦ0000306fl 070ｹ0000000 06036ｦ0 090200000ｸ803. 0400 07 006ｦ0apple0408000007 apple6ｦ00902090ｹ00906 6 5 07036 500036ｦ003 006ｦ0 oddball- 006 5096ｦ000030ｸ8 09 apple046 70900 6 509 0ｸ8 060309030402076 5 006ｦ0 09000300apple6 56ｦ0026ｦ003 6ｦ0006ｦ00209096ｦ0 200ｨC1000 09 0ｨC1000 0306. ｨｦ0305ｨｬ05050004ｨ 00 ｨｺ00ｨ ｨｬ00ｨｲｨｪ02ｨｪｨｦ05 ｨｨapple076 500 0000000ｸ800 07080ｸ8030ｸ8 0807016ｦ0096ｦ00209, 0ｹ00307 08apple090300apple0007 1/3 026ｦ009000ｸ803 000006apple0700076 5 09apple0903000200000706 0107apple0ｸ8 (34%) 09 6 709080807016ｦ003086ｦ0 (37%) 01apple070209016ｦ0 0807016ｦ0090ｸ86 56ｦ00ｸ903 046 50002090ｹ000-000900 07036 500036ｦ0 006ｦ0 006 5096ｦ000030ｸ8 6 5 08apple070ｷ300 04apple00 oddball (0602040ｹ06ｦ006000ｸ800 096ｦ00300000ｸ8). 0502fl 08026ｦ00603090ｹ0000ｸ803 000006apple0700076 5 09apple09-03000200000706 0107apple0ｸ8 046 50002090ｹ000000900 apple006ｦ0010ｷ30909 08apple0709060307 0903 6 5 09000300apple6 56ｦ0026ｦ003 200ｨC500 09 200ｨC1000 0306 (apple0906. 1, 036ｦ0 02. 1), 02fl 000006apple0700076 5 6 709080807016ｦ003086ｦ0 ｨC 300ｨC550 0306 (apple0906. 2, 036ｦ0 02. 2). 050403036ｦ0apple0007 6 5 apple6ｦ00902090ｹ0000ｸ803 0600apple09fl03 07080ｸ803076 5 006ｦ0-0ｹ06ｦ00200000ｸ806 apple6ｦ009applefl (40ｨC120 0306 0703 006ｦ00ｹ06ｦ0026ｦ0 0603090304026ｦ0) 016ｦ001 02fl 000006apple0700076 5 0107apple0ｸ8, 036ｦ001 09 080702fl CA1 6 709080807016ｦ003086ｦ0 0000 0807016ｦ0096ｦ002 09006ｦ00ｹ00903076 707 0909030000000009fl 6 50002090ｹ009000ｸ8 apple6ｦ009applefl 6ｦ0 006ｦ0 oddball- 006 5096ｦ000030ｸ8 0807 06apple6ｦ06 5000000090ｸ9 06 07036 500036ｦ00309 006ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8 (036ｦ0 02. 1 09 2). 05 0ｷ300020703 006ｦ00ｹ06ｦ00200000ｸ806 apple6ｦ009applefl 0706036ｦ003070ｹ00007 06036ｦ0 09020000 6 507 6 5060003 0600apple09fl03 07080ｸ803076 5 016ｦ001 02fl 07 0ｸ70006 6 7apple0408080ｸ8 0906060200 076 56ｦ000000ｸ803 000006apple0700076 5, 036ｦ001 09 02fl 6 7apple0408080ｸ8 08026ｦ00603090ｹ0000ｸ803 010200030701. 016ｦ001 0ｸ80207 07 006ｦ0apple0408000007 6 5 08apple00 0ｸ8 040ｸ70006 apple6ｦ0 070300 [5], 0107apple07030107026ｦ0-03000003000ｸ806 006ｦ00ｹ06ｦ00200000ｸ806 apple6ｦ009applefl 0307080007 006ｦ0096 56ｦ00300 apple6ｦ009applefl- 0703 apple6ｦ00902090ｹ009fl, 0700 0703apple6ｦ0086ｦ00003 09000207apple036ｦ00ｷ3090ｸ9 07 apple6ｦ00902090ｹ009fl03 6 5 0807 6ｦ06 56ｦ000030ｸ803 0603090304026ｦ003 070300070609030002000007 0207006ｦ0 09 0702080000 0ｸ80300 06036ｦ0 090200000ｸ803. 01apple070300 03076 707, 0ｹ1036ｦ0 026ｦ0096ｦ0 07036 500036ｦ0 0000060003 09000207apple036ｦ00ｷ3090ｸ9 07 090003000006096 50007060309 060309-0304026ｦ0 [1].
13366 ｨｨ0702fl0006010906 09 apple. 050600 000006apple07000ｸ8 apple6ｦ00902090ｹ009fl 00076 50706 (09apple0903000200000706) 0107apple0ｸ8 09 6 50600 000006apple07000ｸ8 apple6ｦ00902090ｹ009fl 6 709080807016ｦ003086ｦ0 07 026ｦ0 6ｦ00ｸ903 09 000003090ｹ0000706 0603apple04010304apple0706 06086ｦ00601076 5076 707 apple6ｦ009applefl 6ｦ0, 6 5 0107-0307apple0703 6 50ｸ8060701070ｹ06ｦ006030703000ｸ806 apple6ｦ009applefl fl6 502fl000306fl 0300apple0706 0300080603090304020000076 707 apple6ｦ00902090ｹ009fl [5]. ｨｴ0307 0709006ｦ00ｹ06ｦ00003, 0ｹ00307 000006apple07000ｸ8 apple6ｦ00902090ｹ009fl 08apple0906040306036 5040ｸ903 0000 030702000107 6 5 0007-6 50706 0107apple00, 0007 09 6 5 6 709080807016ｦ0030800 (08apple00 080702070809030002000007 6 5 08070200 CA1). ｨｺ006ｦ0010ｷ30909 000006apple0700076 5 apple6ｦ00902090ｹ009fl 0000 07 006ｦ0apple04-08096 56ｦ00ｸ903 apple6ｦ0096 5090309fl 08apple096 50ｸ8016ｦ00009fl, 000006030703applefl 006ｦ0 0300076 707-01apple6ｦ00300040ｸ9 060309030402fl0ｷ3090ｸ9 (40 + 40 + 24 + 216 + 40 + 40). 070703fl apple006ｦ0010ｷ30909 006ｦ0 6 50ｸ8060701076 500apple07fl03000ｸ806 06036ｦ000 6ｦ0apple03000ｸ806 060309030402 6 5 040602076 509fl03 oddball 09 0000 046 76ｦ0066ｦ00ｸ903, apple006ｦ0010ｷ30909 006ｦ0 apple00 010900 006 5096ｦ00003000ｸ800 0603090304020ｸ8 000006010702000107 046 50002090ｹ0096 56ｦ0-0ｸ90306fl 09 0ｹ10307 046 50002090ｹ000000900 0ｹ06ｦ00603090ｹ00007 060703apple6ｦ000fl000306fl 08apple09 0909070209apple076 56ｦ000000703 08apple0903000000000909 006 5096ｦ0000300076 707 0603090304026ｦ0 6 5 0107000ｷ300 07080ｸ8036ｦ0. 016ｦ001076 5 0300036ｦ000090903 6 50709apple6ｦ006036ｦ00009fl apple006ｦ0010ｷ30906 006ｦ0 apple00 010900 006 5096ｦ00003000ｸ800 0603090304020ｸ8 08apple09 070306040306036 50909 08apple096 50ｸ8016ｦ00009fl apple006ｦ0010ｷ30906 006ｦ0 086ｦ0apple6ｦ002020002000007 006ｦ000070609030ｸ800 06036ｦ000 6ｦ0apple03000ｸ800 0603090304020ｸ8? 050709030708000ｸ803 07 ｨBfl06000000090003 060204080903 0207apple- 0309apple076 56ｦ0000900 086ｦ003fl0300076 707 060200 6ｦ0 06036ｦ000 6ｦ0apple03000ｸ803 0603090304-02076 5, 0807 6ｦ06 56ｦ000030ｸ803 06 6 50ｸ80607010706 6 500apple07fl0300070603000ｸ9 6 5 086ｦ0apple6ｦ0-096 70300 oddball. ｨｨapple09 000606036 50909 06036ｦ000 6ｦ0apple0300076 707 0603090304026ｦ0 0700 06076 5086ｦ0 6ｦ00003 0607 060200 0703, apple6ｦ0090007060300 0903 apple6ｦ06 5006ｦ0 0004020ｸ9 09 07036 50003 000006apple07006ｦ0 apple6ｦ00902090ｹ009fl 0708apple00 0002fl000306fl apple6ｦ00900070603000ｸ9 030008 04 0207000703 09 06036ｦ000 6ｦ0apple03000ｸ803 0603090304020703. ｨｨapple09 00060603-6 50909 006 5096ｦ0000300076 707 0603090304026ｦ0 060200, 0706036ｦ06 5020000000ｸ806 06036ｦ000-6ｦ0apple03000ｸ803 0603090304020703, 0000 06076 5086ｦ0 6ｦ00003 06 006 5096ｦ00003000ｸ803 060309-0304020703. ｨ 03 apple6ｦ00902090ｹ00900 060709 6ｦ00003 07080702000903000200000ｸ806 06096 7-006ｦ002, 01070307apple0ｸ806 0604030309apple04000306fl 06 06096 7006ｦ0020703 0800apple0ｷ30008-03096 500076 707 apple6ｦ00902090ｹ009fl, 6 50ｸ8096 56ｦ000000ｸ803 000606036 5090003 006 5096ｦ00003-00076 707 0603090304026ｦ0. 0207080007 08apple00 0807020708090300, 0ｹ00307 0406090200000900 06086ｦ00601076 5076 707 07036 500036ｦ0 006ｦ0 006 5096ｦ00003000ｸ806 060309030402 6 5 040602076 509fl03 oddball 6ｦ0006ｦ002076 7090ｹ00007 00006 76ｦ003096 50007060309 apple6ｦ00606076 7026ｦ006076 56ｦ0-0009fl, 07 006ｦ0apple04080000000706 6 7apple0408080706 ｨｺ. 046ｦ06ｦ0036ｦ00000006ｦ0 [2, 12ｨC 17] 08apple09 09060807020009076 56ｦ0000909 096 50401076 50ｸ803 060309030402076 5. ｨｨapple00 080702070800000900 07 030703, 0ｹ00307 046 50002090ｹ000000900 06086ｦ0060107-6 5076 707 apple6ｦ009applefl 6ｦ0 006ｦ0 006 5096ｦ00003000ｸ800 0603090304020ｸ8 6ｦ0006ｦ002076 7090ｹ00007 0300036ｦ00009090304 00006 76ｦ003096 50007060309 apple6ｦ00606076 7026ｦ006076 56ｦ00009fl, 06036ｦ06 50903 00076 50ｸ806 6 50708apple0706 07 000006apple0700000ｸ803 0300036ｦ0000909036ｦ003 0207apple0309apple07-6 56ｦ00009fl 060200 6ｦ0 09 000006apple070000076 707 0300036ｦ0000909036ｦ0 0602090ｹ0000009fl 060200-6ｦ0 086ｦ003fl0309 06 000606036 5040ｸ90ｸ70903 0603090304020703. 00060209 0909 07036 500-036ｦ0 006ｦ0 006 5096ｦ00003 6 5 040602076 509fl03 oddball 6 50ｸ80ｹ000060300 07036 50003 006ｦ0 0909070209apple076 56ｦ000000ｸ806 006 5096ｦ00003, 0307 080702040ｹ00903 6 50002090ｹ0090004 apple6ｦ009-02090ｹ009fl 030008 04 07036 500030703 006ｦ0 06036ｦ000 6ｦ0apple03 09 006 5096ｦ00003. ｨｮ0003 0702000400 070302090ｹ00900 006 5096ｦ000036ｦ0 0703 06036ｦ000 6ｦ0apple036ｦ0, 030003 070200-0400 036ｦ0016ｦ0fl apple6ｦ0090007060300. ｨｴ030703 06086ｦ00601076 50ｸ806 apple6ｦ009applefl apple6ｦ00902090ｹ009fl 030008 04 06036ｦ000 6ｦ0apple030703 09 006 5096ｦ000030703 060709 6ｦ00003 MMN (mismatch negativity, 00006 76ｦ003096 50007060300 apple6ｦ00606076 7026ｦ006076 56ｦ0-0009fl), 07 006ｦ0apple0408000000040ｸ9 ｨｺ. 046ｦ06ｦ0036ｦ00000000703 [2] 08apple09 0906060200-076 56ｦ0000909 096 50401076 50ｸ803 060309030402076 5. 0507080706036ｦ06 502flfl 080702040ｹ00000000ｸ800 006ｦ00309 6ｦ000000ｸ800 06 6ｦ000000ｸ80309 ｨｺ. 046ｦ06ｦ0036ｦ00000006ｦ0, 0307080007 06 00026ｦ00300 6 50ｸ86 507 07 6 50403 0309086ｦ003 apple6ｦ00902090ｹ00906: 0800apple- 0ｷ3000803096 5000ｸ800 apple6ｦ00902090ｹ009fl 09090300applefl0ｸ90306fl apple6ｦ000000903 06086ｦ0060107-6 50ｸ803 apple6ｦ009applefl 0703, 01070307apple0ｸ806 0107appleapple000209apple040003 06 026ｦ0090706 05ｨｨ N1. 05086ｦ00601076 50ｸ806 apple6ｦ009applefl 250ｨC600 0306 6 5 07036 5000300 006ｦ0 006 509-6ｦ000030ｸ8 fl6 502fl000306fl 060403030706 6 50307apple090ｹ000076 707 apple6ｦ009applefl 6ｦ0 09 apple6ｦ009-02090ｹ009fl 030008 04 060200 0703 0703 006 5096ｦ000036ｦ0 09 06036ｦ000 6ｦ0apple036ｦ0, 0107-0307apple0700 apple6ｦ06 50007 N2 + MMN. ｨｪ6ｦ0010903 07 apple6ｦ0090703, 01apple070300 0800apple0ｷ3000803096 5000ｸ803 apple6ｦ00902090ｹ00906 (N1) 6 50ｸ80ｹ0090602fl000306fl 000ｸ700 apple6ｦ00902090ｹ00900 030008 04 060200 0703 0703 06036ｦ000 6ｦ0apple036ｦ0 09 6ｦ00103046ｦ00200-000ｸ803 006 5096ｦ000030703. 056ｦ000000ｸ800, 080702040ｹ00000000ｸ800 6 5 0ｹ1030706 apple6ｦ0 070300 0807 000006apple07-006ｦ003, 000006010702000107 0000 06076 5086ｦ0 6ｦ00ｸ903 06 006ｦ004090309 apple0009040200036ｦ0-036ｦ00309, 080702040ｹ00000000ｸ80309 6 5 6ｦ0006ｦ002076 7090ｹ0000ｸ803 07080ｸ8036ｦ003 06 apple006 709-0603apple6ｦ00ｷ3090006 6 50ｸ8096 56ｦ000000ｸ803 08070300000ｷ3096ｦ002076 5 006ｦ0 01apple0702090100 [4]. ｨｺ6ｦ0000000 0ｸ80207 0807016ｦ0096ｦ00007, 0ｹ00307 6 5 09apple0903000200000706 0107apple00 6 5 0703-6 50003 006ｦ0 apple00 010900 006 5096ｦ000030ｸ8 08apple0709060307 090207 046 50002090ｹ000000900 6ｦ0030802090304 0ｸ8 080901076 5 P1(56ｨC80 0306) 09 P2(120ｨC140 0306), 6ｦ0 6 5 6 709080807016ｦ0030800 ｨC 080901076 5 P1, N1, P2 (80ｨC180 0306) 0807 06apple6ｦ06 5000000090ｸ9 06 07036 500036ｦ00309 006ｦ0 oddball- 006 5096ｦ000030ｸ8 09 07 09-00070ｹ0000ｸ800 006 5096ｦ000030ｸ8. 016ｦ001 0307080007 07 ｨBfl0600090300 0ｹ10309 0000-06030ｸ801076 50109? 020ｸ8 0807026ｦ06 76ｦ00003, 0ｹ00307 080706010702000104 6 5 070600076 500 6 50ｸ8096 56ｦ00000076 707 08070300000ｷ3096ｦ0026ｦ0 0200080903 060403036ｦ0 0609006ｦ00803090ｹ00006-010903 08070300000ｷ3096ｦ002076 5 0300076 70903 000006apple0700076 5, 0307 6 5 000903 03076 70403 08apple07fl6 502fl030006fl 0307 0402fl0ｷ30909 0607 060307apple07000ｸ8 apple000309010402flapple000ｸ803 09 apple046 70903 0603apple04010304apple. 05 0307 0800 6 5apple0003fl 09096 50006030007, 0ｹ00307 06086ｦ00601076 50ｸ806 apple6ｦ009applefl 000006apple07006ｦ0 6 507090009016ｦ00003 02090400 08apple09 07-0603090800000909 0807apple076 7076 5076 707 04apple076 500fl 030003 apple6ｦ00000076 707 0807030000-0ｷ3096ｦ0026ｦ0, 09 00060209 006ｦ00ｹ06ｦ00200000ｸ806 026ｦ009000ｸ806 apple6ｦ009applefl 0807fl6 5090206fl, 0307 0700 0702080000 0ｸ80300 070300070609030002000007 04060307060ｹ0096 50ｸ803, 0800apple00 6ｦ06 56ｦ0fl 066 500 000009fl 07 020909090ｹ00006010903 066 5070606036 56ｦ003 0603090304-02076 5 090209 07 apple6ｦ00902090ｹ009fl03 030008 04 0603090304026ｦ00309. ｨｬ6 50002090ｹ000-000900 apple006ｦ0010ｷ30909 000006apple0700076 5 006ｦ0 apple00 010900 006 5096ｦ00003000ｸ800 0603090304-020ｸ8 6 5 08apple070ｷ300 04apple00 oddball 016ｦ0066ｦ0000306fl 080709 000903 010703-080700000003076 5 07036 500036ｦ0 (200ｨC500, 200ｨC1000 0306), 0ｹ00307 6 7076 507apple0903, 0807-6 509 0903070304, 07 apple070209 apple000309010402flapple000ｸ803, 020903 090ｹ000-06010903 09 apple046 70903 0807 0107apple01076 50ｸ803 6 50209fl000906 006ｦ0 6ｦ00103096 50007060300 000006apple0700076 5 0107apple0ｸ8 09 6 709080807016ｦ003086ｦ0. ｨｴ0307 09 080700fl030007, 0807-06010702000104 080709 000900 026ｦ0090ｸ8 07036 50003076 5 000006apple0700076 5 0703apple6ｦ0086ｦ0-0ｸ903 0807 6 5apple0003000009 08apple070ｷ30006060ｸ8, 0703000706fl0ｸ7090006fl 01 01076 7000903096 5-000ｸ803 020400010ｷ309fl03 [1]. ｨｨ07 6 507 fl 0903076 709, 0307080007 01070006036ｦ00309apple076 56ｦ00300, 0ｹ00307 000006apple07000ｸ8 09apple0903000200000706 0107apple0ｸ8 09 080702fl CA1 6 709080807016ｦ003086ｦ0 01apple070209016ｦ0 0807016ｦ0096ｦ00209 076 7apple6ｦ000090ｹ0000000040ｸ9 08026ｦ00603090ｹ00007060300, 0ｹ00307 6 50ｸ8apple6ｦ0086ｦ0000306fl, 6 507-0800apple6 50ｸ803, 6 5 030703, 0ｹ00307 08026ｦ00603090ｹ00007-0603000ｸ9 07 026ｦ0 6ｦ00ｸ903 0000 6 50600 000006apple07000ｸ8, 6ｦ0 08apple090300apple0007 03apple000300 010200030701, 09, 6 507-6 50307apple0ｸ803, 04 0ｹ1030903 08026ｦ00603090ｹ0000ｸ803 010200030701 08apple0709060307 0903 046 50002090ｹ000000900 030702000107 080709 000903 0ｹ06ｦ006030006 0703-6 50003076 5 006ｦ0 apple00 010900 006 5096ｦ00003000ｸ800 0603090304020ｸ8 6 5 08apple070ｷ300 04apple00 oddball 0807 06apple6ｦ06 5000000090ｸ9 06 07036 500036ｦ00309 006ｦ0 0ｹ06ｦ006030ｸ800 06036ｦ000-6ｦ0apple030ｸ8. ｨｨ07 0ｷ30002070304 applefl 04 08apple0909006ｦ001076 5 0ｹ10307 0406090200000900 apple6ｦ009applefl 6ｦ0 0307080007 07030000060309 01 08apple07fl6 5020000090ｸ9 07apple0900000309apple07-6 5070ｹ000076 707 apple0002020001066ｦ0 006ｦ0 0000070809 6ｦ000000700 08apple00 ｨBfl6 50200000900 0603090304026ｦ0 [7]. ｨｬ06090200000900 080709 000903 0ｹ06ｦ006030006 07036 500036ｦ0 030708-0007 03apple6ｦ00103076 56ｦ00300 016ｦ001 07 02006 70ｹ000000900 0800apple00apple6ｦ0 07030109 0900-0207apple036ｦ00ｷ30909 08apple09 0000 0800apple00 6ｦ00ｹ000 6 5 apple046 70900 0ｹ06ｦ0060309 0307096 76ｦ0 09 096ｦ008090609 0607 0ｸ8030906 6 5 01apple6ｦ00301076 5apple00030000000706 09 07026 7076 5apple00-030000000706 086ｦ003fl0309. 050105ｨｦ0501 1. ｨｨapple0903000000000900 086ｦ0apple6ｦ0 096 7030ｸ8 oddball (0602040ｹ06ｦ006000706 096ｦ00300000ｸ8) 06 09060807020009076 56ｦ000090003 0ｷ36 50003076 50ｸ803 09apple0903000200000ｸ803 060309030402076 5 apple6ｦ009000706 090003000006096 50007060309 0807016ｦ0096ｦ00207 09006ｦ00ｹ009-030700 046 50002090ｹ000000900 080709 000903 apple006ｦ0010ｷ30906 04 08026ｦ00603090ｹ0000ｸ803 010200030701 006ｦ0 apple00 010900 006 5096ｦ000030ｸ8 0807 06apple6ｦ06 5000000090ｸ9 06 07036 500036ｦ0-0309 006ｦ0 0ｹ06ｦ006030ｸ800 06036ｦ000 6ｦ0apple030ｸ8 6 5 09000300apple6 56ｦ0026ｦ003 200ｨC500 09