Tsitologiya 55 (6)

132013 000606020902010608 0606ｨｬ 55, 016 00090608010603 01060204020506080606 ﾁ030508020102 0208000903010608 0108 0506040708060704010707 0204010801060508000607 080903060208 080204010307 0807080704070504 08 08. 08. 08ｨ 0108ｨ 01ｨｨｨｬ0609, 1 00. 08. 05ｨｮ09050609, 08. 00. 08ｨｪｨｨ09ｨｨｨｬ0609 08ｨ 04ｨ ｨｪ09ｨｺｨｨｨｦ ｨｨｨｪ09ｨｰｨｨｨｰｨｮｨｰ ｨ ｨｨ0601ｨｨｨｬｨｨｨｨ ｨｨ ｨ ｨｨ0600ｨｨ04ｨｨｨｺｨｨ 08ｨ 040100 040801, 1050502ｨｺｨｰ0806ｨｪｨｪ04ｨｦ ｨ 01080209: abdfar@mail.ru 08. 08. 08ｨ 0108ｨ 01ｨｨｨｬ0609 ｨｨ 0108. 0005ｨｨ07ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｪｨｮ06 060800ｨ ｨｪｨｨ04ｨ 02ｨｨ06 ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ ｨｺｨｮｨｺｨｮ08ｨｮ0404 08ｨｪｨ 05ｨｨ04ｨｨ080609ｨ 05ｨｨ 0905ｨｨ07ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 09 2 ｪ4 0003ｨ ｨｪｨ 080609ｨｰ ｨｨ 060800ｨ ｨｪｨｨ04ｨ 02ｨｨ06 ｨｺ0502ｨｰ06ｨｺ 070806080609ｨｰｨｺ0609 ｨｺｨｮｨｺｨｮ08ｨｮ0404. 0308ｨｨ 01ｨ 090502ｨｪｨｨｨｨ 2 0003ｨ 06ｨｰｨｬ02 ﾂ02ｨｪ04 070806090902ｨｰ0502ｨｪｨｨ02 ｨｬｨ ｨｰ08ｨｨｨｺ09ｨ ｨｬｨｨｨｰ060106ｨｪ0108ｨｨｨｦ, ｨｺ05ｨ 09ｨｰ0208ｨｨ04ｨ 02ｨｨ07 050502ｨｬ02ｨｪｨｰ0609 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0902ｨｰｨｨ 09 06ｨ 08ｨ 040609ｨ ｨｪｨｨ02ｨｬ ｨｮ ﾂｨ 09ｨｰｨｺ0609 050800ｨ 09ｨｰ060705ｨ 04ｨｬ04 ｨｨ ｨｮ090205ｨｨ ﾂ02ｨｪｨｨ02 09 02ｨｨｨｰ060705ｨ 04ｨｬ02 ﾂｨｨ0905ｨ ｨｬ0205ｨｺｨｨ01 09ｨ ｨｺｨｮ060502ｨｦ. 01ｨ ｨｰ08ｨ ｨｪ09-09ｨｰ060806ｨｪ02 01ｨｨｨｺｨｰｨｨ060906ｨｬ 04ｨ 00ｨｨｨｺ09ｨｨ080609ｨ ｨｪ06 ｨｪｨ ｨｺ06070502ｨｪｨｨ02 090204ｨｨｨｺｨｮ05 09 050502ｨｺｨｰ0806ｨｪｨｪ06-070506ｨｰｨｪ04ｨｬ 090601020803ｨｨｨｬ04ｨｬ, ﾂｨｰ06 ｨｮｨｺｨ 040409ｨ 02ｨｰ ｨｪｨ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ02 ｨｰ08ｨ 00ｨｨｨｺｨ 09 05ｨｪ0106ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ06ｨｦ 09ｨｨ09ｨｰ02ｨｬ02. 040201ｨｮｨｺ02ｨｨ07 050502ｨｺｨｰ0806ｨｪｨｪ06ｨｦ 070506ｨｰｨｪ0609ｨｰｨｨ ｨｬ02ｨｬｨ 08ｨ ｨｪ04 ｨｰ06ｨｪ060705ｨ 09ｨｰｨ 090607080609060301ｨ 05ｨ 09ｨｹ ｨｮｨｬ02ｨｪｨｹ0302ｨｪｨｨ02ｨｬ 09 09ｨ ｨｺｨｮ06050701 ｨｺ0605ｨｨ ﾂ0209ｨｰ09ｨ 0609ｨｬｨｨ0600ｨｨ05ｨｹｨｪ0401 09ｨｺ0506 ﾂ02ｨｪｨｨｨｦ. 030609040302ｨｪｨｨ02 01ｨ 090502ｨｪｨｨ07 0106 4 0003ｨ 0706ｨｺｨ 04ｨ 0506 090604ｨｪｨｨｨｺｨｪ060902ｨｪｨｨ02 ｨ 05ｨｹｨｰ0208ｨ 02ｨｨｨｦ 010209ｨｰ08ｨｮｨｺｨｰｨｨ09ｨｪ060006 01ｨ 08ｨ ｨｺｨｰ0208ｨ 09 ｨｰ06ｨｪ060705ｨ 09ｨｰ02. 01ｨｨ 09 0601ｨｪ06ｨｬ ｨｨ04 09ｨ 08ｨｨｨ ｨｪｨｰ0609 ｨｪ02 090407090507050609ｨｹ ｨｪｨ 08ｨｮ0302ｨｪｨｨ07 0202050609ｨｰｨｪ0609ｨｰｨｨ 0705ｨ 04ｨｬｨ 0502ｨｬｨｬ04, ﾂｨｰ06 070601ｨｰ0902080301ｨ 050609ｨｹ 06ｨｺ08ｨ 09ｨｺ06ｨｦ 0609ｨ ｨｪ0906ｨｬ 000605ｨｮｨ 04ｨｬ. 0308ｨｨ 09ｨ 08060902 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 0106 ｨ ｨｰｨｬ0609000208ｨｪ060006 ﾂｨ 09ｨｰ06ｨｰｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｪ0401 ｨｪｨ 08ｨｮ0302ｨｪｨｨｨｦ ｨｬ02ｨｬｨ 08ｨ ｨｪ04 ｨｰ06ｨｪ060705ｨ 09ｨｰｨ ｨ 0405ｨ 07080607060802ｨｨ06ｨｪｨ 05ｨｹｨｪｨ 08ｨ 04ｨｪｨｨ0202 01ｨ 090502ｨｪｨｨｨｦ: ｨｬｨｨｨｪｨｨｨｬｨ 05ｨｹｨｪ06ｨｦ 0708ｨｨ 09ｨ 08060902 06ｨｰ 2 0106 0 0003ｨ ｨｨ ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪ06ｨｦ 0708ｨｨ 09ｨ 08060902 06ｨｰ 4 0106 0 0003ｨ. 06ｨ ｨｺｨｨｨｬ 06ｨ 08ｨ 0406ｨｬ, ｨｬ04 010205ｨ 02ｨｬ 0904090601 06 ｨｰ06ｨｬ, ﾂｨｰ06 0601ｨｪ06ｨｦ ｨｨ04 ｨｬｨｨ0302ｨｪ02ｨｦ 0905ｨｨ07ｨｪｨｨ07 01ｨ 090502ｨｪｨｨ07 ｨｪｨ ｨｺ0502ｨｰｨｺｨｨ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ 0709050702ｨｰ0907 05ｨｪ0106ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪｨ 07 09ｨｨ09ｨｰ02ｨｬｨ. 080506 ﾂ02090402 09050609ｨ :01ｨ 090502ｨｪｨｨ02, 09ｨｰ08020909, 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺｨ 07 0902ｨｰｨｹ, 070806080609ｨｰｨｺｨｨ ｨｺｨｮｨｺｨｮ08ｨｮ0404. 0604090209ｨｰｨｪ06, ﾂｨｰ06 09ｨｪ0203ｨｪ0202 01ｨ 090502ｨｪｨｨ02 0905ｨｨ0702ｨｰ ｨｪｨ 00ｨｮｨｪｨｺ02ｨｨ06ｨｪｨ 05ｨｹｨｪ04ｨｦ 09ｨｰｨ ｨｰｨｮ09 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨｨ 03ｨｨ0906ｨｰｨｪ0401. 08ｨ ｨｺ ｨｰ0208ｨｬ0601ｨｨｨｪｨ ｨｬｨｨ ﾂ0209ｨｺｨｨｨｦ 07ｨ 08ｨ ｨｬ02ｨｰ08 01ｨ 090502ｨｪｨｨ02 09 ｨ ｨｨ060900020802 0502ｨｬ05ｨｨ 09ｨ 08ｨｹｨｨ08ｨｮ02ｨｰ 06ｨｰ 0.03 0003ｨ ｨｪｨ 09040906ｨｰ02 06ｨｺ060506 10 ｨｺｨｬ 0106 ｨ 06050202 ﾂ02ｨｬ 100 0003ｨ 09 06ｨｺ02ｨ ｨｪ09ｨｺｨｨ01 0005ｨｮｨ ｨｨｨｪｨ 01. 020507 08ｨ 09ｨｰ02ｨｪｨｨｨｦ 0706ｨｺｨ 04ｨ ｨｪ06, ﾂｨｰ06 0607ｨｰｨｨｨｬｨｮｨｬ ｨ ｨｰｨｬ0609000208ｨｪ060006 01ｨ 090502ｨｪｨｨ07 050203ｨｨｨｰ ｨｪｨ 09040906ｨｰｨ 01 1000 ｪ2800 ｨｬ, 000102 ｨｨ 06ｨ ｨｪｨ 08ｨｮ03ｨｨ09ｨ 02ｨｰ0907 ｨｨ01 ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪｨ 07 07080601ｨｮｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｹ. 05 ﾂｨｨｨｰｨ 02ｨｰ0907, ﾂｨｰ06 070601ｨ 090502ｨｪｨｨ02 ｨｰ08ｨ ｨｪ09070608ｨｰｨ, 0006ｨｰ0609ｨｨｨｪｨｰ0204ｨ ｨｨ 080609ｨｰｨ ｨｪｨｨ0302 09ｨｮｨ ｨ 05ｨｹ07ｨｨｨｦ09ｨｺ060006 07060709ｨ 09090704ｨ ｨｪ06 ｨｨｨｬ02ｨｪｨｪ06 09 070609040302ｨｪｨｨ02ｨｬ ｨ ｨｰｨｬ0609000208ｨｪ060006 01ｨ 090502ｨｪｨｨ07 (01ｨ ｨｬｨ 0502ｨｦ, 2004). 030807ｨｬｨ 07 00ｨｮｨｪｨｺ02ｨｨ06ｨｪｨ 05ｨｹｨｪｨ 07 080605ｨｹ 01ｨ 090502ｨｪｨｨ07 08ｨ 0909ｨｬｨ ｨｰ08ｨｨ09ｨ 02ｨｰ0907 070802030102 0909020006 09 ｨｰ06 ﾂｨｺｨｨ 040802ｨｪｨｨ07 0109ｨｨ03ｨｮｨｴ02ｨｦ 09ｨｨ0504 ｨｬｨ 09090609060006 07020802ｨｪ0609ｨ 090601ｨｪ0401 08ｨ 09ｨｰ0906080609 (Knoblauch, Peters, 2010). 00 0209ｨｰ0209ｨｰ0902ｨｪｨｪ0401 ｨｮ09050609ｨｨ0701 01ｨｨｨ 07ｨ 0406ｨｪ 090601ｨｪ060006 0706ｨｰ02ｨｪ02ｨｨｨ 05ｨ ｨｺ0502ｨｰ06ｨｺ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ 09 04ｨ 09ｨｨ09ｨｨｨｬ0609ｨｰｨｨ 06ｨｰ 09ｨｨ01ｨ ｨｺ060502ｨ 0502ｨｰ0907 06ｨｰ ｨC0.5 0106 ｨC(4 ｪ5) M03a ｪ 0708ｨｨ 09ｨｨ05ｨｹｨｪ06ｨｬ 06ｨ 0204090603ｨｨ09ｨ ｨｪｨｨｨｨ (Galm*es et al., 2011). 05ｨｪｨｨ0302ｨｪｨｨ02 01ｨ 090502ｨｪｨｨ07 ｨｪｨｨ0302 00ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨ01 04ｨｪｨ ﾂ02ｨｪｨｨｨｦ 090607080609060301ｨ 02ｨｰ0907 090208ｨｹ0204ｨｪ04ｨｬｨｨ 070608ｨ 0302ｨｪｨｨ07ｨｬｨｨ ｨｬ02ｨｰｨ ｨ 0605ｨｨ04ｨｬｨ ｨｨ 00ｨｨｨ 0205ｨｹ06 08ｨ 09ｨｰ02ｨｪｨｨｨｦ. 010209ｨｬ06ｨｰ0807 ｨｪｨ 0706 ﾂｨｰｨｨ 09ｨｰ060502ｨｰｨｪ0606 ｨｨ09ｨｰ0608ｨｨ06 ｨｨ04ｨｮ ﾂ02ｨｪｨｨ07 01ｨ 05ｨｹｨｪ020006 ｨｰ08ｨ ｨｪ09070608ｨｰｨ 08ｨ 09ｨｰ0906080609, 090607080609 06 0008ｨ 01ｨｨ02ｨｪｨｰｨ 01 01ｨ 090502ｨｪｨｨｨｦ, ｨｰ0802ｨ ｨｮ06ｨｴｨｨ010907 010507 07020802ｨｪ0609ｨ 08ｨ 09ｨｰ0906080609 ｨｬ020301ｨｮ ｨ ｨｺ020207ｨｰ060806ｨｬ ｨｨ 0106ｨｪ060806ｨｬ, 0609ｨｰｨ 02ｨｰ0907 ｨｪ0208020302ｨｪｨｪ04ｨｬ (Knoblauch, Peters, 2010; Turgeon, 2010). 0604ｨｬ02ｨｪ02ｨｪｨｨ02 01ｨ 090502ｨｪｨｨ07 09 ｨｺ0502ｨｰ06 ﾂｨｪ0401 ｨｺ06ｨｬ07ｨ 08ｨｰｨｬ02ｨｪｨｰｨ 01 050203ｨｨｨｰ 09 0609ｨｪ060902 080200ｨｮ050702ｨｨｨｨ ｨｬｨｪ06030209ｨｰ09ｨ 00ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨ01 070806020209090609: ｨｮ09ｨｰｨｹｨｨ ﾂｨｪ0401 0109ｨｨ0302ｨｪｨｨｨｦ (Ache et al., 2010), 080609ｨｰｨ (Abe, Horikoshi, 1995), ｨｪｨ 09ｨｰｨｨｨｦ (00ｨ ｨｦｨｪｨ 08, 1987) ｨｨ ｨｰ. 01. 00 070609050201ｨｪ0202 090802ｨｬ07 09ｨｪｨｨｨｬｨ ｨｪｨｨ02 ｨｺ 01ｨ 090502ｨｪｨｨ06 08ｨ 09ｨｰ02ｨｰ 09 09090704ｨｨ 09 00ｨｨ0706ｨｰ020406ｨｦ 06 09ｨｨ00ｨｪｨ 05ｨｹｨｪ06ｨｦ 00ｨｮｨｪｨｺ02ｨｨｨｨ 090605ｨｪ 01ｨ 090502ｨｪｨｨ07. 0001ｨｨ0108ｨ 0905ｨｨ ﾂ0209ｨｺｨ 07 00ｨｨ0706ｨｰ0204ｨ 03 07080201070605ｨ 00ｨ 02ｨｰ, ﾂｨｰ06 ｨ 0409ｨｰ0806 08ｨ 0907080609ｨｰ08ｨ ｨｪ0706ｨｴｨｨ020907 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 00ｨｨ01080601ｨｨｨｪｨ ｨｬｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｬ0600ｨｮｨｰ ｨ 04ｨｰｨｹ 07020802ｨｪ0609 ﾂｨｨｨｺ06ｨｬ 09ｨｨ00ｨｪｨ 05ｨｹｨｪ06ｨｦ ｨｨｨｪ000608ｨｬｨ 02ｨｨｨｨ 09 ｨ ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨ01 06ｨ ｨｲ02ｨｺｨｰｨ 01 (Dziubinska, 2003). 00 070802010205ｨ 01 070806ｨ 0502ｨｬ04 0905ｨｨ07ｨｪｨｨ07 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 03ｨｨ090402 060800ｨ ｨｪｨｨ04ｨｬ04 ｨｪｨ ｨｨｨ 06050202 ｨｨ04ｨｮ ﾂ02ｨｪ04 05000002ｨｺｨｰ04 09040906ｨｺ060006 01ｨ 090502ｨｪｨｨ07, 0706-09ｨｨ01ｨｨｨｬ06ｨｬｨｮ 0706ｨｰ06ｨｬｨｮ, ﾂｨｰ06 09040906ｨｺ0602 01ｨ 090502ｨｪｨｨ02 03ｨｨ0806ｨｺ06 0708ｨｨｨｬ02ｨｪ0702ｨｰ0907 09 ｨ ｨｨ06ｨｰ0201ｨｪ06050600ｨｨｨｨ 010507 09ｨｰ0208ｨｨ05ｨｨ04ｨ 02ｨｨｨｨ 09ｨ ｨｺ02ｨｨｨｪ ｨｨ ｨｺ06ｨｪ09020809ｨ 02ｨｨｨｨ 07ｨｨｨｴ02090401 07080601ｨｮｨｺｨｰ0609 (Rothschild, Mancinelli, 2001). 00 ｨｬｨｪ060006 ﾂｨｨ090502ｨｪｨｪ0401 08ｨ ｨ 06ｨｰｨ 01, 0904070605ｨｪ02ｨｪｨｪ0401 0708ｨｨ 01ｨ 090502ｨｪｨｨｨｨ 40 ｪ100 0003ｨ ｨｨ 09040302, 09 06ｨ 05ｨ 09ｨｰ0701, 000102 00ｨｮｨｪｨｺ02ｨｨ06ｨｪｨ 05ｨｹｨｪ0402 ｨｪｨ 08ｨｮ0302ｨｪｨｨ07 09ｨｰｨ ｨｪ060907ｨｰ0907 06 ﾂ0209ｨｨ01ｨｪ04ｨｬｨｨ, ｨ 0405ｨｨ 090407090502ｨｪ04 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 09 05ｨｨ07ｨｨ01ｨｪ0401 0905060701 ｨｬ02ｨｬｨ 08ｨ ｨｪ, 01ｨｨ09090602ｨｨｨ 02ｨｨ07 0605ｨｨ0006ｨｬ0208ｨｪ0401 ｨ 0205ｨｺ06090401 ｨｺ06ｨｬ070502ｨｺ090609 ｨｨ 0706ｨｰ020807 ｨｨｨｬｨｨ 000208ｨｬ02ｨｪｨｰｨ ｨｰｨｨ09ｨｪ06ｨｦ ｨ ｨｺｨｰｨｨ09ｨｪ0609ｨｰｨｨ (Gross, Jaenicke, 1994; Molina-Garcia, 2002), ｨ ｨｰｨ ｨｺ0302 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 09 05ｨｺ090708020909ｨｨｨｨ 0002ｨｪ06ｨｬｨ (Fernandes et al., 2004). 01ｨ 08ｨｮ0302ｨｪｨｨ07 09 ｨｺ08060902ｨｪ0609ｨｪ06ｨｦ ｨｨ ｨｪ020809ｨｪ06ｨｦ 09ｨｨ09ｨｰ02ｨｬｨ 01 0708ｨｨ 00ｨｨ070208ｨ ｨ 08ｨｨ ﾂ0209ｨｺｨｨ01 ｨｰ08ｨ 09ｨｬｨ 01 0709050706ｨｰ0907 0708ｨｨ ﾂｨｨｨｪ06ｨｦ 00ｨｨｨ 0205ｨｨ 03ｨｨ0906ｨｰｨｪ0401 (05ｨ 05ｨｹ02ｨｬｨ ｨｪ ｨｨ 0108., 1979). 06ｨｬ0206ｨｰ0907 01ｨ ｨｪｨｪ0402 06 ｨｰ06ｨｬ, ﾂｨｰ06 00ｨｨ070208ｨ ｨ 08ｨｨ ﾂ0209ｨｺｨｮ06 00ｨｨｨ 0205ｨｹ 060800ｨ ｨｪｨｨ04ｨｬ0609 ｨｬ0603ｨｪ06 0708020106ｨｰ0908ｨ ｨｰｨｨｨｰｨｹ 0708020109ｨ 08ｨｨｨｰ0205ｨｹｨｪ06ｨｦ 06ｨ 08ｨ ｨ 06ｨｰｨｺ06ｨｦ ｨｨ01 09ｨｰ08020909060904ｨｬｨｨ 000608ｨｬ06ｨｪｨ ｨｬｨｨ (Tanino et al., 1992). 00 02020506ｨｬ 09050201ｨｮ02ｨｰ 06ｨｰｨｬ02ｨｰｨｨｨｰｨｹ, ﾂｨｰ06, ｨｪ0209ｨｬ06ｨｰ0807 ｨｪｨ ｨ ｨｺｨｰｨｮｨ 05ｨｹｨｪ0609ｨｰｨｹ 070806ｨ 0502ｨｬ04, ｨｨ04 09090201 07ｨ 08ｨ ｨｬ02ｨｰ080609 09ｨｪ0203ｨｪ02ｨｦ 0908020104 0905ｨｨ07ｨｪｨｨ02 ｨｨｨｬ02ｨｪｨｪ06 01ｨ 090502ｨｪｨｨ07 ｨｪｨ ｨ ｨｨ0601ｨｨｨｬｨｨ06 ｨｨ 00ｨｨ04ｨｨ06050600ｨｨ06 03ｨｨ090401 060800ｨ ｨｪｨｨ04ｨｬ0609, 09 ｨｰ06ｨｬ ﾂｨｨ090502 08ｨ 09ｨｰ02ｨｪｨｨｨｦ, ｨｨ04ｨｮ ﾂ02ｨｪ06 09 ｨｪｨ ｨｨｨｬ02ｨｪｨｹ0302ｨｦ 09ｨｰ020702ｨｪｨｨ (01ｨ ｨｬｨ 0502ｨｦ, 2004). 030605ｨｰ06ｨｬｨｮ 020205ｨｹ06 ｨｪｨ 0302ｨｦ 08ｨ ｨ 06ｨｰ04 ｨ 040506 060202ｨｪｨｨｨｰｨｹ 0102ｨｦ09ｨｰ09ｨｨ02 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ ｨｮ05ｨｹｨｰ08ｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｮ ｨｺ0502ｨｰ06ｨｺ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ. 414

130005ｨｨ07ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｪｨｮ06 060800ｨ ｨｪｨｨ04ｨ 02ｨｨ06 ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ ｨｺｨｮｨｺｨｮ08ｨｮ0404 415 04ｨｨ09. 1. 0005ｨｨ07ｨｪｨｨ02 09ｨｪ0203ｨｪ020006 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 080609ｨｰ 070806080609ｨｰｨｺ0609 ｨｺｨｮｨｺｨｮ08ｨｮ0404. ｨ ｪ ﾂ02080204 5 ﾂ 0706090502 070609040302ｨｪｨｨ07 01ｨ 090502ｨｪｨｨ07 0106 4 0003ｨ 08ｨ 04ｨｪｨｨ02ｨ 0105ｨｨｨｪ04 070806080609ｨｰｨｺ0609 ｨｬ020301ｨｮ ｨｺ06ｨｪｨｰ080605ｨｹｨｪ04ｨｬｨｨ ｨｨ 060704ｨｰｨｪ04ｨｬｨｨ 06ｨ 08ｨ 0402ｨ ｨｬｨｨ ｨ 0405ｨ ｨｪ02010609ｨｰ06090208ｨｪ06ｨｦ; ｨ ｪ ﾂ02080204 4 09ｨｮｨｰ 0706090502 09ｨｪｨｨ0302ｨｪｨｨ07 01ｨ 090502ｨｪｨｨ07 0106 ｨ ｨｰｨｬ0609000208ｨｪ060006 0105ｨｨｨｪｨ ｨｪｨ 010402ｨｬｨｪ06ｨｦ ｨｨ 0706010402ｨｬｨｪ06ｨｦ ﾂｨ 09ｨｰ02ｨｦ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ 090609ｨｰｨ 09050705ｨ 06ｨｺ060506 76 ｨｨ 68 % 06ｨｰ ｨｺ06ｨｪｨｰ08060507 090606ｨｰ0902ｨｰ09ｨｰ0902ｨｪｨｪ06. 00ｨ ｨｰ0208ｨｨｨ 05 ｨｨ ｨｬ02ｨｰ0601ｨｨｨｺｨ 02ｨ ｨｲ02ｨｺｨｰ06ｨｬ ｨｨ09090502010609ｨ ｨｪｨｨｨｦ 0905ｨｮ03ｨｨ05ｨｨ 4 ｪ7-01ｨｪ0209ｨｪ0402 08ｨ 09ｨｰ02ｨｪｨｨ07 ｨｺｨｮｨｺｨｮ08ｨｮ0404 Zea mays L. 090608ｨｰ0609 08ｨｪ06ｨｰｨ ｨｨ 08ｨｮｨ ｨ ｨｪｨｹ. 06ｨｰｨｨ0605ｨｨ080609ｨ ｨｪｨｪ0402 08ｨ 09ｨｰ02ｨｪｨｨ07 090408ｨ ｨｴｨｨ09ｨ 05ｨｨ09ｨｹ 00ｨｨ0108060706ｨｪｨｪ04ｨｬ ｨｬ02ｨｰ060106ｨｬ ｨｪｨ 09060106070806090601ｨｪ06ｨｦ 09060102 0708ｨｨ ｨｺ06ｨｬｨｪｨ ｨｰｨｪ06ｨｦ ｨｰ02ｨｬ070208ｨ ｨｰｨｮ0802 09 0705ｨ 09ｨｰｨｨｨｺ06090401 100-ｨｬｨｨ0505ｨｨ05ｨｨｨｰ0806090401 02ｨｬｨｺ0609ｨｰ0701. 020507 ｨ ｨｪｨ 05ｨｨ04ｨ 0905ｨｨ07ｨｪｨｨ07 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 080609ｨｰ06090402 0708060202090904 ｨｨｨｪｨｰｨ ｨｺｨｰｨｪ0402 08ｨ 09ｨｰ02ｨｪｨｨ07 09 0908020102 090408ｨ ｨｴｨｨ09ｨ ｨｪｨｨ07 09ｨｬ0209ｨｰ02 09 ｨｺ06ｨｪｨｰ02ｨｦｨｪ020806ｨｬ 0706ｨｬ02ｨｴｨ 05ｨｨ 09 ｨｺｨ ｨｬ0208ｨｮ 09040906ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｨ 07060109020800ｨ 05ｨｨ 0906040102ｨｦ09ｨｰ09ｨｨ06 01ｨ 090502ｨｪｨｨ07 2 ｨｨ05ｨｨ 4 0003ｨ 09 ｨｰ02 ﾂ02ｨｪｨｨ02 5 ﾂ. 0806ｨｪｨｰ080605ｨｹｨｪ0402 06ｨ 08ｨ 040204 0706ｨｬ02ｨｴｨ 05ｨｨ 09 ｨｰｨ ｨｺｨｮ06 0302 ｨｺｨ ｨｬ0208ｨｮ 0708ｨｨ ｨ ｨｰｨｬ0609000208ｨｪ06ｨｬ 01ｨ 090502ｨｪｨｨｨｨ. 0004ｨ 0608 090205ｨｨ ﾂｨｨｨｪ04 01ｨ 090502ｨｪｨｨ07, ｨ 05ｨｨ04ｨｺ06ｨｦ ｨｺ 09020801ｨｪ02ｨｦ 0008ｨ ｨｪｨｨ0202 00ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺ060006 01ｨｨｨ 07ｨ 0406ｨｪｨ, ｨ 0405 07080601ｨｨｨｺｨｰ0609ｨ ｨｪ 030205ｨ ｨｪｨｨ02ｨｬ 070605ｨｮ ﾂｨｨｨｰｨｹ ｨｪｨ ｨｨｨ 0605ｨｹ03ｨｨｨｦ ｨｺ06ｨｪｨｰ08ｨ 09ｨｰ 060704ｨｰｨｪ0401 ｨｨ ｨｺ06ｨｪｨｰ080605ｨｹｨｪ0401 06ｨ 08ｨ 04020609. 02ｨ 090502ｨｪｨｨ02 09060401ｨ 09ｨ 05ｨｨ 09 0706ｨｬ06ｨｴｨｹ06 07ｨｪ0209ｨｬ0609ｨｨ09ｨｰ02ｨｬ04, 090609ｨｰ0607ｨｴ02ｨｦ ｨｨ04 ｨ ｨ 050506ｨｪ0609 0903ｨ ｨｰ060006 09060401ｨｮ01ｨ, 080201ｨｮｨｺｨｰ0608ｨ 01ｨ 090502ｨｪｨｨ07 ｨｨ 050502ｨｺｨｰ0806ｨｺ05ｨ 07ｨ ｨｪ0609 070601ｨ ﾂｨｨ ｨｨ 09ｨ 080609ｨ 01ｨ 090502ｨｪｨｨ07, 06ｨ ｨｲ0201ｨｨｨｪ02ｨｪｨｪ0401 07ｨｪ0209ｨｬ06ｨｬｨ 00ｨｨ09ｨｰ08ｨ 0507ｨｬｨｨ 09 ｨｺｨ ｨｬ020806ｨｦ 01ｨ 090502ｨｪｨｨ07 (08ｨｪｨｨ09ｨｨｨｬ0609 ｨｨ 0108., 2012). 00 08070102 060704ｨｰ0609 00ｨｨｨｺ09ｨ 02ｨｨ06 06ｨ 08ｨ 04020609 010507 050502ｨｺｨｰ0806ｨｪｨｪ06ｨｦ ｨｬｨｨｨｺ080609ｨｺ0607ｨｨｨｨ 070806090601ｨｨ05ｨｨ ｨｪ0207060908020109ｨｰ0902ｨｪｨｪ06 09 ｨｺｨ ｨｬ020802 ｨ 0204 09ｨ 080609ｨ 01ｨ 090502ｨｪｨｨ07 0706090502 05ｨｰｨ 07ｨ 090401020803ｨｨ09ｨ ｨｪｨｨ07 06ｨ 08ｨ 04020609 070601 01ｨ 090502ｨｪｨｨ02ｨｬ (30 ｨｬｨｨｨｪ). 00 0108ｨｮ00ｨｨ01 09ｨ 08ｨｨｨ ｨｪｨｰｨ 01 00ｨｨｨｺ09ｨ 02ｨｨ06 070806090601ｨｨ05ｨｨ 0706090502 09ｨ 080609ｨ 01ｨ 090502ｨｪｨｨ07 0106 ｨ ｨｰｨｬ0609000208ｨｪ060006. 020507 050502ｨｺｨｰ0806ｨｪｨｪ06-ｨｬｨｨｨｺ080609ｨｺ0607ｨｨ ﾂ0209ｨｺ060006 ｨｨ09090502010609ｨ ｨｪｨｨ07 ｨ 07ｨｨｨｺｨ 05ｨｹｨｪ0402 0008ｨ 00ｨｬ02ｨｪｨｰ04 ｨｺ0608ｨｪ07 0105ｨｨｨｪ06ｨｦ 7 ｪ8 ｨｬｨｬ 00ｨｨｨｺ09ｨｨ080609ｨ 05ｨｨ 12 ﾂ 09 2.5%-ｨｪ06ｨｬ 08ｨ 09ｨｰ09060802 0005ｨｮｨｰｨ 08ｨ 05ｨｹ010200ｨｨ01ｨ ｨｪｨ 0.1 00 00060900ｨ ｨｰｨｪ06ｨｬ ｨ ｨｮ00020802 (0801 7.2) ｨｨ 070609ｨｰ00ｨｨｨｺ09ｨｨ080609ｨ 05ｨｨ 2 ﾂ 09 1%-ｨｪ06ｨｬ OsO 4 ｨｪｨ ｨｰ06ｨｬ 0302 ｨ ｨｮ00020802 09 0106ｨ ｨ 090502ｨｪｨｨ02ｨｬ 09ｨ 01ｨ 08060404 (34 ｨｬ00/ｨｬ05). 02ｨ 08ｨ 040204 010200ｨｨ0108ｨ ｨｰｨｨ080609ｨ 05ｨｨ 09 08ｨ 09ｨｰ090608ｨ 01 0907ｨｨ08ｨｰｨ 09060408ｨ 09ｨｰｨ 06ｨｴ02ｨｦ ｨｺ06ｨｪ0202ｨｪｨｰ08ｨ 02ｨｨｨｨ (30 ｪ70 %) ｨｨ ｨ 0202ｨｰ06ｨｪｨ (70 ｪ100 %) 09 070609050201ｨｮ06ｨｴｨｨｨｬ 04ｨ ｨｬ02ｨｴ02ｨｪｨｨ02ｨｬ ｨ 0202ｨｰ06ｨｪｨ 06ｨｺｨｨ09ｨｹ06 07080607ｨｨ0502ｨｪｨ (45 ｨｬｨｨｨｪ). 050200ｨｬ02ｨｪｨｰ04 ｨｺ0608ｨｪ07 07080607ｨｨｨｰ0409ｨ 05ｨｨ 050706ｨｺ09ｨｨ01ｨｪ06ｨｦ 09ｨｬ060506ｨｦ 060706ｨｪ-812 (010208ｨｬｨ ｨｪｨｨ07) ｨｨ 070605ｨｨｨｬ0208ｨｨ040609ｨ 05ｨｨ 0708ｨｨ 37, 45 ｨｨ 57 縺05 0706 24 ﾂ. 030605ｨｮｨｰ06ｨｪｨｺｨｨ02 ｨｨ ｨｮ05ｨｹｨｰ08ｨ ｨｰ06ｨｪｨｺｨｨ02 0908020404 070605ｨｮ ﾂｨ 05ｨｨ ｨｪｨ ｨｬｨｨｨｺ0806ｨｰ06ｨｬ02 LKB-III (01090202ｨｨ07), ｨｺ06ｨｪｨｰ08ｨ 09ｨｰｨｨ080609ｨ 05ｨｨ 08ｨ 09ｨｰ09060806ｨｬ ｨｮ08ｨ ｨｪｨｨ05-ｨ 0202ｨｰｨ ｨｰｨ (Tander, 1990) ｨｨ 02ｨｨｨｰ08ｨ ｨｰ06ｨｬ 0909ｨｨｨｪ02ｨ (Reynolds, 1963). 03080207ｨ 08ｨ ｨｰ04 07080609ｨｬｨ ｨｰ08ｨｨ09ｨ 05ｨｨ 09 0607ｨｰｨｨ ﾂ0209ｨｺ06ｨｬ ｨｬｨｨｨｺ080609ｨｺ060702 NU-2 (010208ｨｬｨ ｨｪｨｨ07) ｨｨ 09 050502ｨｺｨｰ0806ｨｪｨｪ06ｨｬ ｨｬｨｨｨｺ080609ｨｺ060702 JEM-1200 0309 (080706ｨｪｨｨ07). 020704ｨｰ04 070806090601ｨｨ05ｨｨ 09 3-ｨｺ08ｨ ｨｰｨｪ06ｨｦ ｨ ｨｨ06050600ｨｨ ﾂ0209ｨｺ06ｨｦ 070609ｨｰ0608ｨｪ0609ｨｰｨｨ. 01ｨ 08ｨｨ09ｨｮｨｪｨｺｨ 01 0708ｨｨ09020102ｨｪ04 09080201ｨｪｨｨ02 04ｨｪｨ ﾂ02ｨｪｨｨ07 09 ｨｨ01 09ｨｰｨ ｨｰｨｨ09ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0603ｨｨｨ ｨｺ06ｨｦ. 040204ｨｮ05ｨｹｨｰｨ ｨｰ04 05060005ｨ 09ｨｪ06 070605ｨｮ ﾂ02ｨｪｨｪ04ｨｬ 01ｨ ｨｪｨｪ04ｨｬ, ｨｮ090205ｨｨ ﾂ02ｨｪｨｨ02 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 0708ｨｨ090601ｨｨ0506 ｨｺ 09ｨｪｨｨ0302ｨｪｨｨ06 09ｨｺ06080609ｨｰｨｨ 080609ｨｰｨ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ. 010207060908020109ｨｰ0902ｨｪｨｪ06 ﾂ02080204 5 ﾂ 0706090502 ｨｪｨ 0106030102ｨｪｨｨ07 09 ｨ ｨ 0806ｨｺｨ ｨｬ020802 09080201ｨｪ0707 0105ｨｨｨｪｨ ｨｺ0608ｨｪ07 ｨｨ 0706ｨ 0200ｨ ｨ 0405ｨ ｨｪ02010609ｨｰ06090208ｨｪ06 ｨｪｨｨ0302, ﾂ02ｨｬ ｨｮ ｨｺ06ｨｪｨｰ080605ｨｹｨｪ0401 08ｨ 09ｨｰ02ｨｪｨｨｨｦ (08ｨｨ09. 1, ｨ ), ｨｪ06 ｨｪｨ 4-02 09ｨｮｨｰ 08ｨ 04ｨｪｨｨ02ｨ 090609ｨｰｨ 09050705ｨ 06ｨｺ060506 70 % 06ｨｰ ｨｺ06ｨｪｨｰ08060507 ｨｨ ｨ 0405ｨ 010609ｨｰ06090208ｨｪ06ｨｦ (08ｨｨ09. 1, ｨ ). 08ｨｪｨ 05ｨｨ04 01ｨｨｨｪｨ ｨｬｨｨｨｺｨｨ 080609ｨｰｨ 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ02ｨｰ 06 ｨ 0409ｨｰ0806ｨｬ ｨｨ 070605ｨｪ06ｨｬ 09060909ｨｰｨ ｨｪ06090502ｨｪｨｨｨｨ 09ｨｺ06080609ｨｰｨｨ 080609ｨｰ06090401 070806020209090609 09 0.44 0106 2.6 09ｨｬ/09ｨｮｨｰ ﾂ02080204 1 09ｨｮｨｰ 0706090502 070802ｨｺ08ｨ ｨｴ02ｨｪｨｨ07 0906040102ｨｦ09ｨｰ09ｨｨ07 01ｨ 090502ｨｪｨｨ07 (08ｨｨ09. 2). 030605ｨｮ ﾂ02ｨｪｨｪ0402 01ｨ ｨｪｨｪ0402 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ06ｨｰ 06 09040906ｨｺ06ｨｦ ﾂｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｨ 07080602020909ｨ 080609ｨｰｨ 08ｨ 09ｨｰ070302ｨｪｨｨ02ｨｬ ｨｺ 00ｨｨ070208ｨ ｨ 08ｨｨｨｨ. 080502ｨｰｨｺｨｨ ｨｺ06ｨｪｨｰ080605ｨｹｨｪ0401 08ｨ 09ｨｰ02ｨｪｨｨｨｦ 09 ｨｺ06ｨｪ0202 0406ｨｪ04 08ｨ 09ｨｰ070302ｨｪｨｨ07 (5 ｨｬｨｬ 06ｨｰ ｨｺ06ｨｪ ﾂｨｨｨｺｨ ｨｺ0608ｨｪ07) ｨｨｨｬ0205ｨｨ ｨｺ08ｨｮ07ｨｪｨｮ06 0202ｨｪｨｰ08ｨ 05ｨｹｨｪｨｮ06 09ｨ ｨｺｨｮ0605ｨｹ, 07010806 01ｨｨ09ｨｺ0609ｨｨ01ｨｪ06ｨｦ 000608ｨｬ04 ｨｨ ｨｰ06ｨｪｨｺｨｨｨｦ 0708ｨｨ09ｨｰ02ｨｪｨｪ04ｨｦ 090506ｨｦ 02ｨｨｨｰ060705ｨ 04ｨｬ04 (08ｨｨ09. 3, ｨ, ｨ ). 0601 0705ｨ 04ｨｬｨ 0502ｨｬｨｬｨ ｨ 0405ｨ ｨｮｨｬ020802ｨｪｨｪ06 090605ｨｪｨｨ09ｨｰ06ｨｦ ｨｨ 070506ｨｰｨｪ06 0708ｨｨ050200ｨ 05ｨ ｨｺ ｨｺ0502ｨｰ06 ﾂ- 04ｨｨ09. 2. 0005ｨｨ07ｨｪｨｨ02 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 09ｨｺ06080609ｨｰｨｹ 080609ｨｰｨ ｨｺ0608ｨｪ02ｨｦ 070806080609ｨｰｨｺ0609 ｨｺｨｮｨｺｨｮ08ｨｮ0404. 1 ｪ ｨｺ06ｨｪｨｰ080605ｨｹ, 2 ｪ 01ｨ 090502ｨｪｨｨ02 4 ｨｬ03ｨ 09 ｨｰ02 ﾂ02ｨｪｨｨ02 5 ﾂ. 07ｨｺｨ 04ｨ ｨｪ04 09080201ｨｪ0707 09ｨｺ06080609ｨｰｨｹ (ｨｬ02ｨｰ0601 ｨｪｨ ｨｨｨｬ02ｨｪｨｹ03ｨｨ01 ｨｺ09ｨ 0108ｨ ｨｰ0609) ｨｨ 09ｨｺ06080609ｨｰｨｹ 080609ｨｰｨ 09 1-02 09ｨｮｨｰ 0706090502 0708ｨｨ05060302ｨｪｨｨ07 01ｨ 090502ｨｪｨｨ07. 05ｨｺ06080609ｨｰｨｹ 080609ｨｰｨ 09ｨｪｨｨ03ｨ 05ｨ 09ｨｹ 0708ｨｨ 0906040102ｨｦ09ｨｰ09ｨｨｨｨ 09 2.6 0106 0.44 09ｨｬ/09ｨｮｨｰ ｨｨ 070605ｨｪ0609ｨｰｨｹ06 09060909ｨｰｨ ｨｪｨ 0905ｨｨ09ｨ 05ｨ 09ｨｹ 0106 ｨｺ06ｨｪｨｰ080605ｨｹｨｪ0401 04ｨｪｨ ﾂ02ｨｪｨｨｨｦ ｨｪｨ 09050201ｨｮ06ｨｴｨｨ02 09ｨｮｨｰｨｺｨｨ 0706090502 070802ｨｺ08ｨ ｨｴ02ｨｪｨｨ07 0906040102ｨｦ09ｨｰ09ｨｨ07.

13416 08. 08. 08ｨ 0108ｨ 01ｨｨｨｬ0609 ｨｨ 0108. 04ｨｨ09. 3. 0705ｨｹｨｰ08ｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨ ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ ｨｺ06ｨｪｨｰ080605ｨｹｨｪ0401 08ｨ 09ｨｰ02ｨｪｨｨｨｦ, 04ｨ 00ｨｨｨｺ09ｨｨ080609ｨ ｨｪｨｪ0401 0708ｨｨ ｨ ｨｰｨｬ0609000208ｨｪ06ｨｬ 01ｨ 090502ｨｪｨｨｨｨ. 08010806 (07) 01ｨｨ09ｨｺ0609ｨｨ01ｨｪ06ｨｦ 000608ｨｬ04 (ｨ ) 08ｨ 09070605060302ｨｪ06 09 ｨｰ06ｨｪｨｺ06ｨｬ 0708ｨｨ09ｨｰ02ｨｪｨｪ06ｨｬ 09050602 02ｨｨｨｰ060705ｨ 04ｨｬ04 (ｨ ). 01ｨｨｨ 05060705ｨ 04ｨｬｨ 09080201ｨｪ02ｨｦ 050502ｨｺｨｰ0806ｨｪｨｪ06ｨｦ 070506ｨｰｨｪ0609ｨｰｨｨ 090601020803ｨｨｨｰ ｨｬｨｪ060006 ﾂｨｨ090502ｨｪｨｪ0402 070605ｨｨ0906ｨｬ04 (09, 00). 060502ｨｬ02ｨｪｨｰ04 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0902ｨｰｨｨ (0509) ｪ 02ｨｨ09ｨｰ0208ｨｪ04 ｨｨ 01ｨｨｨｺｨｰｨｨ060906ｨｬ04 ｨ 0707ｨ 08ｨ ｨｰｨ 010605ｨｹ0103ｨｨ ｪ 08ｨ 0907080201020502ｨｪ04 08ｨ 09ｨｪ06ｨｬ0208ｨｪ06 0706 06ｨ ｨｲ02ｨｬｨｮ 02ｨｨｨｰ0604060507 (09, 00). 09ｨｺ ｪ 09ｨ ｨｺｨｮ0605ｨｹ, ｨｺ00 ｪ 01ｨｨｨｺｨｰｨｨ060906ｨｬ04 ｨｺ06ｨｬ070502ｨｺ09ｨ 010605ｨｹ0103ｨｨ, ｨｺ09 ｪ ｨｺ0502ｨｰ06 ﾂｨｪｨ 07 09ｨｰ02ｨｪｨｺｨ, ｨｬ ｪ ｨｬｨｨｨｰ060106ｨｪ0108ｨｨ07. ｨｪ06ｨｦ 09ｨｰ02ｨｪｨｺ02 (08ｨｨ09. 3, ｨ, 09). 01ｨｨｨ 05060705ｨ 04ｨｬｨ ｨｨｨｬ0205ｨ 09080201ｨｪ0606 050502ｨｺｨｰ0806ｨｪｨｪｨｮ06 070506ｨｰｨｪ0609ｨｰｨｹ ｨｨ 090601020803ｨ 05ｨ 04ｨ 09ｨｨｨｰｨｺｨｨ 070605ｨｨ0906ｨｬ (08ｨｨ09. 3, 09, 00). 02ｨｺ08ｨｮ00050402 0608ｨｰ060106ｨｺ09ｨ 05ｨｹｨｪ0402 ｨｬｨｨｨｰ060106ｨｪ0108ｨｨｨｨ, ｨｺ060806ｨｰｨｺｨｨ02 02ｨｨ09ｨｰ0208ｨｪ04 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0902ｨｰｨｨ (0605) ｨｨ 01ｨｨｨｺｨｰｨｨ060906ｨｬ04 ｨ 0405ｨｨ 01ｨｨ0000ｨｮ04ｨｪ06 ｨｨ 08ｨ 09ｨｪ06ｨｬ0208ｨｪ06 08ｨ 0907080201020502ｨｪ04 0706 06ｨ ｨｲ02ｨｬｨｮ 02ｨｨｨｰ0604060507 (08ｨｨ09. 3, ｨ ｪ00). ﾁｨｨ090506 ﾂ0205ｨｪ06 ﾂｨｪ0401 090204ｨｨｨｺｨｮ05 ｨｮ 02ｨｨ09ｨｰ0208ｨｪ 01ｨｨｨｺｨｰｨｨ060906ｨｬ ｨｪ02ｨｬｨｪ060006 ﾂｨｨ090502ｨｪｨｪ06. 0606ｨｪ060705ｨ 09ｨｰ ｨｺ0502ｨｰ06ｨｺ ｨｺ060804 ﾂｨ 09ｨｰ06 0609ｨｬｨｨ0600ｨｨ0502ｨｪ ｨｨ ｨ 01ｨｺ08ｨｮ09ｨｰｨｨ080609ｨ ｨｪ 050502ｨｺｨｰ0806ｨｪｨｪ06-070506ｨｰｨｪ04ｨｬ 0902ｨｴ0209ｨｰ0906ｨｬ. 08ｨｪｨ 050600ｨｨ ﾂｨｪ0602 0902ｨｴ0209ｨｰ0906 09 09ｨｨ0102 01050607ｨｹ0209 ｨｨ 0008ｨ ｨｪｨｮ05 090407090507050609ｨｹ ｨｨ 09 0706050609ｨｰｨｨ 0202ｨｪｨｰ08ｨ 05ｨｹｨｪ06ｨｦ 09ｨ ｨｺｨｮ0605ｨｨ (08ｨｨ09. 3, ｨ, 00). 00 ｨｮ09050609ｨｨ0701 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 2 0003ｨ (30 ｨｬｨｨｨｪ) ｨｪｨ ｨ 050601ｨ 05ｨｨ 06ｨｺ08ｨｮ000502ｨｪｨｨ02 07010208 (08ｨｨ09. 4, ｨ ) ｨｨ ｨｺ05ｨ 09ｨｰ0208ｨｨ04ｨ 02ｨｨ06 050502ｨｬ02ｨｪｨｰ0609 0605 09 06ｨ 08ｨ 040609ｨ ｨｪｨｨ02ｨｬ ｨｮ ﾂｨ 09ｨｰｨｺ0609 050800ｨ 09ｨｰ060705ｨ 04ｨｬ04 (08ｨｨ09. 4, ｨ, 01). 01ｨ ｨ ｨｮ01ｨ ｨｪｨｨ02 0706050609ｨｰｨｨ 0902ｨｰｨｨ ｨ 040506 ｨｪ0204ｨｪｨ ﾂｨｨｨｰ0205ｨｹｨｪ04ｨｬ. 0201ｨｪ06090802ｨｬ02ｨｪｨｪ06 09040709050705ｨｨ09ｨｹ 070806090902ｨｰ0502ｨｪｨｨ02 ｨｬｨ ｨｰ08ｨｨｨｺ09ｨ ｨｬｨｨｨｰ060106ｨｪ0108ｨｨｨｦ (08ｨｨ09. 4, 09) ｨｨ 08ｨ 0907ｨ 01 070605ｨｨ0906ｨｬ ｨｪｨ ｨｬ06ｨｪ060906ｨｬ04 (08ｨｨ09. 4, 00, 01). 07090205ｨｨ ﾂ02ｨｪｨｨ02 050502ｨｺｨｰ0806ｨｪｨｪ06ｨｦ 070506ｨｰｨｪ0609ｨｰｨｨ 02ｨｨｨｰ0604060507 090607080609060301ｨ 050609ｨｹ 080609ｨｰ06ｨｬ 09 02ｨｨｨｰ060705ｨ 04ｨｬ02 ﾂｨｨ0905ｨ ｨｬ0205ｨｺｨｨ01 07080609ｨ ｨｺｨｮ060502ｨｦ (08ｨｨ09. 4, ｨ ). 01ｨ ｨｰ08ｨ ｨｪ09-09ｨｰ060806ｨｪ02 01ｨｨｨｺｨｰｨｨ060906ｨｬ ｨｪｨ ｨｺｨ 0705ｨｨ09ｨ 05ｨｨ09ｨｹ 090204ｨｨｨｺｨｮ0504 09 050502ｨｺｨｰ0806ｨｪｨｪ06-070506ｨｰｨｪ04ｨｬ 090601020803ｨｨｨｬ04ｨｬ (08ｨｨ09. 4, 00). 040201ｨｮｨｺ02ｨｨ07 050502ｨｺｨｰ0806ｨｪｨｪ06ｨｦ 070506ｨｰｨｪ0609ｨｰｨｨ ｨｬ02ｨｬｨ 08ｨ ｨｪ04 ｨｰ06ｨｪ060705ｨ 09ｨｰｨ ｨｺ0608080205ｨｨ080609ｨ 05ｨ 09 ｨｮｨｬ02ｨｪｨｹ0302ｨｪｨｨ02ｨｬ 09 09ｨ ｨｺｨｮ06050701 ｨｺ0605ｨｨ ﾂ0209ｨｰ09ｨ 0609ｨｬｨｨ0600ｨｨ05ｨｹｨｪ0401 09ｨｺ0506 ﾂ02ｨｪｨｨｨｦ (08ｨｨ09. 4, ｨ, 09, 01). 0308ｨｨ 01ｨ 090502ｨｪｨｨｨｨ 4 0003ｨ 01ｨ 08ｨ ｨｺｨｰ0208 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｪ0401 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨｨｦ 09 02ｨｨｨｰ0604060502, 01ｨｨｨｺｨｰｨｨ060906ｨｬｨ 01 ｨｨ ｨｬｨｨｨｰ060106ｨｪ0108ｨｨ0701 (08ｨｨ09. 5, ｨ ｪ09) 09 02020506ｨｬ ｨ 0405 ｨ ｨｪｨ 050600ｨｨ ﾂ02ｨｪ ｨ 05ｨｹｨｰ0208ｨ 02ｨｨ07ｨｬ, 090407090502ｨｪｨｪ04ｨｬ 0708ｨｨ 01ｨ 090502ｨｪｨｨｨｨ 2 0003ｨ (08ｨｨ09. 4, ｨ ｪ01). 0701090602ｨｪｨｨ02 090205ｨｨ ﾂｨｨｨｪ04 01ｨ 090502ｨｪｨｨ07 ｨｨｨｪ01ｨｮ02ｨｨ080609ｨ 0506 090604ｨｪｨｨｨｺｨｪ060902ｨｪｨｨ02 ｨ 05ｨｹｨｰ0208ｨ 02ｨｨｨｦ 09 ｨｰ06ｨｪ060705ｨ 09ｨｰ02 (08ｨｨ09. 5, 09, 09ｨｰ080205ｨｺｨｨ). 01ｨ 050502ｨｺｨｰ0806ｨｪ060008ｨ ｨｬｨｬｨ 01 06ｨｪｨｨ 09040709050705ｨｨ09ｨｹ ｨｺｨ ｨｺ 06ｨ 05ｨ 09ｨｰｨｨ 070208000608ｨ 02ｨｨｨｦ, ﾂｨｰ06 ｨｬ06000506 ｨｮｨｺｨ 040409ｨ ｨｰｨｹ ｨｪｨ ｨｪｨ 08ｨｮ0302ｨｪｨｨ02 0202050609ｨｰｨｪ0609ｨｰｨｨ ｨｬ02ｨｬｨ 08ｨ ｨｪ04 0202ｨｪｨｰ08ｨ 05ｨｹｨｪ06ｨｦ 09ｨ ｨｺｨｮ0605ｨｨ (08ｨｨ09. 5, 09). 040204ｨｺ0602 09ｨｪｨｨ0302ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 0106 ｨ ｨｰｨｬ0609000208ｨｪ060006 0708ｨｨ090601ｨｨ0506 ｨｺ ｨｮ090205ｨｨ ﾂ02ｨｪｨｨ06 0706090802030102ｨｪｨｨｨｦ ｨｰ06ｨｪ060705ｨ 09ｨｰｨ (08ｨｨ09. 6), ｨｺ0605ｨｨ ﾂ0209ｨｰ0906 ｨｺ06ｨｰ06080401 ｨ 040506 070807ｨｬ06 07080607060802ｨｨ06ｨｪｨ 05ｨｹｨｪ06 08ｨ 04ｨｪｨｨ0202 01ｨ 090502ｨｪｨｨｨｦ ｪ ｨｬｨｨｨｪｨｨｨｬｨ 05ｨｹｨｪ06ｨｦ 09 09ｨ 08ｨｨｨ ｨｪｨｰ02 2 1 7 0 0003ｨ (08ｨｨ09. 6, ｨ, 09ｨｰ080205ｨｺｨｨ) ｨｨ ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪ06ｨｦ 0708ｨｨ 4 1 7 0 0003ｨ (08ｨｨ09. 6, ｨ, 09ｨｰ080205ｨｺｨｨ). 0308ｨｨ 05ｨｰ06ｨｬ 01ｨ 08ｨ ｨｺｨｰ0208 0108ｨｮ00ｨｨ01 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨｨｦ (ｨ 00080200ｨ 02ｨｨ07 0902ｨｰｨｨ ｨｨ 01ｨｨｨｺｨｰｨｨ060906ｨｬ 09 090204ｨｨｨｺｨｮ05ｨ ｨｬｨｨ, 070806090902ｨｰ0502ｨｪｨｨ02 ｨｬｨｨｨｰ060106ｨｪ0108ｨｨｨｦ), 090407090502ｨｪｨｪ0401 0708ｨｨ 00ｨｨｨｺ09ｨ 02ｨｨｨｨ 06ｨ 08ｨ 04020609 09 ｨｮ09050609ｨｨ0701 070609040302ｨｪｨｪ060006 01ｨ 090502ｨｪｨｨ07, 09ｨｮｨｴ0209ｨｰ0902ｨｪｨｪ06 ｨｪ02 ｨｨ04ｨｬ02ｨｪ07050907 (08ｨｨ09. 6, 09, 00). 01ｨｨ 09 0601ｨｪ06ｨｬ ｨｨ04 ｨｨ04ｨｮ ﾂｨ 02ｨｬ0401 09ｨ 08ｨｨｨ ｨｪｨｰ0609 ｨｪｨ ｨｬｨｨ ｨｪ02 090407090507050609ｨｹ ｨｪｨ 08ｨｮ0302ｨｪｨｨ07 0202050609ｨｰｨｪ0609ｨｰｨｨ 0705ｨ 04ｨｬｨ 0502ｨｬｨｬ04, ﾂｨｰ06 070601ｨｰ0902080301ｨ 050609ｨｹ 06ｨｺ08ｨ 09ｨｺ06ｨｦ ｨｨｨｪ01ｨｨｨｺｨ ｨｰ060806ｨｬ 0202050609ｨｰｨｪ0609ｨｰｨｨ 07060008ｨ ｨｪｨｨ ﾂｨｪ06ｨｦ ｨｬ02ｨｬｨ 08ｨ ｨｪ04 0609ｨ ｨｪ0906ｨｬ 000605ｨｮｨ 04ｨｬ (01ｨ ｨｪｨｪ0402 ｨｪ02 0708ｨｨ09020102ｨｪ04). 02ｨ 09ｨｮ030102ｨｪｨｨ02 05060005ｨ 09ｨｪ06 070605ｨｮ ﾂ02ｨｪｨｪ04ｨｬ 01ｨ ｨｪｨｪ04ｨｬ, 070609040302ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 09020102ｨｰ ｨｺ ｨｰ0608ｨｬ060302ｨｪｨｨ06 080609ｨｰｨ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ (08ｨｨ09. 1, 2). 00 05ｨｨｨｰ0208ｨ ｨｰｨｮ0802 ｨｨｨｪ00ｨｨｨ ｨｨ080609ｨ ｨｪｨｨ02 080609ｨｰｨ 06ｨｰｨｬ02 ﾂｨ 02ｨｰ0907 ｨｺｨ ｨｺ 07020809ｨ 07 0802ｨ ｨｺ02ｨｨ07 ｨｪｨ 070609040302ｨｪｨｨ02 0609ｨｬ06ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 (Bartels, Sunkar, 2005; Munns, 2011), 0708ｨｨ ﾂ02ｨｬ ﾂｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｹ 080609ｨｰｨ 0706ｨ 0200ｨ ｨｺ 0609ｨｬ06ｨｰｨｨ ﾂ0209ｨｺ06ｨｬｨｮ 09ｨｰ08020909ｨｮ 06ｨｺｨ 04ｨ 05ｨ 09ｨｹ 09040302, ﾂ02ｨｬ ｨｮ ｨｺ0608ｨｪ07 (Bartels, Sunkar, 2005). 00 ｨｪｨ 0302ｨｬ

0005ｨｨ07ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｪｨｮ06 060800ｨ ｨｪｨｨ04ｨ 02ｨｨ06 ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ ｨｺｨｮｨｺｨｮ08ｨｮ0404 417 04ｨｨ09. 4. 0705ｨｹｨｰ08ｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨ ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ 0406ｨｪ04 08ｨ 09ｨｰ070302ｨｪｨｨ07, 04ｨ 00ｨｨｨｺ09ｨｨ080609ｨ ｨｪｨｪ0401 0708ｨｨ 09ｨｪ0203ｨｪ02ｨｬ 01ｨ 090502ｨｪｨｨｨｨ 2 0003ｨ (30 ｨｬｨｨｨｪ). 02ｨｺ08ｨｮ000502ｨｪｨｨ02 07010208 (ｨ ) 090607080609060301ｨ 050609ｨｹ ｨ 00080200ｨ 02ｨｨ02ｨｦ 02ｨｨ09ｨｰ0208ｨｪ 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0902ｨｰｨｨ (ｨ ), 070806090902ｨｰ0502ｨｪｨｨ02ｨｬ ｨｬｨ ｨｰ08ｨｨｨｺ09ｨ ｨｬｨｨｨｰ060106ｨｪ0108ｨｨｨｦ (09), 08ｨ 0907ｨ 0106ｨｬ 070605ｨｨ0906ｨｬ ｨｨ ｨｪｨ ｨｺ06070502ｨｪｨｨ02ｨｬ ｨｰ08ｨ ｨｪ09070608ｨｰｨｪ0401 090204ｨｨｨｺｨｮ05 (00, 01). 02ｨ 0604ｨｪｨ ﾂ02ｨｪｨｨ07 ｨｰ02 0302, ﾂｨｰ06 ｨｨ ｨｪｨ 08ｨｨ09. 3. 0905ｨｮ ﾂｨ 02 ｨｪｨ ｨ 050601ｨ 050609ｨｹ 07080607060802ｨｨ06ｨｪｨ 05ｨｹｨｪ0602 09ｨｪｨｨ0302ｨｪｨｨ02 09ｨｺ06080609ｨｰｨｨ 080609ｨｰ06090401 070806020209090609 ｨｺ0608ｨｪ07 ｨｨ 0706ｨ 0200ｨ (08ｨｨ09. 1, ｨ, ｨ ), 0708ｨｨ ﾂ02ｨｬ 0708ｨｨ 01ｨ 090502ｨｪｨｨ0701 ｨｪｨ 070608070106ｨｺ ｨｪｨｨ0302 090205ｨｨ ﾂｨｨｨｪ04, 0708ｨｨ ｨｺ06ｨｰ060806ｨｦ ｨｪｨ ｨ 050601ｨ 02ｨｰ0907 070802ｨｺ08ｨ ｨｴ02ｨｪｨｨ02 01020502ｨｪｨｨ07 ｨｺ0502ｨｰ06ｨｺ ｨｮ 05ｨｮｨｺｨ 08ｨｨ06ｨｰ (Iwahashi et al., 2005). 06ｨｰｨｨ ｨｪｨ ｨ 05060102ｨｪｨｨ07 070604090605ｨｨ05ｨｨ 04ｨ ｨｺ0506 ﾂｨｨｨｰｨｹ, ﾂｨｰ06 0708ｨｨ ﾂｨｨｨｪ06ｨｦ ｨｰ0608ｨｬ060302ｨｪｨｨ07 080609ｨｰｨ 09 ｨｪｨ 03ｨｨ01 060704ｨｰｨ 01 0709050702ｨｰ0907 ｨｨｨｪ00ｨｨｨ ｨｨ080609ｨ ｨｪｨｨ02 080609ｨｰｨ 08ｨ 09ｨｰ070302ｨｪｨｨ02ｨｬ. 08ｨｪｨ 05ｨｨ04 0406ｨｪ04 08ｨ 09ｨｰ070302ｨｪｨｨ07 ｨｺ0608ｨｪ07 0102ｨｦ09ｨｰ09ｨｨｨｰ0205ｨｹｨｪ06 09040709ｨｨ05 09ｨｮｨｴ0209ｨｰ0902ｨｪｨｪ0402 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 ｨｺ0502ｨｰ06 ﾂｨｪ06ｨｦ 060800ｨ ｨｪｨｨ04ｨ 02ｨｨｨｨ 060704ｨｰｨｪ0401 08ｨ 09ｨｰ02ｨｪｨｨｨｦ 0708ｨｨ 00ｨｨ070208ｨ ｨ 08ｨｨｨｨ (08ｨｨ09. 4, 5, 6). 06ｨｰｨｨ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 090408ｨ 03ｨ 05ｨｨ09ｨｹ 09 ｨｮｨｬ02ｨｪｨｹ0302ｨｪｨｨｨｨ 09 ｨｺ0502ｨｰｨｺｨ 01 090601020803ｨ ｨｪｨｨ07 070605ｨｨ0906ｨｬ 09 02ｨｨｨｰ0604060502, ｨｺ05ｨ 09ｨｰ0208ｨｨ04ｨ 02ｨｨｨｨ 050502ｨｬ02ｨｪｨｰ0609 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0902ｨｰｨｨ ｨｨ 01ｨｨｨｺｨｰｨｨ060906ｨｬ ｨ 0707ｨ 08ｨ ｨｰｨ 010605ｨｹ0103ｨｨ (08ｨｨ09. 4, ｨ, 00; 5, ｨ ｪ09). 01ｨ 070208ｨｨ000208ｨｨｨｨ 01ｨｨｨｺｨｰｨｨ060906ｨｬ 06ｨｰｨｬ02 ﾂｨ 050609ｨｹ ｨｪｨ ｨｺ06070502ｨｪｨｨ02 ｨｰ08ｨ ｨｪ09070608ｨｰｨｪ0401 07ｨｮ040408ｨｹｨｺ0609, 090601020803ｨ ｨｴｨｨ01 050502ｨｺｨｰ0806ｨｪｨｪ06-070506ｨｰｨｪ0602 0902ｨｴ0209ｨｰ0906 (08ｨｨ09. 5, 09). 05060906ｨｺｨｮ07ｨｪ0609ｨｰｨｹ 05ｨｰｨｨ01 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｪ0401 ｨ 05ｨｹｨｰ0208ｨ 02ｨｨｨｦ 090609ｨｬ0209ｨｰｨｪ06 09 ｨｬ06080006050600ｨｨ ﾂ0209ｨｺｨｨｨｬｨｨ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07ｨｬｨｨ 0106ｨｪ0108ｨｨ06ｨｬｨ (08ｨｨ09. 4, 09; 5, ｨ ) 070802030102 0909020006 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ02ｨｰ 06 ｨｰ0608ｨｬ060302ｨｪｨｨｨｨ ｨｬ02ｨｰｨ ｨ 0605ｨｨ ﾂ0209ｨｺｨｨ01 070806020209090609, 09ｨｺ0506 ﾂｨ 07 ｨ ｨｨ0609ｨｨｨｪｨｰ0204 ｨ 0205ｨｺ0609 ｨｨ ｨｰ08ｨ 00ｨｨｨｺ 09 ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ06ｨｦ 09ｨｨ09ｨｰ02ｨｬ02, ﾂｨｰ06 010608060306 09060005ｨ 09ｨｮ02ｨｰ0907 09 ｨｨｨｪ00ｨｨｨ ｨｨ080609ｨ ｨｪｨｨ02ｨｬ 080609ｨｰ06090401 070806020209090609 (08ｨｨ09. 1, 2). 08ｨ ｨｺｨｰ, ﾂｨｰ06 06ｨ ｨｪｨ 08ｨｮ0302ｨｪｨｪ0402 ｨ 05ｨｹｨｰ0208ｨ 02ｨｨｨｨ (08ｨｨ09. 4, 5) 09060108ｨ ｨｪ0706ｨｰ0907 0708ｨｨ 0102ｨｺ06ｨｬ0708020909ｨｨｨｨ (08ｨｨ09. 6), 0909ｨｨ0102ｨｰ0205ｨｹ09ｨｰ09ｨｮ02ｨｰ 06 070807ｨｬ06ｨｬ 0905ｨｨ07ｨｪｨｨｨｨ 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ ｨｮ05ｨｹｨｰ08ｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｮ 08ｨ 09ｨｰｨｨｨｰ0205ｨｹｨｪ06ｨｦ ｨｺ0502ｨｰｨｺｨｨ; 0102ｨｺ06ｨｬ0708020909ｨｨ07 ｨｮ090205ｨｨ ﾂｨｨ09ｨ 05ｨ ﾂｨ 09ｨｰ06ｨｰｨｮ ｨｪｨ 08ｨｮ0302ｨｪｨｨｨｦ ｨｬ02ｨｬｨ 08ｨ ｨｪ04 ｨｰ06ｨｪ060705ｨ 09ｨｰｨ, ｨｪ06 ｨｪ02 0905ｨｨ0705ｨ ｨｪｨ 06ｨ ｨｴｨｨｨｦ 01ｨ 08ｨ ｨｺｨｰ0208 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨｨｦ. 01ｨ ｨｨｨ 06050202 ｨｨｨｪｨｰ02080209ｨｪ04ｨｬｨｨ 09 ｨｰ06 ﾂｨｺｨｨ 040802ｨｪｨｨ07 0905ｨｨ07ｨｪｨｨ07 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 03ｨｨ090402 060800ｨ ｨｪｨｨ04ｨｬ04 0708020109ｨｰｨ 09050706ｨｰ0907 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 0906 09ｨｪｨｮｨｰ0802ｨｪｨｪ02ｨｦ 09ｨｨ09ｨｰ02ｨｬ02 02ｨｨｨｰ060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺｨｨ01 ｨｬ02ｨｬｨ 08ｨ ｨｪ. 05060005ｨ 09ｨｪ06 ｨｬｨｪ02ｨｪｨｨ06 01ｨ ｨｬｨ 050207 (01ｨ ｨｬｨ 0502ｨｦ, 2004), 05ｨｪ0106ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪｨ 07 0902ｨｰｨｹ, ｨ ｨｮ01ｨｮ ﾂｨｨ 04ｨ ｨｬｨｺｨｪｨｮｨｰ06ｨｦ 09ｨｨ09ｨｰ02ｨｬ06ｨｦ, 01060503ｨｪｨ 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ040609ｨ ｨｰｨｹ0907 ｨｬｨ ｨｺ09ｨｨｨｬｨ 05ｨｹｨｪ06ｨｦ ﾂｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｹ06 ｨｺ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ06 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07, ﾂｨｰ06 ｨｨ 09040709050702ｨｰ0907 09 ｨｪｨ 03ｨｨ01 060704ｨｰｨ 01. 030802030102 0909020006 ﾂｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｹ 090408ｨ 03ｨ 05ｨ 09ｨｹ 09 ｨｺ05ｨ 09ｨｰ0208ｨｨ04ｨ 02ｨｨｨｨ 02ｨｨ09ｨｰ0208ｨｪ 0605. 0604ｨｪｨ ﾂｨ 05ｨｹｨｪ06 08ｨ 09ｨｪ06ｨｬ0208ｨｪ06 08ｨ 0907080201020502ｨｪｨｪｨ 07 0706 06ｨ ｨｲ02ｨｬｨｮ ｨｺ0502ｨｰｨｺｨｨ 0902ｨｰｨｹ 000608ｨｬｨｨ080609ｨ 05ｨ ｨ 00080200ｨ ｨｰ04, ﾂｨ 09ｨｰ06 09 ｨ 06050202 ｨｨ05ｨｨ ｨｬ02ｨｪ0202 01ｨ 06ｨｰｨｨ ﾂｨｪ06 08ｨ 09070605060302ｨｪｨｪ04ｨｬｨｨ 02ｨｨ09ｨｰ0208ｨｪｨ ｨｬｨｨ (08ｨｨ09. 4, ｨ, 01; 5, ｨ ). 02ｨ 04 ﾂｨｪ06 02ｨｨ09ｨｰ0208ｨｪ04 0802ｨｰｨｨｨｺｨｮ05ｨｮｨｬｨ ｨｺ060506ｨｺｨ 05ｨｨ040609ｨ ｨｪ04 09 050502ｨｬ02ｨｪｨｰｨ ｨｬｨｨ ｨ ｨｺｨｰｨｨｨｪ0609060006 02ｨｨｨｰ0609ｨｺ020502ｨｰｨ ｨｨ 09 000608ｨｬ02 0002ｨｺ09ｨ 0006ｨｪｨ 05ｨｹｨｪ06ｨｦ 0902ｨｰｨｨ 08ｨ 09ｨｪ06ｨｬ0208ｨｪ06 08ｨ 0907080201020502ｨｪ04 09 ｨｺ0502ｨｰ06 ﾂｨｪ06ｨｬ 06ｨ ｨｲ02ｨｬ02 (Snapp et al., 2003; Sparkes, 2011). 0308ｨｨ ｨｨ09090502010609ｨ ｨｪｨｨｨｨ ｨｮ05ｨｹｨｰ08ｨ ｨｰ06ｨｪｨｺｨｨ01 090802040609 ｨｰｨ ｨｺｨ 07 0902ｨｰｨｹ 09040709050702ｨｰ0907 09 09ｨｨ0102 01ｨｨ0000ｨｮ04ｨｪ06 08ｨ 09070605060302ｨｪｨｪ0401 09 06ｨ ｨｲ02ｨｬ02 ｨｺ0502ｨｰｨｺｨｨ ｨｮ070506ｨｴ02ｨｪｨｪ0401 02ｨｨ09ｨｰ0208ｨｪ (08ｨｨ09. 3, 09, 00). 0306ｨｺｨ 04ｨ ｨｪ06, ﾂｨｰ06 ｨｮ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨ ｨｺｨｰ06ｨｬｨｨ0604ｨｨｨｪ0609ｨ 07 09ｨｨ09ｨｰ02ｨｬｨ ｨｨ0008ｨ 02ｨｰ 0609ｨｪ0609ｨｪｨｮ06 080605ｨｹ 09 060800ｨ ｨｪｨｨ04ｨ 02ｨｨｨｨ ｨｨ 01ｨｨｨｪｨ ｨｬｨｨｨｺ02 050502ｨｬ02ｨｪｨｰ0609 05ｨｪ0106ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ06ｨｦ 09ｨｨ09ｨｰ02ｨｬ04 ｨｨ ｨｰ08ｨ 00ｨｨｨｺ02 0202 070806ｨｨ04090601ｨｪ0401 ｪ ｨｰ08ｨ ｨｪ09070608ｨｰｨｪ0401 090204ｨｨｨｺｨｮ05 (Sparkes, 2011). 08ｨ ｨｺｨｰ ｨｺ05ｨ 09ｨｰ0208ｨｨ04ｨ 02ｨｨｨｨ 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0902ｨｰｨｨ (08ｨｨ09. 4, ｨ ; 5, ｨ ), 01ｨｨｨｺｨｰｨｨ060906ｨｬ ｨ 0707ｨ 08ｨ ｨｰｨ 010605ｨｹ0103ｨｨ (08ｨｨ09. 4, 00) ｨｨ ｨｰ08ｨ ｨｪ09070608ｨｰｨｪ0401 090204ｨｨｨｺｨｮ05 ｨｬ060302ｨｰ ｨｮｨｺｨ 040409ｨ ｨｰｨｹ ｨｪｨ ﾂｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｹ ｨ ｨｺｨｰｨｨｨｪ06-ｨｬｨｨ0604ｨｨｨｪ0609060006 ｨｺ06ｨｬ070502ｨｺ09ｨ ｨｺ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07ｨｬ 01ｨ 090502ｨｪｨｨ07. 06ｨ ｨｺｨ 07 ﾂｨｮ0909ｨｰ09ｨｨｨｰ0205ｨｹｨｪ0609ｨｰｨｹ 0102ｨｦ09ｨｰ09ｨｨｨｰ0205ｨｹｨｪ06 0706ｨｺｨ 04ｨ ｨｪｨ ｨｪｨ ｨｺ0502ｨｰｨｺｨ 01 ｨｺｨｮ05ｨｹｨｰｨｮ0804, ｨｪ06 0708ｨｨ 01ｨ 090502ｨｪｨｨ0701, ｨｪｨ 070608070106ｨｺ 070802090403ｨ 06ｨｴｨｨ01 ｨｰ02, ﾂｨｰ06 ｨｨ09070605ｨｹ040609ｨ ｨｪ04 ｨｪｨ ｨｬｨｨ (Kawarai et al., 2006). 0805ｨ 09ｨｰ0208ｨｨ04ｨ 02ｨｨ07 050502ｨｬ02ｨｪｨｰ0609 0902ｨｰｨｨ 09 06ｨ 08ｨ 040609ｨ ｨｪｨｨ02ｨｬ ｨｮ ﾂｨ 09ｨｰｨｺ0609 050800ｨ 09ｨｰ060705ｨ 04ｨｬ04 ｨｰｨｨ07ｨｨ ﾂｨｪｨ 010507 0902ｨｺ0802ｨｰｨｨ08ｨｮ06ｨｴｨｨ01 ｨｺ0502ｨｰ06ｨｺ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨｨ 03ｨｨ0906ｨｰｨｪ0401 (Snapp et al., 2003). 08ｨ ｨｺ 0708ｨ 09ｨｨ0506, 000608ｨｬｨｨ080609ｨ ｨｪｨｨ02 ｨｺ05ｨ 09ｨｰ02080609 0605 09 ｨｰｨ ｨｺｨｨ01 ｨｺ0502ｨｰｨｺｨ 01 ｨｪｨ 010601ｨｨｨｰ0907 070601 ｨｺ06ｨｪｨｰ08060502ｨｬ ｨｺ0502ｨｰｨｺｨｨ, ｨ 09ｨ ｨｬｨｨ 06ｨｪｨｨ 0708020109ｨｰｨ 090502ｨｪ04 ｨｮ070506ｨｴ02ｨｪｨｪ04ｨｬｨｨ 02ｨｨ09ｨｰ0208ｨｪｨ ｨｬｨｨ, ｨｮ07ｨ ｨｺ0609ｨ ｨｪｨｪ04ｨｬｨｨ 09 070506ｨｰｨｪ0402 07ｨ ｨｺ02ｨｰ04. 06ｨｪ06ｨｦ 01ｨ 08ｨ ｨｺｨｰ0208 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨｨｦ 090407090502ｨｪ 09 ｨｪｨ 03ｨｨ01 060704ｨｰｨ 01. 060502ｨｬ02ｨｪｨｰ04 ｨｨ04ｨｬ02ｨｪ0705ｨｨ ｨｬ0209ｨｰ06 09090602ｨｦ 0506ｨｺｨ 05ｨｨ04ｨ 02ｨｨｨｨ ｨｨ 06ｨ 08ｨ 0406-

418 08. 08. 08ｨ 0108ｨ 01ｨｨｨｬ0609 ｨｨ 0108. 04ｨｨ09. 5. 0005ｨｨ07ｨｪｨｨ02 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 4 0003ｨ (30 ｨｬｨｨｨｪ) ｨｪｨ ｨｮ05ｨｹｨｰ08ｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｮ ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ ｨｺｨｮｨｺｨｮ08ｨｮ0404. 0800080200ｨ 02ｨｨ07 05ｨｪ01060705ｨ 04ｨｬｨ ｨｰｨｨ ﾂ0209ｨｺ06ｨｦ 0902ｨｰｨｨ, 01ｨｨｨｺｨｰｨｨ060906ｨｬ, ｨｰ08ｨ ｨｪ09070608ｨｰｨｪ0401 090204ｨｨｨｺｨｮ05 (ｨ ) ｨｨ 070806090902ｨｰ0502ｨｪｨｨ02 ｨｬｨ ｨｰ08ｨｨｨｺ09ｨ ｨｬｨｨｨｰ060106ｨｪ0108ｨｨｨｦ (ｨ ) 090607080609060301ｨ 06ｨｰ0907 ｨｪｨ 08ｨｮ0302ｨｪｨｨ07ｨｬｨｨ 0202050609ｨｰｨｪ0609ｨｰｨｨ ｨｰ06ｨｪ060705ｨ 09ｨｰｨ (09). 02ｨ 08ｨ 040204 04ｨ 00ｨｨｨｺ09ｨｨ080609ｨ ｨｪ04 09 ｨｺｨ ｨｬ020802 01ｨ 090502ｨｪｨｨ07. 02ｨ 0604ｨｪｨ ﾂ02ｨｪｨｨ07 ｨｰ02 0302, ﾂｨｰ06 ｨｨ ｨｪｨ 08ｨｨ09. 3. 090409ｨ 05ｨｨ ｨ 00080200ｨ ｨｰ04. 00 ｨｪｨ 0302ｨｬ 0905ｨｮ ﾂｨ 02 ｨ 00080200ｨ 02ｨｨ07 050502ｨｬ02ｨｪｨｰ0609 ｨｪ0609ｨｨ05ｨ, 09ｨｺ06080202, 01ｨ 06ｨｰｨｨ ﾂｨｪ04ｨｦ ｨｨ ｨｪ02ｨｮ0708ｨ 09050702ｨｬ04ｨｦ 01ｨ 08ｨ ｨｺｨｰ0208. 00 ｨｪｨ 09ｨｰ0607ｨｴ0202 090802ｨｬ07 080605ｨｹ 0605 ｨｺｨ ｨｺ 060800ｨ ｨｪ02050504, 06ｨｰ0902ｨｰ09ｨｰ0902ｨｪｨｪ06ｨｦ 04ｨ 080701 09ｨ 03ｨｪ02ｨｦ03ｨｨ01 00ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨ01 070806020209090609, 09ｨｺ0506 ﾂｨ 07 09ｨｨｨｪｨｰ0204, 00060501ｨｨｨｪ00 ｨｨ 070609ｨｰｨｰ08ｨ ｨｪ09050702ｨｨ06ｨｪｨｪｨｮ06 ｨｬ0601ｨｨ00ｨｨｨｺｨ 02ｨｨ06 ｨ 0605ｨｹ03ｨｨｨｪ09ｨｰ09ｨ ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ0401 ｨｨ 0902ｨｺ0802ｨｰ0608ｨｪ0401 ｨ 0205ｨｺ0609, 06ｨ ｨｴ02ｨｨ04090209ｨｰｨｪｨ (Samali et al., 2010). 00 ｨｮ09050609ｨｨ0701 07ｨ ｨｰ06050600ｨｨｨｨ 09 ｨｺ0502ｨｰｨｺｨ 01 03ｨｨ0906ｨｰｨｪ0401 ｨｪｨ ｨ 050601ｨ 02ｨｰ0907 0802ｨ ｨｺ02ｨｨ07 09ｨｰ08020909ｨ 0605, ｨｺ06ｨｰ0608ｨ 07 090906ｨｦ09ｨｰ0902ｨｪｨｪｨ ｨｰｨ ｨｺ0302 0708ｨｨ 000608ｨｬｨｨ080609ｨ ｨｪｨｨｨｨ ｨｪ02ｨｦ08060102000208ｨ ｨｰｨｨ09ｨｪ0401, 08ｨ ｨｺ06090401 04ｨ ｨ 06050209ｨ ｨｪｨｨｨｦ ｨｨ 01ｨｨｨ ｨ 02ｨｰｨ. 00 06ｨ ｨｴ02ｨｬ 0905ｨｮ ﾂｨ 02 05ｨｰｨ 0802ｨ ｨｺ02ｨｨ07 04ｨ 07ｨｮ09ｨｺｨ 02ｨｰ0907, ｨｺ060001ｨ 00ｨｨ04ｨｨ06050600ｨｨ ﾂ0209ｨｺｨｨ02 ｨｨ05ｨｨ 09ｨｪ0203ｨｪｨｨ02 ｨｮ09050609ｨｨ07 0905ｨｨ0706ｨｰ ｨｪｨ 0708060202090904 00060501ｨｨｨｪ00ｨ ｨ 0205ｨｺ0609 ｨｨ ｨｪ02040802050402 ｨｨ05ｨｨ ｨｪ020708ｨ 09ｨｨ05ｨｹｨｪ06 0906ｨ 08ｨ ｨｪｨｪ0402 ｨ 0205ｨｺｨｨ ｨｪｨ ｨｺｨ 0705ｨｨ09ｨ 06ｨｰ0907 09 0706050609ｨｰｨｨ 0902ｨｰｨｨ (Fulda et al., 2010). 02ｨｪｨ 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ04ｨｮ02ｨｰ0907 ｨ ｨｺｨｰｨｨ09ｨ 02ｨｨ02ｨｦ ｨｰ080201 09ｨｰ08020909-0802020207ｨｰ06080609 09 0605: PKR-07060106ｨ ｨｪ06ｨｦ ER-ｨｺｨｨｨｪｨ 0404 0605 (PERK), ｨ ｨｺｨｰｨｨ09ｨｨ08ｨｮ06ｨｴ020006 ｨｰ08ｨ ｨｪ09ｨｺ08ｨｨ0702ｨｨ06ｨｪｨｪ060006 00ｨ ｨｺｨｰ0608ｨ 6 (ATF6) ｨｨ ｨｨｨｪ0604ｨｨｨｰ060504ｨ 09ｨｨ09ｨｨｨｬ060006 000208ｨｬ02ｨｪｨｰｨ 1 (Ire1), ｨｺ06ｨｰ06080402 09 09090606 06 ﾂ02080201ｨｹ 0906090502 ﾂ02ｨｪ04 09 0802ｨ ｨｺ02ｨｨ06 ｨｺ0502ｨｰｨｺｨｨ ｨｪｨ 0102ｨｪｨ ｨｰｨｮ08ｨｨ080609ｨ ｨｪｨｪ0402 ｨ 0205ｨｺｨｨ (UPR, unfolded protein reaction). 050609ｨｬ0209ｨｰｨｪ06 ｨｰ08ｨｨ 0902ｨｰ09ｨｨ 06ｨｰ0902ｨｰｨ ｨ 0506ｨｺｨｨ08ｨｮ06ｨｰ ｨｰ08ｨ ｨｪ09050702ｨｨ06, ｨｮ090205ｨｨ ﾂｨｨ09ｨ 06ｨｰ 05ｨｺ090708020909ｨｨ06 03ｨ 07020806ｨｪ0609 ｨｨ ｨｮ09ｨｨ05ｨｨ09ｨ 06ｨｰ 0605-04ｨ 09ｨｨ09ｨｨｨｬ0402 07ｨｮｨｰｨｨ 01020008ｨ 01ｨ 02ｨｨｨｨ 070605ｨｨ070207ｨｰｨｨ010609. 030905ｨｨ ｨｬ06ｨｴｨｪ0609ｨｰｨｹ 0802ｨ ｨｺ02ｨｨｨｨ UPR ｨｪ02010609ｨｰｨ ｨｰ06 ﾂｨｪｨ 010507 0102ｨｰ06ｨｺ09ｨｨｨｺｨ 02ｨｨｨｨ 0902ｨｺ0802ｨｰｨｨ08ｨｮ02ｨｬ0401 07080601ｨｮｨｺｨｰ0609, 04ｨ 07ｨｮ09ｨｺｨ 02ｨｰ0907 0605-ｨｨｨｪ01ｨｮ02ｨｨ08ｨｮ02ｨｬｨ 07 00ｨｨｨ 0205ｨｹ ｨｺ0502ｨｰ06ｨｺ. 020507 08ｨ 09ｨｰ02ｨｪｨｨｨｦ ｨｬ060502ｨｺｨｮ050708ｨｪｨ 07 09ｨｨ00ｨｪｨ ｨｰｨｮ08ｨ 010507 08ｨ 0409ｨｨｨｰｨｨ07 0802ｨ ｨｺ02ｨｨｨｨ UPR 0106 ｨｺ06ｨｪ02ｨ ｨｪ02 08ｨ 0903ｨｨ00080609ｨ ｨｪｨ. 0306ｨｺｨ 04ｨ ｨｪ06, ﾂｨｰ06 ｨ ｨｺｨｰｨｨ09ｨ 02ｨｨ07 bzip17 ｪ ｨ ｨｪｨ 0506-00ｨ ATF6 ｪ 070806ｨｨ09010601ｨｨｨｰ 0708ｨｨ 090605020906ｨｬ 09ｨｰ0802090902 (Liua, Howell, 2010). 00 2012 00. (Varadarajan et al., 2012) 0607ｨｨ09ｨ ｨｪ 07080602020909 ｨ 0409ｨｰ0806ｨｦ ｨｨ 06ｨ 08ｨ ｨｰｨｨｨｬ06ｨｦ 0802060800ｨ ｨｪｨｨ04ｨ 02ｨｨｨｨ 0605, ｨｺ06ｨｰ060804ｨｦ 07080201030209ｨｰ09ｨｮ02ｨｰ 000608ｨｬｨｨ080609ｨ ｨｪｨｨ06 09ｨｰ0802090906090401 0802ｨ ｨｺ02ｨｨｨｦ 09 ｨｺ0502ｨｰｨｺｨ 01 ｨｨ 0708020109ｨｰｨ 09050702ｨｰ 0906ｨ 06ｨｦ ｨｬｨ 0903ｨｰｨ ｨ ｨｪ0602 ｨｨ 09070202ｨｨ00ｨｨ ﾂｨｪ0602 0802ｨｬ06010205ｨｨ080609ｨ ｨｪｨｨ02 09ｨｨ09ｨｰ02ｨｬ04 ｨｬ02ｨｬｨ 08ｨ ｨｪ 0902ｨｰｨｨ (EMR, endoplasmic reticulum membrane remodeling). 06ｨｰ06ｨｰ 07080602020909 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ04ｨｮ02ｨｰ0907 ｨｺ05ｨ 09ｨｰ0208ｨｨ04ｨ 02ｨｨ02ｨｦ 0605 09 ｨｺ08ｨｮ07ｨｪ0402 09ｨｺ06070502ｨｪｨｨ07 ｨ 00080200ｨｨ080609ｨ ｨｪｨｪ0401 02ｨｨ09ｨｰ0208ｨｪ. 0306 09090602ｨｦ ｨｬ06080006050600ｨｨｨｨ 06ｨｪｨｨ ｨｪｨ 0706ｨｬｨｨｨｪｨ 06ｨｰ ｨ 00080200ｨ ｨｰ04 02ｨｨ09ｨｰ0208ｨｪ 0902ｨｰｨｨ, 090407090502ｨｪｨｪ0402 ｨｪｨ ｨｬｨｨ 09 ｨｮ09050609ｨｨ0701 00ｨｨ070208ｨ ｨ 08ｨｨｨｨ (08ｨｨ09. 4, ｨ ; 5, ｨ ). 0306 ｨｬｨｪ02ｨｪｨｨ06 ｨ 09ｨｰ06080609, 05ｨｰ06ｨｰ 06ｨｰ0902ｨｰ 0902ｨｰｨｨ 050906050602ｨｨ06ｨｪｨｪ06 ｨｺ06ｨｪ09020809ｨ ｨｰｨｨ0902ｨｪ, 090607080609060301ｨ 02ｨｰ0907 070802ｨｺ08ｨ ｨｴ02ｨｪｨｨ02ｨｬ ｨｰ08ｨ ｨｪ09050702ｨｨｨｨ ｨｨ 0902ｨｺ080202ｨｨｨｨ, ｨｪ06 09 06ｨｰ05ｨｨ ﾂｨｨ02 06ｨｰ 010608060306 ｨｨ04090209ｨｰｨｪ06ｨｦ 0802ｨ ｨｺ02ｨｨｨｨ 09ｨｰ08020909ｨ 0605 (UPR) 070605ｨｪ0609ｨｰｨｹ06 06ｨ 08ｨ ｨｰｨｨｨｬ (Varadarajan et al., 2012). 01ｨ ｨ 050601ｨ 02ｨｬ0402 ｨｪｨ ｨｬｨｨ ｨｬ06080006050600ｨｨ ﾂ0209ｨｺｨｨ02 ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07 ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ07, 070802ｨｺ08ｨ ｨｴ02ｨｪｨｨ02 020006 080609ｨｰｨ ｨｨ 06ｨ 08ｨ ｨｰｨｨｨｬ0609ｨｰｨｹ 00ｨｮｨｪｨｺ02ｨｨ06ｨｪｨ 05ｨｹｨｪ0401 01ｨ 08ｨ ｨｺｨｰ0208ｨｨ09ｨｰｨｨｨｺ 0708ｨｨ 0102ｨｺ06ｨｬ0708020909ｨｨｨｨ ｨ 06050202 090606ｨｰ0902ｨｰ09ｨｰ09ｨｮ06ｨｰ 05ｨｰ06ｨｦ 06ｨ 08ｨ ｨｰｨｨｨｬ06ｨｦ 0802ｨ ｨｺ02ｨｨｨｨ 0902ｨｰｨｨ (Varadarajan et al., 2012), ｨｪ02030205ｨｨ 00ｨｺ05ｨ 0909ｨｨ ﾂ0209ｨｺ06ｨｦ03 0802ｨ ｨｺ02ｨｨｨｨ 09ｨｰ08020909ｨ 0605 (Fulda et al., 2010). 060608ｨｬ060302ｨｪｨｨ02 ｨｰ08ｨ 00ｨｨｨｺｨ ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ060006 ｨｬｨ ｨｰ0208ｨｨｨ 05ｨ 09 08ｨ 09ｨｰｨｮｨｴｨｮ06 09ｨ ｨｺｨｮ0605ｨｹ ｨｬ060302ｨｰ ｨ 04ｨｰｨｹ 0708ｨｨ ﾂｨｨｨｪ06ｨｦ 070607090502ｨｪｨｨ07 070208000608ｨ 02ｨｨｨｦ 09 ｨｰ06ｨｪ060705ｨ 09ｨｰ02, ﾂｨｰ06 09 09090606 06 ﾂ02080201ｨｹ ｨｬ060302ｨｰ 090209ｨｰｨｨ ｨｺ 070802ｨｺ08ｨ ｨｴ02ｨｪｨｨ06 080609ｨｰｨ ｨｺ0502ｨｰ06ｨｺ 08ｨ 09ｨｰ070302ｨｪｨｨ02ｨｬ. 0102ｨｰ 0906ｨｬｨｪ02ｨｪｨｨｨｦ, ﾂｨｰ06 0202050609ｨｰｨｪ0609ｨｰｨｹ ｨｰ06ｨｪ060705ｨ 09ｨｰｨ, ｨ 090502010609ｨ ｨｰ0205ｨｹｨｪ06, 0907060906ｨ ｨｪ0609ｨｰｨｹ ｨｺ0502ｨｰｨｺｨｨ 0002ｨｪ0208ｨｨ080609ｨ ｨｰｨｹ ｨｰ08ｨ ｨｪ09ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ04ｨｦ 0706ｨｰ02ｨｪ02ｨｨｨ 05 ｨｨ06ｨｪ0609 09060106080601ｨ, ｨｪ02-

130005ｨｨ07ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｪｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨｪｨｮ06 060800ｨ ｨｪｨｨ04ｨ 02ｨｨ06 ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ ｨｺｨｮｨｺｨｮ08ｨｮ0404 419 04ｨｨ09. 6. 0705ｨｹｨｰ08ｨ 09ｨｰ08ｨｮｨｺｨｰｨｮ08ｨ ｨｺ0502ｨｰ06ｨｺ ｨｺ0608ｨｪ02ｨｦ 0708ｨｨ ｨ 0409ｨｰ0806ｨｬ 09ｨ 08060902 09ｨｪ0203ｨｪ020006 01ｨ 090502ｨｪｨｨ07 0106 ｨ ｨｰｨｬ0609000208ｨｪ06ｨｦ 090205ｨｨ ﾂｨｨｨｪ04. ｨ, 09 ｪ 09ｨ 080609 01ｨ 090502ｨｪｨｨ07 09 2 0003ｨ 0106 ｨ ｨｰｨｬ0609000208ｨｪ060006; ｨ, 00 ｪ 09ｨ 080609 01ｨ 090502ｨｪｨｨ07 09 4 0003ｨ 0106 ｨ ｨｰｨｬ0609000208ｨｪ060006. ﾁｨ 09ｨｰ06ｨｰｨ 08ｨ 040804090609 (09ｨｰ080205ｨｺｨｨ) ｨｰ06ｨｪ060705ｨ 09ｨｰｨ ｨｨ 0102000608ｨｬｨ 02ｨｨｨｦ ｨｺ0502ｨｰ06 ﾂｨｪ0401 09ｨｰ02ｨｪ06ｨｺ 070807ｨｬ06 07080607060802ｨｨ06ｨｪｨ 05ｨｹｨｪｨ ｨ ｨｬ0705ｨｨｨｰｨｮ0102 09ｨ 08060302ｨｪｨｪ060006 01ｨ 090502ｨｪｨｨ07. 02ｨ 08ｨ 040204 04ｨ 00ｨｨｨｺ09ｨｨ080609ｨ ｨｪ04 0708ｨｨ ｨ ｨｰｨｬ0609000208ｨｪ06ｨｬ 01ｨ 090502ｨｪｨｨｨｨ. 02ｨ 0604ｨｪｨ ﾂ02ｨｪｨｨ07 ｨｰ02 0302, ﾂｨｰ06 ｨｨ ｨｪｨ 08ｨｨ09. 3. 06ｨ 010601ｨｨｨｬ04ｨｦ 010507 09ｨｰ0608ｨｨ ﾂｨｪ060006 ｨｰ08ｨ ｨｪ09070608ｨｰｨ 0609ｨｬ06ｨｰｨｨｨｺ0609, 0709050706ｨｰ0907 ｨｺ08ｨｨｨｰｨｨ ﾂｨｪ04ｨｬｨｨ 010507 080609ｨｰ06090401 070806020209090609 09 0406ｨｪ02 08ｨ 09ｨｰ070302ｨｪｨｨ07. 06ｨ ｨｺｨｨｨｬ 06ｨ 08ｨ 0406ｨｬ, ｨｮ090205ｨｨ ﾂ02ｨｪｨｨ02 00ｨｨ01080609ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 0106 06ｨ 05ｨ 09ｨｰ02ｨｦ 09020801ｨｪ02ｨｦ 0008ｨ ｨｪｨｨ0204 00ｨｨ04ｨｨ06050600ｨｨ ﾂｨｪ0401 01ｨ 090502ｨｪｨｨｨｦ (2 ｪ4 0003ｨ ) 0708ｨｨ090601ｨｨｨｰ ｨｺ 09ｨｮｨｴ0209ｨｰ0902ｨｪｨｪ04ｨｬ ｨｨ04ｨｬ02ｨｪ02ｨｪｨｨ07ｨｬ 09 05ｨｪ0106ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ06ｨｦ 09ｨｨ09ｨｰ02ｨｬ02, 0708060709050706ｨｴｨｨｨｬ0907 09 ｨ 05ｨｹｨｰ0208ｨ 02ｨｨｨｨ 060800ｨ ｨｪ020505, ｨｮ ﾂｨ 09ｨｰ09ｨｮ06ｨｴｨｨ01 09 ｨｬ02ｨｬｨ 08ｨ ｨｪｨｪ06ｨｬ ｨｰ08ｨ 00ｨｨｨｺ02, ｨｨ 000608ｨｬｨｨ080609ｨ ｨｪｨｨｨｨ 09 ｨｺ0502ｨｰｨｺｨ 01 08ｨ 09ｨｰ02ｨｪｨｨｨｦ 0802ｨ ｨｺ02ｨｨｨｨ 09ｨｰ08020909ｨ 0605. 04ｨ ｨ 06ｨｰｨ 0904070605ｨｪ02ｨｪｨ 0708ｨｨ 00ｨｨｨｪｨ ｨｪ09060906ｨｦ 07060101020803ｨｺ02 04060909ｨｨｨｦ09ｨｺ060006 0006ｨｪ01ｨ 00ｨｮｨｪ01ｨ ｨｬ02ｨｪｨｰｨ 05ｨｹｨｪ0401 ｨｨ090902010609ｨ ｨｪｨｨｨｦ (07080602ｨｺｨｰ 08-04-01258). 0507ｨｨ0906ｨｺ 05ｨｨｨｰ0208ｨ ｨｰｨｮ0804 08ｨｪｨｨ09ｨｨｨｬ0609 08. 00., 05ｨｮ09050609 00. 08., 05ｨｮｨｦｨｺ0609 00. 08. 2012. 0209ｨｪｨ 09ｨｰｨｺｨ ｨｺ ｨｨ09090502010609ｨ ｨｪｨｨ07ｨｬ ｨｬｨ 09090607020802ｨｪ0609ｨ 070601 0905ｨｨ07ｨｪｨｨ02ｨｬ 09ｨｰｨ ｨｰｨｨ ﾂ0209ｨｺ060006 ｨｨ 01ｨｨｨｪｨ ｨｬｨｨ ﾂ0209ｨｺ060006 01ｨ 090502ｨｪｨｨ07 ｨｪ0207060908020109ｨｰ0902ｨｪｨｪ06 09 01ｨ ｨｰ ﾂｨｨｨｺ02 0008ｨ 01ｨｨ02ｨｪｨｰｨｪ060006 080004. 02ｨ ｨｰ ﾂｨｨｨｺｨｨ ｨｨ 09ｨｨ09ｨｰ02ｨｬ04. 7 : 64 ｪ67. 00ｨ ｨｦｨｪｨ 08 04. 1987. 0209ｨｨ0302ｨｪｨｨ07 ｨｮ 08ｨ 09ｨｰ02ｨｪｨｨｨｦ. 00.: 05ｨｪｨ ｨｪｨｨ02. 176 09. 01ｨ ｨｬｨ 0502ｨｦ 07. 00. 2004. 0608ｨ ｨｪ09070608ｨｰｨｪｨ 07 09ｨｨ09ｨｰ02ｨｬｨ 090609ｨｮ01ｨｨ09ｨｰ0401 08ｨ 09ｨｰ02ｨｪｨｨｨｦ. 0503ｨ.: 060401-0906 05.-0302ｨｰ0208ｨ ｨｮ0800. ｨｮｨｪ-ｨｰｨ. 424 c. 05ｨ 05ｨｹ02ｨｬｨ ｨｪ 01. 09., 08ｨｮ ﾂｨｮｨｺ 01. 08., 01ｨｮ080002ｨｪｨｨ010402 08. 01. 1979. 0209ｨｪ060904 00ｨｨ070208ｨ ｨ 08ｨｨ ﾂ0209ｨｺ06ｨｦ 00ｨｨ04ｨｨ06050600ｨｨｨｨ 09.: 0000020108. 318 09. Abe F., Horikoshi K. 1995. Hydrostatic pressure promotes the acidification of vacuoles in Saccharomyces cerevisiae. FEMS Microbiol. Lett. 130 : 307 ｪ312. Ache P., Bauer H., Kollist H., Khaled A. S., Al-Rasheid, Lautner S., Hartung W., Hedrich R. 2010. Stomatal action directly feeds back on leaf turgor: new insights into the regulation of the plant water status from non-invasive pressure probe measurements. Plant J. 62 : 1072 ｪ1082. Bartels D., Sunkar R. 2005. Drought and salt tolerance in plants. Critical Reviews in Plant Sciences. 24 : 23 ｪ58. Dziubinska H. 2003. Ways of signal transmission and physiological role of electrical potentials in plants. Acta Soc. Bot. Pol. 72 : 309 ｪ318. Fernandes P. M., Domitrovic T., Kao C. M., Kurtenbach E. 2004. Genomic expression pattern in Saccharomyces cerevisiae cells in response to high hydrostatic pressure. FEBS Lett. 556 : 153 ｪ160. Franks P. J. 2003. Use of the pressure probe in studies of stomatal function. J. Exp. Bot. 54 : 1495 ｪ1504. Fulda S., Gorman A. M., Hori O., Samali A. 2010. Cellular stress responses: cell survival and cell death. Int. J. Cell Biol. Article ID 214074, p. 23. Galmｨｦs J., Ribas-Carbｨｮ M., Medrano H., Flexas J. 2011. Rubisco activity in Mediterranean species is regulated by the chloroplastic CO 2 concentration under water stress. J. Exp. Bot. 62 : 653 ｪ665. Gross M., Jaenicke R. 1994. Proteins under pressure. The influence of high hydrostatic pressure on structure, function and assembly of proteins and protein complexes. Eur. J. Biochem. 221 : 617 ｪ630. Iwahashi H., Odani M., Ishidou E., Kitagawa E. 2005. Adaptation of Saccharomyces cerevisiae to high hydrostatic pressure causing growth inhibition. Nature. 579 : 2847 ｪ2852. Kawarai T., Arai S., Furukawa S., Ogihara H., Yamasaki M. 2006. High-hydrostatic-pressure treatment impairs actin cables and budding in Saccharomyces cerevisiae. J. Biosci. Bioeng. 101 : 515 ｪ518. Knoblauch M., Peters W. S. 2010. Mｨｹnch, morphology, microfluidics ｪ our structural problem with the phloem. Plant, Cell and Environment. 33 : 1439 ｪ1452. Liua J., Howell S. H. 2010. Endoplasmic reticulum protein quality control and its relationship to environmental stress responses in plants. Plant Cell. 22 : 2930 ｪ2942.

13420 08. 08. 08ｨ 0108ｨ 01ｨｨｨｬ0609 ｨｨ 0108. Molina-Garcia A. D. 2002. The effect of hydrostatic pressure on biological systems. Biotechnol. Genet. Eng. Rev. 19 : 3 ｪ53. Munns R. 2011. Plant adaptations to salt and water stress: differences and Commonalities. Adv. Bot. Res. 57 : 1 ｪ32. Reynolds E. S. 1963. The use of lead citrate at high ph as an electron opaque stain in electron microscopy. Cell Biol. 17 : 208 ｪ212. Rothschild L. J., Mancinelli R. L. 2001. Life in extreme environments. Nature. 409 : 1092 ｪ1101. Samali A., Gerald U. F., Deegan S., Gupta S. 2010. Methods for monitoring endoplasmic reticulum stress and the unfolded protein response. Int. J. Cell Biol. 2010 : 1 ｪ12. Snapp E. L., Hegde R. S., Francolini M., Lombardo F., Colombo S., Pedrazzini E., Borgese N., Lippincott-Schwartz J. 2003. Formation of stacked ER cisternae by low affinity protein interactions. J. Cell Biol. 163 : 257 ｪ269. Sparkes I. 2011. Recent advances in understanding plant myosin function: life in the fast lane. Mol. Plant. 4 : 805 ｪ812. Tander B. 1990. Improved uranyl acetate staining for electron microscopy. J. Electron. Microsc. Techn. 16 : 81 ｪ82. Tanino K. K., Chen H. H., Fuchigami L. H., Weiser C. J. 1992. Abscisic acid increases terrestrial plant cell resistance to hydrostatic pressure. Plant Physiol. 98 : 745 ｪ748. Turgeon M. 2010. The puzzle of phloem pressure. Plant Physiol. 154 : 578 ｪ581. Varadarajan S., Bampton E. T., Smalley J. L., Tanaka K., Caves R. E., Butterworth M., Wei J., Pellecchia M., Mitcheson J., Gant T. W., Dinsdale D., Cohen G. M. 2012. A novel cellular stress response characterised by a rapid reorganisation of membranes of the endoplasmic reticulum. Cell Death and Differentiation. 19 : 1896 ｪ1907. 030609ｨｰｨｮ07ｨｨ05ｨ 28 I 2013 EFFECT OF HYDROSTATIC PRESSURE ON STRUCTURAL ORGANIZATION OF THE MAIZE ROOT CELLS F. A. Abdrakhimov, 1 M. A. Suslov, A. V. Anisimov Institute of Biochemistry and Biophysics, Kazan Scientific Center RAS; 1 e-mail: abdfar@mail.ru We analyzed the effect of hydrostatic pressure at 2 ｪ4 MPa level on the growth and organization of corn seedlings cells. At pressure value of 2 MPa, we observed the clarification of the mitochondrial matrix, clusterization of endoplasmic reticulum elements with formation of ergastorplasm sites and the increasing number of small vacuoles in the cytoplasm. We detected the accumulation of vesicles with electron-dense content on the trans side of dictyosomes, which indicates a change in the endomembranous system traffic. The reduction of electronic density of tonoplast was accompanied by a reduction of osmiophilic inclusions in the vacuoles. Increasing the pressure up to 4 MPa showed the emergence of destructive alterations in the tonoplast. None of the variants showed any violation of plasmalemma integrity, as confirmed by coloring with Evans blue. When we reset the hydrostatic pressure to atmospheric pressure, the frequency of structural distortions of the tonoplast membrane was proportional to the pressure difference: the minimum was at reset from 2 to 0 MPa and the maximum was at reset from 4 to 0 MPa. Thus, it has been concluded, that the endomembraneous system is among the targets of pressure effect on the plant cells. K e y w o r d s: pressure, stress, endoplasmic reticulum, corn seedlings.