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08 29 6 7. 05. 05. 05040ｷ3076 5*, ***, 04. ｨ. 05040ｷ3076 56ｦ0**, *** *ｨ 060309030403 08apple07 0203 0ｹ1010702076 70909 09 0ｹ16 507020ｸ90ｷ30909 0903. 02.04. 056 5apple0ｷ3076 56ｦ0 ｨｺ0204, 117077 020706016 56ｦ0, 0906010906 08apple070608., 33 **02070601076 506010906 6 7070604 6ｦ0apple06036 50ｸ806 04096 5apple06090303 0903. 02.05. 0703070706076 56ｦ0, ｨｴ01070ｷ303apple, 119992 020706016 56ｦ0, 09060109 6 707apple0ｸ8 ***ｨｺ0706060906060107-05036ｦ306010906 03apple0708090ｹ6010906 0ｷ303apple, 6ｧ8 3, Street 3/2, 10 Distr., Hochimin, Vietnam E-mail: vvsuntsov@rambler.ru ｨｨ070603040809026ｦ0 6 5 apple 6ｦ10ｷ3090ｸ9 19.01.29 6 7. 05 030702010402flapple07-6 703090ｹ6010903 09060602 076 56ｦ9fl03 08070602 6 707 06fl0309020309fl, 08apple0701026ｦ309apple040306fl 066ｦ2036ｦｷ3090707 08apple070906030708 09 6 50709 04 090302fl 0ｹ4030ｸ8 Yersinia pestis 0703 08066 5 070304 apple01040209076 707 030901apple07 6ｦ0 Y. pseudotuberculosis O:1b. ｨｨapple09 0ｹ1030703 08apple070ｷ30606 6 509 0707 apple6ｦ9076 56ｦ9fl 066 5fl090ｸ86 56ｦｸ903 06 6 707apple090907036ｦ20ｸ803 6 50603apple6ｦ96 56ｦ903 6 5 6 70703 6ｦ0-0ｷ30603apple6ｦ20706 0207apple030ｸ8 0ｹ10109076 70ｸ803 6 703090ｹ6010903 0603apple04010304apple, 6 5 0ｹ06ｦ60307060309 6 50403 06080ｷ30902090ｹ6010903 08026ｦ90309 pfra 09 ppst. ｨｨapple 026ｦ06 76ｦ306fl 6ｦ203apple6ｦ3096 56ｦ0fl ｰ 6ｦ0apple6 50906016ｦ0fl ｱ 0307 02 6 7apple6ｦ0 046ｦ2090603090ｹ601076 707 08apple070906030708 09fl 6 50709 04 090302fl 0ｹ4030ｸ8, 08070603apple076ｦ0fl 6ｦ0 0ｹ1010702076 7090ｹ6010706 0706076 5. ｨｨapple076 5 07 06apple6ｦ06 509 6 50403 0ｹ16 507020ｸ90ｷ309070ｸ803 060ｷ36ｦ0apple096 5 ｨC 066ｦ2036ｦｷ309070906030601076 707 09 6 7apple6ｦ0 046ｦ2090603090ｹ601076 707, 08apple 08070ｹ309 0703 6ｦ306fl 08070602 0304. 07apple6ｦ0 046ｦ2090903 090209 066ｦ2036ｦｷ30907090903? 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Pourcel et al., 24; Song et al., 24; Zhou et al., 24a, b, c; Cui et al., 28; Li et al., 28), 09 0307apple03090ｹ6010706 0706076 50706 fl6 502fl0306fl 086ｦ0apple6ｦ0 096 7036ｦ0 07ｨｨ07. 07apple6ｦ0 046ｦ2090603090ｹ6016ｦ0fl 0307 02 060709 6ｦ06ｦ0 6ｦ309 6 5 01070301-0603 08070207080906 05ｨｪｨｴ 6ｦ0 0706076 5 0ｹ1010702076 7090ｹ6010903 6ｦ0-020ｸ9 0906 09 09070702076 707-086ｦ0apple6ｦ909030702076 7090ｹ6010903 036ｦ3apple096ｦ2076 5, 080702040ｹｸ803 6 5 0903apple6ｦ2076ｦ9096ｦ306010703 (ｨｪ04-6 50906010703) 08apple09apple07 0703 09 03apple0708090ｹ6010703 01apple0ｸ806090703 (05036ｦ306010703) 6ｦ3apple0708076 70703 070ｹ06ｦ06 76ｦ3 0ｹ4030ｸ8. 050709 04 090302 0ｹ4030ｸ8 ｨC 07 02096 76ｦ30ｸ806 086ｦ0apple6ｦ90903 09010903 0302010903 07apple076 50ｸ803 6 7apple0ｸ80904076 5 09 096ｦ60ｷ307 apple6ｦ90ｸ803. 0207-030307 6ｦ309 0807016ｦ96ｦ7, 0ｹ307 07 096 5apple6 709apple076 56ｦ2 0703 036ｦ2076 509apple040203076 707 066ｦ8apple070907070709076 707 6 50709 04 090302fl 08066 5 070304 apple010402096ｦ0 Y. pseudotuberculosis O:1b, 0109-040ｹ76 707 086ｦ0apple6ｦ909036ｦ0 0409apple0701076 707 01apple046 76ｦ0 066ｦ30ｸ803 apple6ｦ90707 - apple6ｦ90ｸ803 08096 507030ｸ803, 0702 20 030ｸ806. 0203 6ｦ96ｦ0 (Achtman et al., 1999, 24; Skurnik et al., 20). ｨｦ 020902076 703090ｹ6010706 03070207 07060309 6 50709 04 090302fl 0ｹ4-030ｸ8 066 509 030206036 5040ｸ903 036ｦ108 03076 709 0409apple070107 09096 50603-0ｸ8 030901apple07 090702076 7090ｹ60109, 0ｹ10809 03090702076 7090ｹ60109, 0ｹ10809090707030702076 7090ｹ60109 09 0ｹ1010702076 7090ｹ60109 6ｦｸ8 09 026ｦ1030ｸ8 (05fl0302076 5, 1989; 05040ｷ3076 5, 05040ｷ3076 56ｦ0, 26). 050709-0901026ｦ0 6ｦ96ｦ0apple036ｦ0fl 6ｦ40ｹ06ｦ0fl 0903apple096 76ｦ0: 016ｦ109 0607 0ｸ80309fl 08apple07090907040209 6ｦ0 6ｦ406 08026ｦ3 6 5 06030702 0703 6ｦ2-0703 08apple0704020703 ｨC 6 5 01070ｷ3 08070602 6 707 0901076 5076 707 08apple0907 6ｦ0 09 6 5 0807060202 0901076 507 6 5apple03fl, 01070307apple0ｸ8 08apple09-6 50209 01 6 507090901076 5090ｸ9 01apple6ｦ6 07086ｦ6076 707 01apple076 5fl076 707 086ｦ0apple6ｦ909036ｦ0 0909 06026ｦ0 076 509apple040203076 707 0109040ｹ76 707 030901apple07-6ｦ0 09 6 707 0409apple0701070304, 08070ｹ309 6 506066 503070304 apple6ｦ608apple070603apple6ｦ0-090ｸ9; 016ｦ1076 56ｦ0 08apple09apple07 6ｦ0 06020103096 50ｸ803 026ｦ10307apple076 5, 6 50ｸ8096 56ｦ06 5040903 096 5apple6 70ｷ3090ｸ9 0ｹ1030903 030901apple07 076 5; 6 7 09 016ｦ0-010903 07 apple6ｦ90703 0ｹ10307 08apple07090907040207? ｨｨ07090601 0ｹ16 507020ｸ90ｷ309070ｸ803 0107apple06 030901apple07 6ｦ0 0ｹ4030ｸ8 06060ｹ06ｦ6 6 5 0403 6 5 0706076 50703 030702010402flapple0ｸ8 6 703090109 6 5 apple6ｦ3016ｦ3 01070ｷ3080ｷ30909 07ｨｨ07 6 5 0703apple0ｸ86 5 0703 01026ｦ606090ｹ6010903 060306036 50ｸ803 6ｦ401: 09070702076 70909, 0ｹ1010702076 70909, 086ｦ0apple6ｦ9090307-02076 70909, 09076 7076 7apple6ｦ20909, 020909090702076 70909, 086ｦ20703070207-6 70909, 086ｦ207010209036ｦ30702076 70909 09 apple046 70903, 6 5 0307 6 5apple03fl 016ｦ1 0ｷ30207060307 080709036ｦ9 03076 70704apple076 56 50ｸ803 08apple070ｷ30606076 5 0ｹ16 507020ｸ90ｷ30909 08apple090ｷ30908096ｦ207 6 5070903070807 6 5 apple6ｦ3016ｦ3 016ｦ1076 707-0209 07 07 076 707 0909 apple6ｦ76 707 6ｦ8apple6ｦ06 50209fl 0907-02076 70909. ｨｬ06090209fl0309 08apple0ｹ906020ｸ803 01026ｦ606090ｹ6010903 6ｦ0-0401 01 6ｦｹ06ｦ204 XXI 6 5. 0ｸ80207 6ｦ107080207 0601070ｹ7 03070806036 507 026ｦ103076 5, 0807096 50702096 5040903 080702040ｹ903 0706036ｦ0-03070ｹ7 06076 7026ｦ6076 56ｦｸ8 09 04 0903020ｸ8 016ｦ0apple03090ｸ8 09-6ｦ3090109 apple6ｦ96 5090309fl 08apple09apple07 0ｸ8 6 5 01070ｷ3 08070602 6 707-0901076 5076 707 08apple0907 6ｦ0 09 6 5 6 70702070ｷ3 ｨｨ6ｦ26ｦ0apple0103090109. ｨｴ0307 0608070607 06036 5076 56ｦ207 6 50706060709 6ｦ90ｸ9 09060307apple0909 0207apple0309apple076 56ｦ0-09fl apple03apple07-09 06076 5apple030ｸ803 6ｦ0apple6ｦ2076 5 09 6 50403apple096 509 07-6 50706 0603apple04010304apple0ｸ8 03076 70903 06076 5apple030ｸ803 6 509 076 5 6 7apple0ｸ80904-076 5 09 08090ｸ70403 ｨC 0706076 50ｸ803 030709fl6 5 0ｹ403076 707 030901apple07 6ｦ0, 070607 07060306 0903 0ｹ1010702076 70909, 066 5fl096ｦｸ803 06 6ｦ603040802-0903 09 0701070ｹ06ｦ903 0901076 5076 707 6 5apple0309 09, 016ｦ1 ｨ ｨ 0505ｨｪｨ ｨｹ ｨｺ0204. 05ｨｺｨ ｨｹ 03ｨ ｨｦｨｦ07ｨ ｨｮ050102ｨｹ 6ｧ8 6 29
13ｨｨｨｺｨ 0409ｨ ｨｨ 05ｨ 05ｨｦｨｦ03ｨｺ02ｨ ｨｦ050204ｨ ｨｹ 02ｨ 01ｨｺｨｦ0302 ｨｮｨｬ0201 Yersinia pestis 647 0602 06036 509, ｨC 09060307apple0909 060306036 50ｸ803 08apple09apple07 0ｸ803 070ｹ06ｦ06 7076 5 0ｹ4030ｸ8. 07026ｦ06 50706 08apple090ｹ90706 06076 5apple03076 707 0209 apple06036 56ｦ0 0207 6 5 6 50708apple07066ｦ3 08apple070906030708 09fl 09 0ｹ16 507020ｸ90ｷ30909 030901apple07 6ｦ0 0ｹ4-030ｸ8 080706020408090207 6 70204 070107 08apple0703096 507apple0ｹ9 07 076 707 0909 0706076 50ｸ803 08070207080906 01026ｦ606090ｹ6010706 0307apple0909 08apple09apple07-0706 070ｹ06ｦ06 7076 507060309 0ｹ4030ｸ8 ｨC 07 6 70702076 7090ｹ6010706 apple6 507060309 6 50709 04 090302fl ｨC 06 6 50ｸ86 507 6ｦ309 076 506040903 0207-09060602 076 56ｦ0-0906 6ｦ103apple0906 apple07 6ｦ0 Yersinia, 0706036ｦ06 502fl0ｸ90ｸ7090309 09 036ｦ206046 707 06070309fl 6 5 6 707 020902076 703090ｹ6010706 0307-0207 07060309. ｨ 030ｸ90306fl 09 apple046 709 08apple090ｹ90ｸ8. 046ｦ0 0706076 56ｦ909 0207-6ｦｸ803 6 50ｸ8fl6 502 08apple 0701 0ｹ403076 707 030901apple07 6ｦ0, - 026ｦｸ90306fl 08apple 080702070809fl 07 6 5apple0309 090204apple016ｦｷ30909 08apple 01076 50706 09 070ｹ0apple06 0207apple03 09 096 5apple6 70ｷ30909 0706076 5-0ｸ803 02090203090ｹ6010903 02090906 ( 09076 56ｦ0apple076 5, 0807 6 509 076 5) 6 509-6ｦ0 Y. pestis. 0207-09060602 076 56ｦ9fl 0807096 50702090209 08070603apple070903 ｰ040107apple0ｸ8 ｱ 09 ｰ07apple090309apple076 56ｦｸ8 ｱ 6 76ｦ2076 7090ｹ60109 apple6 5fl apple07 6ｦ0 Yersinia, 030703fl 0ｸ8 09 060807apple0ｸ8, 07 07060307060ｸ8 07 060408 09fl, 09 096ｦ20708090209 0706076 50ｸ8 0209-02076 7076 7apple6ｦ2090ｹ601076 707 6ｦ06ｦ209096ｦ0 6 50403apple096 509 076 5076 707 apple6ｦ9-0707 apple6ｦ909fl 6 50709 04 090302fl 0ｹ4030ｸ8, 0ｹ06 707 03076 7026ｦ0 0807096 507-020903 06 01026ｦ606090ｹ6016ｦ0fl 0307apple09fl 08apple09apple07 0706 070ｹ06ｦ06 707-6 507060309 0ｹ1030706 07020909. 05040ｸ706036 50ｸ806 0706036ｦ30701 08apple0902070809fl 0207-03- 0307 076 5 01 020902076 7030901 030901apple07 6ｦ0 Y. pestis 06070603070903 6 5 030703, 0ｹ307 apple090402036ｦ30ｸ8 6ｦ06ｦ209096ｦ0, 08apple076 507 0903076 707 0807 0703-020ｸ803 6 70ｸ803 0603apple04010304apple6ｦ3 (0207-036ｦ0apple01apple6ｦ3, 6 7-0ｸ803 02apple6ｦ06 703036ｦ3), 08apple0706fl0306fl 080706apple 06036 507 6ｦ0 020902076 7090ｸ9 09 036ｦ106070703090ｸ9 0909040ｹ06ｦ30706 6 7apple0408080ｸ8. ｨｴ0307 03070803 08apple096 507 0903 01 06apple090ｸ803 0906016ｦ809fl03, 070607 07 6 5 030703 0602040ｹ06ｦ0, 01076 7 6ｦ0 090608070209040ｸ90306fl 09-090ｹｸ8 0ｹ10109030802flapple0ｸ8 090209 07020409 6 50ｸ8 07apple0109. 0208 04 0303 090307 0ｹ10307, 07020406 0ｹ06ｦ6030ｸ9 0909-096ｦ0 07apple076 7076 509090ｸ8 09 03apple04 07030107060309 0207-030307 0901, 036ｦ0apple6ｦ103apple- 07 08070ｹ309 02fl 6 50603 apple6ｦ0 0703 0807 020902076 7030901 09 02090207-6 7076 7apple6ｦ20909 (02 apple6ｦ30607, 27). 0ｸ7 07 09 0706036ｦ30701 0207-030307 076 5 06070603070903 6 5 030703, 0ｹ307 6ｦ0 0307080702076 7090ｸ9 080702040ｹ06ｦ30ｸ803 020902076 703090ｹ6010903 apple6 56 5 096ｦｹ9030207 6 50209fl09 07016ｦ90ｸ86 56ｦ3 6 50ｸ8 07apple 030702010402flapple076 707 036ｦ0apple01apple6ｦ0; 6 707 6ｦ0 016 56ｦ30ｸ806 6 50ｸ8 07apple 08apple 06036ｦ06 502fl03 070204040ｸ9 08apple07 020304. 05 0807-0603apple070909 020902076 70906 0ｹ403076 707 030901apple07 6ｦ0 030307 0ｸ8 06 apple6ｦ90ｸ80309 036ｦ0apple01apple6ｦ309 6ｦｸ903 096ｦ30307 apple6ｦ902090ｹ06ｦｸ90ｸ709-06fl apple090402036ｦ30ｸ8 (Achtman et al., 24; Pourcel et al., 24; Gu et al., 26; Torrea et al., 26; Vergnaud et al., 27; Cui et al., 28; Li et al., 28). ｨｺ090402036ｦ30ｸ8 0207-09060602 076 56ｦ906 6 50709 04 090302fl 0ｹ4-030ｸ8 060207apple0309apple076 56ｦ209 08070ｹ309 6 50607 0ｸ7 0309, 0ｹ307 066 5070606036 56ｦ0 6 707 6 50ｸ80607010706 6 509apple04020307060309 09 086ｦ3076 707-060309 6 5070909010209, 080703090307 040ｹ06ｦ60309fl 0304036ｦｷ30906, apple010703 09-6ｦｷ30906 09 020ｷ30906, 08040303 6 50603apple6ｦ96 56ｦ9fl 6 5 6 70703 08apple - 01076 50706 0207apple030ｸ8 0ｹ10109076 70ｸ803 6 703090ｹ6010903 0603apple04010304apple apple6ｦ9076 707 apple6ｦ903apple6ｦ0 ｨC 0307 09020ｸ803 6 7076 5, 6 70ｸ803 0207-01076 5, 03apple6ｦ608070907076 5, 08026ｦ90309 ｨC 0909 6 50406 06apple 0ｸ8 09/090209 0703 016ｦ10903-0209 07 0109040ｹｸ803 030901apple0707apple6 76ｦ90903076 5, 0807apple0706 036ｦ106070703090ｹ60109 6ｦ2010903 0703 0ｹ403076 707 030901apple07-6ｦ0. ｨｦ 6ｦ107 09 6 5 07 0706 0909 apple6ｦ0 0703 0807 07 076 707 08026ｦ06ｦ0 04016ｦ90ｸ86 56ｦ306fl 6ｦ0 080ｸ806 090603070ｹ901 6 50603apple6ｦ96 56ｦ30ｸ803 6 703090ｹ6010903 0603apple04010304apple 09 apple6ｦ601apple0ｸ86 56ｦ306fl 0603-06036 50ｸ806 08apple070ｷ30606 0903 6 5 apple09fl 6 5 076 50707 apple6ｦ9076 56ｦ0-040ｸ9 0ｹ403040ｸ9 0102030104. 01 08apple0903apple04, apple6ｦ601apple0ｸ86 56ｦ306fl 08apple090ｷ30908 ｰ6 5 apple09fl ｱ ｰ06080ｷ30902090ｹ6010903 ｱ 08026ｦ90309 ppst 09 pfra: 06080ｷ30902090ｹ60109 08026ｦ90309 0ｸ8, 0807 0708apple 02-090ｸ9, 0207apple0309apple040ｸ90306fl 0ｹ1 076 707 ｨC 080703070304 0709 09 0608-0ｷ30902090ｹｸ8, 09 060209 6 7076 507apple0903 07 0903 096ｦ90306036 5076 56ｦ909 (acquisition) 0703 09096 506030ｸ803 0807016ｦ0 0ｹ408apple07 0ｸ803 0207apple03, 0307 06080ｷ30902090ｹ601090309 0903 6ｦ90ｸ86 56ｦ3 0602 0403. 01apple04080ｸ8 07 076ｦ1030ｸ8 09090309fl 6 707036ｦ0 6 50709 04-090302fl 0ｹ4030ｸ8, 01apple040307 03fl0ｸ90ｸ709 03036ｦ0 0702090ｹ60109 6 5096ｦ903070703070409fl 07 09070ｹ706 030901apple07 0706 0102030109 06 6 50406 06apple 0706, 0703apple6ｦ86ｦｸ903 066ｦ2036ｦｷ3090709060306010906 0807 0307 01 08apple07 0203 6 509 0707 apple6ｦ9076 56ｦ9fl 04 08apple07016ｦ0apple090703 (apple0906. 1). 020901apple07 0ｸ8 0ｹ4030ｸ8, 0ｷ309apple01040209apple040ｸ90ｸ709 6 5 0807080402fl0ｷ309fl03 0807026 50109 03apple6ｦ0 036ｦ0 (ｨ 6ｦ0 6ｦ6016ｦ2, 026ｦｹ804apple09fl, 0207-6 7070209fl), 030307 apple0ｸ9030706 (046ｦ0) 09 07 0ｸ801076 50706 (056ｦ06 706036ｦ0, 016ｦ06 5016ｦ9), 07 026ｦ0 6ｦｸ903 01070307apple0ｸ80309 09070309-03090ｹ601090309 066 5070606036 56ｦ309, 06 0209086ｦｸ90ｸ7090309 0903 06 08066 5 07-0304 apple010402090ｸ803 030901apple07 0703 (0507036ｦ0apple6ｦ0 06010906, 1998; Zhou et al., 24a, b). ｨｨ07 0703 020ｸ803 0207-08apple09096ｦ16ｦ3 03076 709 09060602 076 56ｦ30209 036ｦ108 06 0209086ｦｸ903 08066 5 070304- apple01040209076 707 030901apple07 6ｦ0 06 6 50709 04 090302fl0309 0ｹ4030ｸ8 0909 0807080402fl0ｷ30906 0807026 50701 016ｦ06 5016ｦ96ｦ0, 05apple 06 02090909 09 0109-036ｦ0fl (060902090808076 5, 20; 010401026 56ｦ0 09 apple., 22; Motin et al., 22; Achtman et al., 24; Song et al., 24; Zhou et al., 24a, b, c; Vergnaud et al., 27; Cui et al., 28; Li et al., 28). 050307 06036 507 0807 0907030903090ｹ6010903 09 030702010402flapple0ｸ803 08apple09096ｦ16ｦ3 apple07 090207 0309 07 080702-6 5070ｹ3 09076 56ｦ0apple 016ｦ1 08apple0703080403070ｹ706 0207apple03 0308 04 08apple 01076 50ｸ803 08066 5 070304 apple010402090ｸ803 030901apple07 0703 09 070ｹ0apple0903 0ｹ4030ｸ803, 6ｦ0 030901apple07 076 5 0909 6 50603 0807026 5070ｹ903 070ｹ06ｦ06 7076 5, 6ｦ8 6 7076 7apple6ｦ2090ｹ60109 04 6ｦ20ｸ803 apple046 7 0703 apple046 76ｦ0 06036ｦ209 070307060903 01 07 070304 09076 56ｦ0apple04 090209 0807-6 509 04 ｨC 0807026 5070ｹ304 (pestoides, microti, microtus, minor): 036ｦ10903 07 apple6ｦ90703 6ｦ0apple6ｦ2 0807 6 509 6ｦ0 Y. pestis ｰpestoides ｱ 07016ｦ96ｦ206fl 03appleapple090307apple096ｦ207 apple6ｦ907apple6 56ｦｸ803 6ｦ0 060107020107 ｰ070603apple076 5076 5 ｱ, 6ｦ20107 04 6ｦ20ｸ803 apple046 7 0703 apple046 76ｦ0. 05apple046 709 apple6ｦ90707 apple6ｦ90ｸ8 060ｷ36ｦ0apple0909 096 5apple6 70ｷ30909 08066 5 070304 apple01040209076 707 09 0ｹ403076 707 030901apple07 076 5, 0607-09 6ｦｸ8 6ｦ0 0706076 5 0207-0807 0307 6ｦ0, 6 5 08apple090ｷ30908 0807 07-0ｸ8 08apple096 5 070304 6ｦ0 apple0906. 1. 076ｦ0apple6ｦ103apple0ｸ80309 070607 070603fl0309 060ｷ36ｦ0apple09fl 066ｦ2-036ｦｷ30907076 707 08apple070906030708 09fl 09 0ｹ16 507020ｸ90ｷ30909 030901apple07 6ｦ0 0ｹ4030ｸ8 03070807 6ｦ96 56ｦ3 0602 040ｸ90ｸ709. 1. 0502040ｹ06ｦ607 6 507090901076 509 6 509 0707 apple6ｦ9040ｸ90ｸ706 0304036ｦｷ30909 (036ｦ1apple070304036ｦｷ30909, 06090603030706 0304036ｦｷ30909, 07ｨｨ07) 6 7-0209 07 6 5 0309apple 090703070609030207 6 7076 7apple6ｦ2090ｹ601076 707 0306036ｦ0, 060706036ｦ06 56ｦ0 09 0603apple04010304apple0ｸ8 06apple 0ｸ8 ( 09076 7070ｷ307096ｦ0). 2. ｨｦ03apple090ｷ36ｦ9 0807060308076 707 0104030402fl03096 5076 707 036ｦ0- apple6ｦ103apple6ｦ0 0207apple0309apple076 56ｦ9fl 04 0ｹ403076 707 030901apple07 6ｦ0 0ｹ16 507020ｸ9-0ｷ309070ｸ803 076 50406036 5. 3. ｨｦ0307apple6 56ｦ70603 08apple07 02030ｸ8 0ｹ16 507020ｸ90ｷ30909 6 50709 04 09-0302fl 0703 0ｹ16 507020ｸ90ｷ30909 030709fl096ｦ0 09 08apple07060ｹ9016ｦ0. ｨ ｨ 0505ｨｪｨ ｨｹ ｨｺ0204. 05ｨｺｨ ｨｹ 03ｨ ｨｦｨｦ07ｨ ｨｮ050102ｨｹ 6ｧ8 6 29
13648 05040ｷ3076 5, 05040ｷ3076 56ｦ0 Yersinia pestis pestoides Yersinia pestis ant med ori Yersinia pseudotuberculosis O:1b II 020902076 703090ｹ60107 0607 0ｸ80309 ｨC 6 50603apple6ｦ96 56ｦ9 08026ｦ90309 0ｸ8 ppst 05090309 080603090ｷ3096ｦ0 I 020902076 703090ｹ60107 0607 0ｸ80309 ｨC 6 50603apple6ｦ96 56ｦ9 08026ｦ90309 0ｸ8 pfra1 05090309 Fra1 ｨｺ0906. 1. 0503036ｦ0 066ｦ2036ｦｷ30907076 707 08apple070906030708 09fl 09 0ｹ16 507020ｸ90ｷ30909 6 50709 04 090302fl 0ｹ4030ｸ8 08040303 ｰ6 50603apple6ｦ96 56ｦ9fl ｱ 06080ｷ30902090ｹ6010903 08026ｦ90309 pfra 09 ppst. 0309076 56ｦ0apple0ｸ8: ant ｨC antiqua; med ｨC medievalis; ori ｨC orientalis. 4. ｨｦ03016ｦ9 0703 036 507apple0ｹ6010706 0207apple030707 apple6ｦ9040ｸ90ｸ706 apple07-0209 060306036 5076 707 0703 07apple6ｦ0 09 6ｦ0 6ｦ803096 5076 707 036ｦ0apple6ｦ103apple6ｦ0 6 509 0707 apple6ｦ9076 56ｦ302076 707 08apple070ｷ306066ｦ0; 0ｹ16 507020ｸ90ｷ309070ｸ806 08apple070ｷ30606 apple6ｦ606036ｦ3apple096 56ｦ306fl 016ｦ1 6ｦ0 6ｦ8036ｦｷ309076 709, 060306036 5070304 0703 07apple04 07036 507 090306fl 6 5 040ｸ706 apple070209 6 5 08apple070906030708 0909 0ｹ4030ｸ8: 076 50ｸ806 0304036ｦ3 0807086ｦ0 6ｦ3 6 5 026ｦ06 70708apple09fl030ｸ8 040602076 509fl 09 06apple6ｦ904 6ｦ3 076 5040ｸ9 0ｹ16 507020ｸ9-0ｷ30907040ｸ9 6 5036 5. 5. ｨｦ03016ｦ9 0703 0807080402fl0ｷ30907076 707 0807 0307 6ｦ0: 0706076 56ｦ0fl 09090ｷ36ｦ0 0ｹ16 507020ｸ90ｷ30907076 707 08apple070ｷ306066ｦ0 ｨC 09090ｹ06ｦ0fl 030901apple07 6ｦ0fl 010203016ｦ0. 6. 050707036 50306036 509 060ｷ36ｦ0apple09fl 0309080702076 7090ｹ6010903 08apple 06036ｦ06 50209fl03 07 070306040306036 50909 08appleapple0ｸ86 5076 707 applefl- 6ｦ0 apple03apple0706080103096 50ｸ803 08apple0703080403070ｹｸ803 0207apple03; 6ｦ06ｦ0-02076 7 0807 07 0706 060903046ｦｷ30909 6 5 086ｦ207030702076 70909 ｨC 01070ｷ308-0ｷ309fl 08appleapple0ｸ86 50906030706 086ｦ207030702076 7090ｹ6010706 02030708090609, 04036 5apple08 6ｦｸ90ｸ706 apple6ｦ207 070306040306036 509 08apple0703080403070ｹ0-0ｸ803 0207apple03, 6ｦ0 066 5fl09 0ｹ103076 707 fl6 50209fl 06 08070207030706 086ｦ207030702076 7090ｹ6010706 02030708090609. 7. ｨｰ09apple070107 08apple09fl036ｦ0fl 6 5 01026ｦ606090ｹ6010903 0ｹ16 507020ｸ90ｷ309-070ｸ803 0307apple09fl03 01070ｷ3080ｷ309fl 08apple6ｦ0 6ｦ8036ｦｷ30909 6 5 0ｹ1030703 060ｷ36ｦ0apple0909 6ｦ307 0903 08apple0902070809fl. ｨ 036ｦ1, 6 509 0707 apple6ｦ9076 56ｦ9 6 50709 04 090302fl 0ｹ4030ｸ8, 016ｦ1 6 707 6 509 fl03 030702010402flapple0ｸ8 6 703090109, 08apple07040207 6 5 07apple6 76ｦ0-090903 016ｦ1076 707-0307 6 509 6ｦ0 0807026 50109 apple07 6ｦ0 Microtus (090209 09076 707 02090901076 707 apple07 6ｦ0) 0703 09090ｹｸ803 030901apple07 0ｸ803 0102030701 6 50709 04 090302fl 08066 5 070304 apple010402096ｦ0 6ｦ0 0706076 5 0602040ｹ06ｦ60ｸ803 01apple04080ｸ803 08apple07 apple6ｦ9076 56ｦ906 6 707036ｦ0 090209 0807 07 apple6ｦ9070304 6 50ｸ8apple6ｦ8090ｸ9 09096 50603076 707 6ｦ309 6ｦ0apple6 509-0906036ｦ0 ｨｺ. 070702 040309 036ｦ0 ｨC 0703 04 6ｦｹ2096 50ｸ803 03070603apple076 5 (Goldscmidt, 1940). 01070ｷ3080ｷ309fl 066ｦ2036ｦｷ30906 08040303 6 50603apple6ｦ96 56ｦ9fl 01apple0408-0ｸ803 6 70ｸ803 020701076 5, 03apple6ｦ608070907076 5 09 08026ｦ90309 04 0903026ｦ0. 040703ｨB0302030ｸ803 066 5070606036 50703 020ｸ9 076 707 07apple6 76ｦ909036ｦ0, 6 5 030703 0ｹ90602 08apple07016ｦ0apple090703076 707, fl6 502fl0306fl 09036 7apple6ｦｷ309fl 6 707 0603apple04010304apple0ｸ803 0ｹ1020303076 5 (08apple09096ｦ0-01076 5) (Kurland, 20). 05076 7026ｦ607 01026ｦ606090ｹ6010903 08apple - 06036ｦ06 50209fl03, 036ｦ1apple070304036ｦｷ30909 6ｦ0 6ｦ803096 50ｸ8, 036ｦ1apple07-0304036ｦ30ｸ8 0809090608070607 0ｸ8 6 5 060306036 50706 06apple 0909-096ｦ0 6ｦ0apple040409fl 0307apple0207020909090702076 7090ｹ6010903 0107appleapple- 02fl0ｷ30906 09 010707apple 096ｦｷ30906 07apple6 76ｦ76 5 (07apple6 76ｦ709 076 5, 07apple6 76ｦ0-0202) 09 0ｷ3020706030ｸ803 07apple6 76ｦ90903076 5. ｨ 06 6 56ｦ807 096ｦ3030903, 0ｹ307 0309 07 066ｦ2036ｦｷ309-070703 6 507090901076 50909 030901apple07 6ｦ0 0ｹ4030ｸ8 6 506-036ｦ109 fl6 502fl0306fl 06076 5apple0407 06060807apple0ｸ803 06apple 09 0307020104-02flapple0ｸ803 6 7030901076 5. ｨｪ6ｦ1, ｨｺ (Wren, 23) ｰ016 56ｦ307-6 50ｸ806 06016ｦｹ701 6 5 036 7076 509 07016ｦ0 (quantum leap in eye blink) ｱ 08066 5 070304 apple01040209076 707 030901apple07 6ｦ0 O:1b 6 5 6 50709-04 090302fl 0ｹ4030ｸ8 apple6ｦ606036ｦ3apple096 56ｦ3 6 5 6 70702076 7090ｹ6010703 036ｦ604036ｦ0 6 5apple0309 09, 6 5 036ｦ10703 0602040ｹ06ｦ0, 6 50ｸ80307 0903 096ｦ0 apple6ｦ30109 0807fl0309fl 0807060308076 707 6 7apple6ｦ0 046ｦ2076 707 06036ｦ76 5-0209fl 6 509 6ｦ0. 04 6ｦ06 507 070804 020901076 56ｦ06ｦ0 06036ｦ3fl 09096 506030ｸ803 0109036ｦ6-06010903 030702010402flapple0ｸ803 6 7030901076 5, 6 5 01070307apple0706 6ｦ06 50307apple0ｸ8 08apple 06036ｦ06 5090209 0307 02 0807060308076 707 0207apple0309apple076 56ｦ9fl 6 703090ｹ6010903 0603apple04010304apple, 0107 09apple040ｸ90ｸ70903 066 5070606036 507 6 509apple04-020307060309 030901apple07 6ｦ0 0ｹ4030ｸ8 08070602 0703 0209fl 09060307 07-6 707 0102076ｦ0 0703 0807080402fl0ｷ30909 08066 5 070304 apple01040209076 707 030901apple07 6ｦ0 (Zhou, Yang, 29). 050903080307036ｦ3090ｹ7, 0ｹ307-06030703applefl 6 5 0ｷ3020703 6ｦ0 0ｹ101020103090ｹ6010906 0807 0307 6 5 6 50708apple0706 08apple070906030708 09fl 0ｹ4030ｸ8, 6 50ｸ8apple6ｦ9096 504090606fl 6 5 03036ｦ90603090ｹ6010703 06076 5030ｸ70909 086ｦ0apple6ｦ0 096 7030ｸ8 07ｨｨ07 09 06020103076 7-096ｦ0, 030702010402flapple0ｸ8 6 703090109, 6ｦ1070ｷ3, 6ｦｹ96ｦｸ903 070ｸ7040ｸ76ｦ3 07 0307 0903070603 090902070809fl 066 5070903 6 5096 702fl 076 5 6 5 03apple03096ｦ3 05ｨｪｨｴ (08apple6ｦ0 6ｦ8036ｦｷ309fl, 6 5apple0309016ｦ207 6ｦ0-0602 076 56ｦ9 08apple09096ｦ1076 5, apple0602 6 7076 5, 09090702fl0ｷ309fl 010207-076 5 09 6 50403apple096 509 076 50ｸ803 0207apple03, 6 707030908090ｹ6010906 09 02-07030908090ｹ6010906 060706036ｦ06 5 0807080402fl0ｷ30909, 0807030701 6 7076 5, 06- ｨ ｨ 0505ｨｪｨ ｨｹ ｨｺ0204. 05ｨｺｨ ｨｹ 03ｨ ｨｦｨｦ07ｨ ｨｮ050102ｨｹ 6ｧ8 6 29
13ｨｨｨｺｨ 0409ｨ ｨｨ 05ｨ 05ｨｦｨｦ03ｨｺ02ｨ ｨｦ050204ｨ ｨｹ 02ｨ 01ｨｺｨｦ0302 ｨｮｨｬ0201 Yersinia pestis 649 020103096 507 08apple0903040ｸ706036 507, 060306036 50ｸ806 0703 07apple). 05 08apple0703096 5076 506 086ｦ0apple6ｦ0 096 703 07ｨｨ07, 0207apple0309apple076 56ｦ9 0ｹ403-076 707 030901apple07 6ｦ0 0909 08066 5 070304 apple01040209076 707 6 5 060707036 503-06036 50909 06 080702070809fl03 05ｨｪｨｴ, 07020807 08apple070307 0903 070307-0609030207 03 0207, 080706030807, 6 7apple6ｦ0 046ｦ207, 07 07 fl096ｦ30207 apple6ｦ06 50703apple07. ｨｮ0307 0ｸ8 08apple09fl03 ｰ 6ｦ0apple6 509-06010906 ｱ 080403 6 509 0707 apple6ｦ9076 56ｦ9fl 6 50709 04 090302fl 0ｹ4030ｸ8, - 07 0307 090307 0807016ｦ96ｦ3 6ｦ2090ｹ9 08apple0307 0706 06apple 0ｸ8 09 0807-016ｦ96ｦ3 090209 02076 7090ｹ60109 07 0706076 56ｦ3 06040ｸ706036 5076 56ｦ9 08apple0703080403070ｹｸ803 0207apple03 016ｦ1 apple6ｦ20ｸ803 090209 apple03apple07-06080103096 50ｸ803 08apple09apple07 0ｸ803 fl6 5020906. 01026ｦ606090ｹ6016ｦ0fl 030307 0702076 709fl 020902076 703090109, 6ｦ0-0909apple040ｸ90ｸ76ｦ0fl06fl 6ｦ0 030307 ｰ03apple0706076 707 086ｦ0apple6ｦ202020909036ｦ0 ｱ, 060403 01070307apple076 707 06070603070903 6 5 06076 7026ｦ6076 56ｦ909 6ｦｸ803 06apple6ｦ06 50903020706 6ｦ06ｦ307030909 09 0307apple020702076 70909, 0ｹ103 apple0907-02076 70909 09 086ｦ207030702076 70909, 090303 01apple6ｦ6 076 7apple6ｦ90ｹ0-0ｸ8 6 5070903070807060309 6 5 apple01070603apple04010ｷ30909 0ｹ16 507020ｸ90ｷ309070ｸ803 08apple070ｷ30606076 5 04 08apple07016ｦ0apple090703. ｨｨ070ｹ103070304, 036ｦ1 016ｦ1 6ｦ0 6ｦ8036ｦ0-0ｷ309fl 0ｹ403076 707 030901apple07 6ｦ0 6ｦ8apple6ｦ06 5026ｦ0 01 06apple 6 707 07 09-036ｦ9fl ｨC 086ｦ0apple6ｦ909036ｦ0apple0706 0609060303 6 7apple0ｸ80904ｨC 0207036ｦ0, 02fl 08070603apple0709fl 060ｷ36ｦ0apple09fl 6 707 08apple070906030708 09fl 09 0ｹ16 507020ｸ9-0ｷ30909 030ｸ8 060706apple 0703070ｹ90209 6 509036ｦ9 6ｦ0 0ｹ1010702076 7090ｹ6010903 6ｦ60801036ｦ3 08apple07 02030ｸ8 ｨC 6 5096ｦ903070703070409fl03 6 50709 04 09-0302fl 06 010701apple030ｸ803 6 509 0703 6 7apple0ｸ809046ｦ0 ｨC 03076 707020601076 707 0604apple016ｦ0-036ｦ0apple 6ｦ06 76ｦ06ｦ0 Marmota sibirica 09 06080ｷ30902090ｹ6010706 02070309 6 5apple6ｦ9090606010903 0604apple01076 5 Oropsylla silantiewi. 050ｸ8-07apple 0ｹ1030903 07 ｨB0103076 5 0ｸ802 0807 06016ｦ96ｦ0 060403030706 096ｦ906, 6ｦ10708020ｸ803 0807 apple6ｦ902090ｹｸ803 6ｦ60801036ｦ3 0307apple0909 ｨｨ6ｦ06 5-02076 50601076 707 (1946) 07 08apple09apple07 0706 070ｹ06ｦ06 7076 507060309 03apple6ｦ6030906-06096 50ｸ803 07020906 0ｹ2076 5016ｦ0 09 08096 507030ｸ803 09 6ｦ4090309 0ｹ1010702076 7090ｹ601090309 6ｦ0 020ｸ9 09fl0309 6 5 0903apple6ｦ20706 02090909. ｨｦ0607 07060309 0ｹ1010702076 70909 03076 707020601076 707 0604apple016ｦ0-036ｦ0apple 6ｦ06 76ｦ06ｦ0 09 02070309 O. silantiewi 0807096 50702090209 060706036ｦ0-6 50903 08apple 06036ｦ06 50209 07 07 060307fl030206036 56ｦ3, 08apple09 01070307apple0ｸ803 0ｹ1010601apple03030ｸ8 09 01apple076 5 030201070809036ｦｸ90ｸ70903 07016ｦ96ｦ0-020906 6 5 03060703 04060307060ｹ96 50703 0107036ｦ103 09 0109040ｹｸ806 030901apple07 0603076 7 06076 5apple040903 0ｹ16 507020ｸ90ｷ309070ｸ806 08apple0307 6 5 08apple090ｷ30908096ｦ207 076 5040ｸ9 06apple 04 07 09036ｦ9fl, 0307 0603 6 5 076 5040ｸ9 6ｦ0 6ｦ803096 5040ｸ9 090704. ｨｨ07 apple07 07 0ｹ1030703 060ｷ36ｦ0apple0906 070809066ｦ0 6ｦ309 apple6ｦ0 (05040ｷ3076 5 09 apple., 1997; 05040ｷ3076 5, 05040ｷ3076 56ｦ0, 20, 26, 28). ｨｦ 0703apple6ｦ86ｦ3 0ｹ1010702076 7090ｹ6010906 0807 0307 01 apple- 01070603apple04010ｷ30909 020902076 70909 apple07 6ｦ0 Yersinia 09 6 509 6ｦ0 Y. pestis 6ｦ0 0706076 5 6ｦ06ｦ209096ｦ0 6ｦ0 6ｦ803096 50ｸ803 066 5070606036 5 030709fl6 5, 0903 08apple07060ｹ901076 5 09 066ｦ3076 707 6 50709 04 090302fl 6ｦ0 0207 6 70207 6ｦ20ｸ803 010209036ｦ3090ｹ6010903 0909030906 08apple09apple07 0ｸ8. ｨｺ090402036ｦ30ｸ8 6ｦ40903 0807026 50ｸ803 09070702076 707-086ｦ0apple6ｦ9090307-02076 7090ｹ6010903 09 0ｹ1010702076 7090ｹ6010903 09060602 076 56ｦ906 026 70209 6 5 0706076 504 08apple090ｷ30908096ｦ207 076 50706 01070ｷ3080ｷ30909 08apple070906030708-09fl 0ｹ4030ｸ8: 030901apple07 Y. pestis 096 5apple6 709apple076 56ｦ2 0703 0109-040ｹ76 707 086ｦ0apple6ｦ909036ｦ0 ｨC 08066 5 070304 apple01040209076 707 030901apple07 6ｦ0 1-6 707 06apple070309086ｦ0 (O:1b) 6 5 01070ｷ3 080709 6 707 0802060603070ｷ3-6ｦ0 (6 5 036ｦ10609036ｦ207 03070207 07 066ｦ0apple036ｦ60107 6 5apple03fl, 22ｨC15 030ｸ806. 0203 6ｦ96ｦ0 ) 6 5 0903apple6ｦ20706 02090909 (apple0906. 2) 6 5 086ｦ0apple6ｦ909036ｦ0apple0706 0609060303 ｰ03076 7070206010906 0604apple0701 ｨC 0207036ｦ0 O. silantiewi ｱ, 01070307apple040ｸ9 03070807 08apple 06036ｦ06 50903 016ｦ1 066 507-07 apple6ｦ907 030308apple6ｦ304apple07-090303040702076 7090ｹ60107 ｰ0ｹ16 507-020ｸ90ｷ3090707 080702 ｱ 06 096ｦ86ｦ9070703 030308apple6ｦ304apple 08apple0903apple- 07 5ｨC37 紊; 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05040ｷ3076 5, 05040ｷ3076 56ｦ0 650 056ｦ3 6ｦ601 07 0807 6 5 0509 0909 03070207 6ｦ9 22ｨC1 07 5 02 03 04 0ｸ8 06 07. 09 01 02 07 08 03 6ｦ96ｦ0 0703. 096ｦ0 02 07 09 6 7 07 04 036ｦ0. 060601 02 6ｦ0 6ｦ0 05 43 1 2 056ｦ36ｦ0 07ｨC07 07 6ｦｲ4 09060601 6ｦ0 09 6ｦｲ4 0907 6ｦ0 3 4 5 ｨｺ0906. 2. ｨｺ6ｦ607 08070906030708 090ｲ4 03090107 6ｦ0 0ｹ4030ｸ8 6 5 09036ｦ20706 02090909. 1 ｨC 06076 5036ｦｲ4 09076ｦ0 6 50ｹ706 03090207030ｸ8; 2 ｨC 6ｦ9 0ｸ8 6ｦ6070ｸ8 05070603070ｹ76 707 08070204046ｦ90ｲ4; 3 ｨC 6ｦ607 08070906030708 090ｲ4 03090107 6ｦ0 0ｹ4030ｸ8 6ｦ0 06076 5030ｸ70706 030903070909 056ｦ36ｦ7-0707 090606010706 6ｦ9 0706 09070ｸ8 09 09070ｸ8 03076 70702030309020ｸ803 080707 ; 4 ｨC 6ｦ6080706036ｦ9 036ｦ10609036ｦ2076 707 0ｹ36 503090ｹ76 707 (066ｦ36ｦ601076 707) 080703070207 6ｦ90ｲ4 07 6 76ｦ90ｷ3 066 50706 0707 09 (43 06.04.); 5 ｨC 6ｦ06ｦ2 086 5090ｹｸ803 080907 0ｸ803 070ｹ06ｦ06 7076 5 0ｹ4030ｸ8 6 5 6ｦ9 0ｸ803 09 0604 6ｦ9 0ｸ803 6ｦ6076ｦ3 6 56ｦ90909. 050302016ｦ309 0807016ｦ96ｦｸ8 6ｦ86ｦ06 502090ｲ4 060306036 50706 0ｹ10106086ｦ60909 03090107 6ｦ0 0909 08070804020ｲ40ｷ30906 0604016ｦ0-036ｦ0 6ｦ06 76ｦ06ｦ0 6 5 01070ｷ3 0802060603070ｷ36ｦ0 09 6 70702070ｷ3. 020ｲ4 080307 6ｦ0 0109040ｹ76 707 086ｦ06ｦ909036ｦ0 01 06040ｸ706036 5076 56ｦ90ｸ9 6 5 02090302 09 01076 509 6ｦ103096 5076 707 0307090ｲ4096ｦ0. 4. 05 066 5070606036 56ｦ3 08066 5 070304 01040209076 707 03090107 6ｦ0: 096 56ｦ9096 507060309 (0608070607 07060309 096ｦ06ｦ86ｦ3 0308020701076 50ｸ803 08096 507030ｸ803 08070602 01040203096 56ｦｷ30909 0809 0909010903 08020ｸ906076 50ｸ803 0303086ｦ3046ｦ3 0701070207 5 紊), 6ｦ2090ｹ909 6 5 010203016ｦ3 6ｦ309026ｦ06 7070ｷ309036ｦｸ803 026ｦ10307076 5, 0608070607 07060309 6ｦ90307086ｦ3060ｲ4 6 5 036ｦ107026ｦ06 76ｦ3, 0608070607 07060309 01 02070309076 56ｦ90ｸ9 090708020701 09 0ｲ4 6ｦ0 046 70903 066 5070606036 5 (Zhou, Yang, 29) ｨC 026 70107 040603070303 086ｦ0 6ｦ8036ｦｷ30909 01 07 09036ｦ90ｸ9 6 5 ｰ03070207 0706 ｱ 01076 509 06080ｲ40ｸ76 707 0604016ｦ36ｦ0 6ｦ06 76ｦ06ｦ0 09 6 5 076 76ｦ90903 02070309 09, 060707036 50306036 507, 01 0807 6ｦ9076 56ｦ90ｸ9 6 5 6 50709 04 0903020ｲ4 0ｹ4030ｸ8. ｨｨ066 5 070304 010402090ｸ806 03090107 0909 0509 0909, 056ｦ26 707 0507060307016ｦ0 09 09036ｦ20706 02090909, 6 5 070302090ｹ9 0703 08070ｹ903, 090303 6 5 6 70703 6 5 01090803090ｹ60109 08026ｦ90309 0ｸ8 (Eppinger et al., 27). ｨｨ07030708, 0ｹ307 090307 0709 0ｸ80209 08 0406036 509016ｦ309 06080ｷ30902090ｹ6010903 08026ｦ90309 0ｹ403076 707 03090107 6ｦ0 ppst 09 pfra. 5. 026ｦ1070ｹ16 507020ｸ90ｷ309070ｸ806 6 76ｦ0 046ｦ2090603090ｹ6010906 036ｦ06ｦ103 0807 6ｦ9076 56ｦ90ｲ4 08066 5 070304 01040209076 707 03090107 6ｦ0 6 5 6 50709 04 0903020ｲ4 0ｹ4030ｸ8: 0ｹ16 507020ｸ90ｷ3090709040ｸ90ｸ70906 03090107 080402 080706030807 0909 0703010ｸ8030706 086ｦ06ｦ909036ｦ706 06090603030ｸ8 ｰ6 70ｸ80904 (08090ｸ76 56ｦ903020ｸ806 036ｦ103)ｨC6 5040ｲ40ｲ4 06 6ｦ0 (08070ｹ06 56ｦ0)ｨC6 70ｸ80904 ｱ 6 5 046 7040ｸ9, 096ｦ3010403040ｸ9 060906030304 ｰ6 70ｸ80904 (02090302070309020709 0ｸ806 01070308020106)ｨC 0207036ｦ0 (08090ｸ76 56ｦ903020ｸ806 036ｦ103)ｨC6 70ｸ80904 ｱ, 0809 0ｹ1030703 03036ｦ0 0702090ｹ60109 08070ｷ306060ｸ8 03090107 6ｦ0 0803080209 08090ｷ30908096ｦ20ｸ8 090903090ｲ4; 6 5010307 6 50908040ｸ76 707 0703 076ｦ0 0ｸ802 6ｦ86ｦ06 502 0807 6 76ｦ0 090304 0303086ｦ304 0703 5 紊 (06070603070ｲ409 6 70204 0701076 707 030708076ｦ0 0604016ｦ0, 090306096 507 6ｦ903070809 0806090307020902076 707 08066 5 070304 01040209076 707 03090107 6ｦ0) 07 37 紊 (04060307060ｹ96 507 0ｹ1016 50902096ｦ207 06070603070ｲ409 0ｹ403076 707 03090107 6ｦ0 6 5 076 76ｦ90903 0308020701076 5076 707 0307090ｲ4096ｦ0). 6. 050403096 509 076 56ｦｲ4 0ｹ16 507020ｸ90ｷ3090ｲ4 03090107 6ｦ0 0ｹ4030ｸ8 08070307 09026ｦ0 0807 030901070ｹ16 507020ｸ90ｷ30907070304 08090ｷ3090804 6ｦ0 6ｦ803096 50706 6ｦ0 096ｦｷ30909. 7. 050ｷ36ｦ906 08 06036ｦ06 5020ｲ403 0ｹ10ｷ30901020708 090ｹ601040ｸ9 0ｹ16 507020ｸ90ｷ30907040ｸ9 0307 02, 6ｦ0 010703070706 03070807 6ｦ606036ｦ3096 56ｦ3 09 6ｦ06ｦ2090909076 56ｦ3 03076 709 6 5070807060ｸ8 0ｹ16 507020ｸ90ｷ309070706 03070909: 06070703070409 036ｦ107-09 030901070ｹ16 507020ｸ90ｷ30909, 6 76ｦ0 046ｦ20909036ｦ0 09 066ｦ2036ｦｷ309070909036ｦ0, 036ｦ1070ｹ16 507020ｸ90ｷ30909 09 020902076 7096ｦ0, 030707-09 080702090209020909; 01076 7030706, 01076 7030706, 016 56ｦ3076 50706 0ｹ16 507020ｸ90ｷ30909; 0702 6 70307 6ｦ3030909 (0307096ｦ90ｹ706 0ｹ16 507020ｸ90ｷ30909) 09 086ｦ0 6ｦ8036ｦｷ30906 6 5 036ｦ1070ｹ16 507020ｸ90ｷ309070ｸ803 0607 0ｸ803090ｲ403, 086ｦ06ｦ202020909030ｸ8, 01076 56 70ｷ30909 09 6 707030708026ｦ90909 0809 02070309076 56ｦ909 076 50ｸ803 0809090ｸ803 020703 09 036ｦ10607076 5 09 03076 709 046 709. 8. 0207 02 0807016ｦ90ｸ86 56ｦ3, 0ｹ307 6ｦ60603070309 07 0ｸ70903 6 507080706076 5 03070909 0ｹ16 507020ｸ90ｷ30909 6 5 010703070ｸ803 0602040ｹ06ｦｲ403 ｨ ｨ 0505ｨｪｨ ｨｹ ｨｺ0204. 05ｨｺｨ ｨｹ 03ｨ ｨｦｨｦ07ｨ ｨｮ050102ｨｹ 6ｧ86 29
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Achtman M., Zurth K., Morelli G. et al. Yersinia pestis, the cause of plague, is a recently emerged clone of Yersinia pseudotuberculosis // PNAS. 1999. V. 96. 6ｧ8 24. P. 14043ｨC14048. ｨ ｨ 0505ｨｪｨ ｨｹ ｨｺ0204. 05ｨｺｨ ｨｹ 03ｨ ｨｦｨｦ07ｨ ｨｮ050102ｨｹ 6ｧ8 6 29