Bull. Ibaraki Nat. Mus., (15): (2012) 13 ( ) Collection and Photographic Data on Dragonflies and Damselflies from Ibaraki Prefecture R
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1 Collection and Photographic Data on Dragonflies and Damselflies from Ibaraki Prefecture Ryo FUTAHASHI, Takehiko YAMANAKA, Yoshinobu UEMURA and Masaki HISAMATSU Reprinted from Bulletin of Ibaraki Nature Museum No.15 December, 2012
2 Bull. Ibaraki Nat. Mus., (15): (2012) 13 ( ) Collection and Photographic Data on Dragonflies and Damselflies from Ibaraki Prefecture Ryo FUTAHASHI, Takehiko YAMANAKA, Yoshinobu UEMURA and Masaki HISAMATSU (Accepted October 20, 2012) Abstract Ninety-one odonate species have so far been reported in Ibaraki Prefecture. Here we give a comprehensive list of dragonflies and damselflies collected from Ibaraki Prefecture based on the collections of Ibaraki Nature Museum and the authors private collections, which consist of 87 species and one hybrid species. We also mention the following four species which are not included in these collections: Stylurus oculatus, Sympetrum uniforme, Libellula angelina, and Tholymis tillarga. The former three species may have become extinct in Ibaraki Prefecture, and the last species seems to be a species migrating from a southern area. Key words: Odonata, dragonfly, damselfly, Ibaraki Prefecture. はじめに, 1970 (1971, 1976, 1979), (1972), (, 1998, 2001, 2004, 2007), (, 2008, 2009, 2010, 2011, 2012). (1997, 2011), 91, (2012a) Sympetrum fonscolombii (Selys, 1840),, Mnais pruinosa Selys, 1853, Asahina (1976) 1,, DNA (National Institute of Advanced Industrial Science and Technology, Higashi, Tsukuba, Ibaraki , Japan) (National Institute for Agro-Environmental Sciences, Kannon-dai, Tsukuba, Ibaraki , Japan) (Toyosato Museum of Entomology, 676 To-higashi, Tsukuba, Ibaraki , Japan) (Maeno Elementary School, 1367 Maeno, Tsukuba, Ibaraki , Japan).
3 14, (Hayashi et al., 2004;, 2004;, 2007)., Mnais costalis Selys, 1869,,,,, 91, Anax parthenope (Selys, 1839) Anax nigrofasciatus Oguma, 1915 ( ) 4 (, 1981;, 1998c;, 2001;, 2012a).,,,,,,,,,,,,,,,,,,, ( ), (2012). 8, ,,, Stylurus oculatus (Asahina, 1949), Sympetrum uniforme (Selys, 1883), Tholymis tillarga (Fabricius, 1798), Libellula angelina Selys, , 5 ( ), ( ), (), ( ), ( ).,. 採集 撮影記録 アオイトトンボ科 Lestidae Calvert, Sympecma paedisca (Brauer, 1877), 1, , ;, 1, ,. (, 2011), 10, (, 2011). ( 1). 図 1. (, , ). Fig. 1. A male of Sympecma paedisca (photographed by R. Futahashi on May 8, 2011 at Omonoi, Tsukuba City). 2. Indolestes peregrinus (Ris, 1916), 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2, , ; 1, , ; 1, , ;, 1, , ;, 1, , ; 1 1, , ; 1 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1, , ;, 1, , ;, 1, ,
4 15 ;, 1, , ; 4 1, , ;, 1 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, ,. ( 2). 図 2. (, , ). Fig. 2. A male-male tandem of Indolestes peregrinus (photographed by R. Futahashi on July 2, 2011 at Handa, Ishioka City). 3. Lestes sponsa (Hansemann, 1823), 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ; 2 1, , ;, 1, , ; 1, , ; 1, , ; 1 1, , ; 2 2, , ;, 2, , ; 1, , ;, 1, , ; 2 1, , ; 2, , ; 2 2, , ;, 1 1, , ; 1, , ;, 1, , ; 1 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1 1, ,. 4. Lestes temporalis Selys, 1883, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1 ( ), , ;, , ;, 1, , ;, 1, , ;, 2, , ;, 1, , ; 1, , ;, 1, , ; 1 1, , ;,, , ; 1 1, , ; 1 1, , ;, 1 1, , ;,, , ;, 1, , ;, 1, , ;, 1, , ;, 2 2, , ; 1, , ;,, , ; 1 ( ), , ;, , ;, 1 1, , ;, , ;, 1, , ; 1, , ;, 1, , ; 1, , ; 1, , ;, 1 1, , ; 1, , ;
5 16, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1 1, , ; 1, , ;, 1 1, ,. 5. Lestes japonicus Selys, 1883, 1 1, , ; 1 1, , ; 1 1, , ; 1 1, , ; 3 1, , ; 3 3, , ; 1, , ; 3 3, , ;, 3 1, ,. (2011),, (, 2009). カワトンボ科 Calopterygidae Selys, Mnais costalis Selys, 1869, 1, , ;, 1, , ;, 1, , ;, 2, , ; 1, , ; 1, , ; 1, , ; 1, , ; 1 1, , ;, , ;, 3 3, , ;, 3, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 2, , ;, 1, , ;, 1, , ;, 2 1, , ; 2, , ;, 2, , ;, 2 (1, 1 ), , ;,, , ;, 2, , ;, 4 1, , ; 1, , ; 1, , ;, 1, , ; 1, , ; 2, ,. 7. Calopteryx japonica Selys, 1869, 1, , ;, 1 1, , ;, 1 1, , ; 1, , ;, 2, , ; 2 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2, ,.,. 8. Calopteryx cornelia Selys, 1853, 1, , ;, 1 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1, , ;, 2 2, , ;, 1 1, , ; 1, , ;, 1, ,. 9. Atrocalopteryx atrata (Selys, 1853), 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ; 1, , ;, 1, , ;, 1 1, , ;, 1, , ;
6 17, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1 1, , ;,, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;,, , ;, 1 1, , ; 2, , ; 1 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2 2, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ;, 1, , ; 1, , ;, 1, , ;, 1 1, , ;, 1, , ; 1, , ;, 3, ,. モノサシトンボ科 Platycnemididae Tillyard et Fraser, Copera annulata (Selys, 1863), 1, , ;, 1 1, , ;, 1, , ; 1, , ;, 1, , ;, 2 2, , ;, 1, , ;, 1, , ;, 1, , ;,, , ;, 1 1, , ;,, ,., 11. Copera tokyoensis Asahina, 1948, 1 1, , ; 1 1, , ; 1 2, , ;, 1, , ; 1 2, , ; 3 2, , ;, , ; 2 1, , ;, 1, , ; 1 1, , ;, 3, , ; 1 1, , ; 1, , ; 1 1, , ; 1, , ; 1 1, , ;, , ;, , ; 1 1, , ; 1 1, , ;, 1, , ; 1, ,.,, 2010 (, 2011)., (, 2000), DNA (, 2011). イトトンボ科 Coenagrionidae Kirby, Ceriagrion melanurum Selys, 1876, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 4 1, , ;, 1 1, , ; 3, , ;, 2, , ; 1 1, , ;,, , ; 2, , ;, , ;, , ;, 1, , ; 1, , ; 1, , ;, 1, ,.
7 18,. 13. Ceriagrion nipponicum Asahina, 1967, 1, ,. (2008). (, 2011), 14. Coenagrion terue (Asahina, 1949), 2, , ;, 2 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1 1, , ; 1 1, , ;, 2, , ; 1, , ;, 1, , ;, 2 1, , ;, 1, , ;, 1, ,., (, 2011).,,. 15. Paracercion plagiosum (Needham, 1930), 3 1, , ;, 1, , ; 1 1, , ; 1, , ; 1 1, , ; 1, , ; 1 2, , ; 1, , ; 3, , ; 1, ,.,, 2008 (, 2011),. 16. Paracercion calamorum (Ris, 1916), 1, , ;, 1, , ; 1, , ;, 2, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ; 1, , ; 1, , ; 2, , ; 2, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 5, , ; 2, , ; 1 1, , ; 1, , ;, , ;, 1, , ; 2, , ;, 1, , ;, 1, , ; 1 1, ,. 17. Paracercion hieroglyphicum (Brauer, 1865), 1 1, , ;, 1, , ;, 2, , ;, 1, , ;, 2 1, ,.,, 18. Paracercion sieboldii (Selys, 1876),, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, ,
8 19 ; 1 1, , ; 2, , ;,, , ;, 1, , ; 1, , ;, 2, , ;, 1, , ;, 1, , ;, 2, , ;, 1, , ; 1, , ;, 3, , ; 1, , ;, 1, , ;, 2 1, , ;, 2 1, , ;, 1, , ;, 1, , ; 1 2, ,.,, , ; 1, , ; 2 2, , ;, 1 1, , ; 1 1, , ;, 1, , ;, 1 1, ,. ( 3)., 19. Paracercion melanotum (Selys, 1876), 2 3, , ; 1, , ;, 2 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 3, , ; 1, ,., (, 2011),. 20. Mortonagrion selenion (Ris, 1916), 2 1, , ;, 1 1, , ;, 1, ,.,,. 21. Mortonagrion hirosei Asahina, 1972, 1 1, , ; 1 1 図 3. (, , ). Fig. 3. Mating of Mortonagrion hirosei (photographed by R. Futahashi on July 10, 2010 at Kamiishizaki, Ibaraki Town). 22. Ischnura senegalensis (Rambur, 1842), 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1 1, , ;, 2 ( 1 ), , ;, 1 1, , ;, 1 1, , ;, 1, , ;, , ;, , ;, 1, , ;, 1, ,
9 20 ;, 1 1, , ; 1, , ; 1, , ; 3 1, , ; 1, , ; 1 1, , ; 1, , ; 1 1, , ; 1, , ; 1 1, , ; 1, , ;, 2 3, , ;, 1, , ; 3 2, , ;, 1 1, ,. 23. Ischnura asiatica Brauer, 1865, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;,, , ;, 1, , ; 1, , ; 1, , ;, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ;, 1, , ; 1, , ;,, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ; 1, , ; 1, , ; 1 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 2, , ; 3 2, , ; 1, , ; 1, , ; 1, , ; 1 1, , ; 1 1, , ; 2 1, , ; 1 1, , ;, 2, , ; 1 1, , ; 1 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, ,. ムカシトンボ科 Epiophlebiidae Muttkowski, Epiophlebia superstes (Selys, 1889), 1 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ;, 2, , ; 2 2, , ;, 2 1, , ;, 1, , ; 1, , ;,, , ;, 1 1, , ; 2 1, ,.,,,,,,,,,, 10 (, 2011). ヤンマ科 Aeshnidae Rambur, Sarasaeschna pryeri (Martin, 1909), 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, ,
10 21 ; 1 1, , ;,, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1 1, , ;, 1 1, , ; 1 1, , ; 1, , ; 1 1, , ;, 1, , ;, 1, , 26. Boyeria maclachlani (Selys, 1883), 1, ,. (, 2011),. 27. Planaeschna milnei (Selys, 1883), 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;,, , ;,, , ;, 2 2, , ; 1, , ;, 1 1, , ; 3 1, ,. (, 1, , ; 1, , ). 28. Aeschnophlebia longistigma Selys, 1883, 1 1, , ;, 1, , ;, 1, , ;,, , ;, ( ), , ;, , ; 1, , ;, 1, , ;,, , ;, 1, , ;, 1, , ; 1 2, , ; 1, , ;, , ; 1, , ;, 2, , ; 1, , ; 1, , ;,, , ;, 1, , ; 1, , ; 1, , ; 2 2, , ; 1, , ; 1, , ; 1 1, , ; 1, , ; 1, , ; 1, , ; 1 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, ,., 29. Aeschnophlebia anisoptera Selys, 1883, 1, , ; 1, , ; 1 1, , ;, 1, ,.,,,,, (, 2011). 30. Gynacantha japonica Bartenef, 1909, 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 3, , ;, 1, , ;, 1, ,., (, 2011). (, 2012),
11 Anaciaeschna martini (Selys, 1897), 1, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ; 1, , ;, 1, , ; 1 1, , ;, 1, , ; 3, , ; 3, , ; 2, , ;, 1, ,.,. 7 ( 4)., 1, , ;, 1, ,. 33. Aeshna mixta Latreille, 1805, 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 2, , ; 1, , ; 1, ,.,. 34. Aeshna crenata Hagen, 1856, 1, , ;, 1, , ;,, , ;, 2, , ;, 1 2, , ; 1, , ;, 1, , ; 1 1, , ;, 1, , ;,, ,. 図 4. (, , ). Fig. 4. A male of Anaciaeschna martini (photographed by R. Futahashi on July 24, 2009 at Higashi, Tsukuba City). 32. Polycanthagyna melanictera (Selys, 1883), ( ), , ;, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2, , ;, 1, , ; 1, , ;, ( ), , ;, , ;, 1 2, , ; 1, , ; 1, , ;, 1, , ; 1, , ; Aeshna juncea (Linnaeus, 1758), 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, ,. 36. Anax parthenope (Selys, 1839), 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, ,
12 23 ; 1, , ; 1, , ; 1, , ;, 1, , ; 2, , ; 1, , ; 1 1, , ; 1, , ;,, , ; 1, , ;, 1, , ; 2, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ; 2, , ;, 1, , ;, 1, , ;, 1, , ;, 1, ,. 37. Anax nigrofasciatus Oguma, 1915, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ;,, , ;, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, ,. 38. Anax guttatus (Burmeister, 1839), 2, , ;, 1, ,., 1998,, (, 1998a, 1998b;, 1998). サナエトンボ科 Gomphidae Rambur, Sinictinogomphus clavatus (Fabricius, 1775), 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 2, , Yamaguchi, S & Aoki, T;, 1 1, , ;,, , ; 1, , ;, 1, , ;,, , ;, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 2, , ; 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2 2, , ; 1, , ; 1 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;,
13 24 5 exs, , ; 7 exs, , ; 8 exs, , ; 1, , ; 3 exs, , ; 27 exs, , ; 1 ex, , ;, 1, ,., 40. Sieboldius albardae Selys, 1886, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, ,. 41. Onychogomphus viridicostus (Oguma, 1926), 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 2, , ;, 1 1, , ;, 1, , ;, 1, ,. 42. Nihonogomphus viridis Oguma, 1926, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ; 1 1, , ; 1, , ;, 1, , ; 1, ,. 43. Davidius fujiama Fraser, 1936, 2, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ;, 1, , ; 1 6, , ;,, , ;, 1, , ; 2 1, , ;, 1, , ;, 1, , ;, 1, ,. 44. Davidius nanus (Selys, 1869), 2, , ;, 1, , ;, 1, , ;, 1, , ; 2, , ; 1, , ;, 1, , ;, 2, , ; 2, , ;, 1, , ;, 1, , ; 2, , ;, 1, ,. 45. Davidius moiwanus (Matsumura et Okumura in Okumura, 1935), 1, , ; 1, , ;, 1, ,. 46. Lanthus fujiacus (Fraser, 1936), 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;,, , ;, 1, , ;, 1 1, , ;, 1, ,.
14 Sinogomphus flavolimbatus (Matsumura in Oguma, 1926), 1, , ;, 1, , ;, 1, ,. 48. Stylogomphus suzukii (Matsumura in Oguma, 1926), 1 1, , ;, 1, , ;,, , ;,, ,. 49. Trigomphus melampus (Selys, 1869), 2, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;,, , ;, 1, , ; 1, , ;, 1, , ; 1, , ; 2, , ; 2, , ; 1, , ; 1, , ; 1, ,. ; 1, , ;, , ; 2 2, , ; 3 3, , ;, 1 1, , ; 1 1, , ; 1 1, , ;, 1, , ;, 1, , ;,, , ;, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ; 1, , ;, 1 ex, , ;, 2, , ;, 1, , ;, 1, , ;, 1, ,.,, (, 2011). 52. Stylurus oculatus (Asahina, 1949) ,,,, (, 2012). 50. Anisogomphus maacki (Selys, 1872), 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;,, ,. 51. Stylurus nagoyanus (Asahina, 1951), 2 2, , ;, 1 1, , ;, 1, , ; 1 2, , 53. Shaogomphus postocularis (Selys, 1869), 1 1, , ; 1, , ; 1 1, , ;, 1, , ;, 1, , ; 1, , ;, 2 2, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2, , ;, 1, , ;
15 26, 1, , ; 1, , ; 1, , ;, 1, , ; 1, , ; 1, , ;, 2, , ; 1, , ;, 2, ,., (, 2011). 54. Asiagomphus pryeri (Selys, 1883), 1, , ;, 1, , ;, 2, , ;, 2, ,.,. 55. Asiagomphus melaenops (Selys, 1854), 2 exs, , ;, 1, , ;, 1, , ;, 1, , ;, 1 2, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1 1, , ; 1 2, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;,, , ;, 1 1, , ;, 1, , ; 1, ,. ムカシヤンマ科 Petaluridae Needham, Tanypteryx pryeri (Selys, 1889), 1, , ;, 1, , ;, 1, , ; 1 1, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 2, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 1 1, , ; 1, , ; 1 1, , ; 1, , ; 1, , ;, 1, , ;, 1, ,. オニヤンマ科 Cordulegastridae Banks, Anotogaster sieboldii (Selys, 1854), 1, , ;, 1 ex, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;,, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, , ;, 1, , ; 1, , ; 1, , ;, 1 1, , ;,, , ;, 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1 1, , ; 1 1, , ; 1, , ;, 1, , ;, 2, , ;, 3, , ; 1, , ;, 1, , ;, 1
16 27, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ;, 3 exs, , ; 2 exs, , ; 2 exs, , ; 1, , ; 1, , ;, 1, , ;, 1, ,. エゾトンボ科 Corduliidae Selys, Epitheca marginata (Selys, 1883), 1, , ;, 1, , ; 1, , ; 1, , ; 3, , ;, 1, , ; 1, , ; 1, ,.,. 59. Somatochlora uchidai Förster, 1909, 1, , ;, 1, , ;, 1, ,. (, 2011),., (, 2010). ヤマトンボ科 Macromiidae Tillyard, Epophthalmia elegans (Brauer, 1865), 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;,, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ; 1, , ; 1, , Obata, K; 2 exs, , ; 1 ex, , ;, 1, , ; 1, , ;, 1, ,. 60. Somatochlora viridiaenea (Uhler, 1858), 1 1, , ;, 1 1, , ;, 1, , ; 1 1, , ; 2, , ;, 1, , ; 1 1, , ; 2, , ; 3, ,. 61. Somatochlora clavata Oguma, 1913, 1, , ; 1, , ;, 1, ,. 63. Macromia daimoji Okumura, 1949, 1, , ;, 1, , ;, 1, , ;, 1, , ; 2, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1 1, ,.,.
17 Macromia amphigena Selys, 1871, 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1, , ;, 1, , ; 1, , ;, 2, , ;, 1, , ;, 1, , ;,, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 2, , ;, 2 exs, , ;, 1, , ;,, , ;, 1, , ;, 1 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, ,. トンボ科 Libellulidae Selys, Rhyothemis fuliginosa Selys, 1883, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 3 2, , ; 1, , ; 1, , ; 1, , ; 1 1, , ;, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 1 1, , ; 1 1, , ; 1 2, , ; 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1 ( ), , ; 3 3, , ; 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ; 3 3, , ; 3 3, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 2, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, ,. 66. Sympetrum fonscolombii (Selys, 1840), 1, ,. (2012a), 2011 ( 5).,,., 図 5. (, , ). Fig. 5. A male of Sympetrum fonscolombii (collected by R. Futahashi on August 22, 2011 at To-higashi, Tsukuba City).
18 Sympetrum darwinianum (Selys, 1883), 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;, 2, , ;, 1, , ;, 1 1, , ; 1 1 ( ), , ;, 1, , ;, 1, , ;, 1, , ; 2 exs, , ;, 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1 2, , ;, 1, , ;, 1 1, , ; 1, , ; 1, , ; 1 1, , ; 1 1, , ; 3 3, , ; 1 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ; 2 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ; 2 3, , ; 1, , ;, 1, , ;, 1, , Kikuchi, S;, 1, , ;, 1, , ; 1, , ; 2, , ;, 2 1, ,.,,. (, 1, , ). 68. Sympetrum risi Bartenef, 1914, 1, , ;, 1, ,. (, 2011),. 69. Sympetrum infuscatum (Selys, 1883), 1, , ;, 1, , ;, 1, , ;, 2, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1 1, , ; 1 1, , ; 1 1, , ;, 1, , ; 1 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2 2, , ; 1, , ; 1 1, , ; 1, , ;, 1, , ;, 1 ( ), , ; 3 3, , ;, 2, , ;,, , ;, 1, , ;, 1 1, , ;, 1, ,
19 30 ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1, , ;, 1, , ; 1, , Kikuchi, S;, 1, , Horiguchi, T; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 2, ,.,,. (, 1, , ; 1, , ). 70. Sympetrum frequens (Selys, 1883), 1, , ;, 1, , ;, 1, , ;, 1, , ; 2, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;,, , ;, , ; 2 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 3 3, , ; 1 1, , ; 1 4, , ; 1 1, , ; 1, , ; 2 2, , ; 1 1, , ; 1 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1 1, , ;, 1, , ;, 1, , ;, 1 4, , Naka,T&Watanabe,K;, 1, , ; 1, , ; 1 1, , ; 1, , ; 1 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 2, , ; 1, , ;,, , ;, 1, , ;, 1, , ; 1, , ;, 2, , ; 1, , ;, 1, ,.,,. 71. Sympetrum baccha Selys, 1884, 1, , ;,1, , ;, 1, , ;, 1 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, ,.,. 72. Sympetrum parvulum (Bartenef, 1912), 1, , ; 2 1, , ; 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ;, 2, , ;, 1, ,., 73. Sympetrum eroticum (Selys, 1883), 1, , ;
20 31, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1, , ; 1, , ;, 1, , ;, 2, , ;, 1, , ;, 2 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1 ( 1 ), , ;, 1, , ; 1, , ; 1, , ;, 1, , ;,, , ;,, , ;, 1, , ;, 10 exs, , ; 2, , ; 1 1, ,. 74. Sympetrum kunckeli (Selys, 1884), 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 2, , ; 1, , ; 1, , ; 1, , ; 1, , ; 1, , ;, 1 1, , ;, 1 1, , ; 1 1, , ; 1 1, , ; 2 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1 1, , ; 1 1, , ; 1, , ; 1, , ;, 1, , ;, 1 1, , ;, 1 1, , ;, 1 1, , ; 3 1, , ; 2 2, , ; 1 1, , ; 1 1, , ; 1, , ;, 1, , ; 1, , Shibayama, H;, 1, , ;, 1, , ; 1 1, , ;, 1, , ; 1, , ;, 1, ,., 75. Sympetrum pedemontanum (Allioni, 1766), 1, , ;, 1, , ;, 1 1, , ; 1 2, , ;, 1, , ;, 1, , ; 1 1, , ;, 1, ,.,. 76. Sympetrum speciosum Oguma, 1915, 1, ,.,. 77. Sympetrum croceolum (Selys, 1883), 1, ,.,, (, 2011). 78. Sympetrum uniforme (Selys, 1883), (, 2011).
21 Tramea virginia (Rambur, 1842), 1, , ; 1, ,. ( 6)., (, 1980;, 2010). 図 6. (, , ). Fig. 6. A female of Tramea virginia (collected by H. Naka on May 29, 2004 at Kannondai, Tsukuba City). 80. Pseudothemis zonata (Burmeister, 1839), 1, , ;, 1, , ; 1, , ;, 1, , ;, 2 1, , ;, 1, , ;, 2, , ;,, , ;, 1 ( ), , ;, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ; 1 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ; 2, , ; 1, , ;, 1, , ; 1 1, , ; 1, , ; 1, , ; 1, , ; 1, , ;, 2 1, , ; 2 1, , ;, 3 1, , ;, 2, , ; 1, , ;, 2 1, , ; 1 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, ,. 81. Deielia phaon (Selys, 1883), 1 ( ), , ; 1, , ; 1 ( ), , ;, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1 ( ), , ;, 1 ( ), , ;, 1, , ; 1, , ; 1, , ;, 1 ( ), , ;, 2, , ;, 1, , ;, , ;,, , ; 2 exs, , ; 1, , ; 1, , ; 1 ( ), , ;, , ;, 1, , ; 1 ( ), , ;, 1, , ;, 1 ( ), , ; 1, , ; 1, , ; 1 ( ), , ;, 1, , ;, 1, , ;
22 33, 1 ( ), , ; 1, , ;, 1 ( ), , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1 ( ), , ;, 1 ( ), , ;, 1, , ; 1, , ;, 3 2, , ; 1, , ; 1, , ; 1 1, , ; 1, , ; 1, , ; 1, , ; 1 1, , ; 1 1, , ; 1, , ; 1 ( ), , ; 1, , ; 2 2 ( 1, 1 ), , ; 1 1, , ;, 1, , ;, 1 1, , ;, 1 ( ), , ; 1, , ; 4 exs, , ; 4 exs, , ; 6 exs, , ; 10 exs, , ; 1 ex, , ; 3 exs, , ; 11 exs, , ; 11 exs, , ; 5 exs, , ; 5 exs, , ;, 2, , ;, 1, , ; 1 1 ( ), , ;, 1, ,. ( ). 82. Tholymis tillarga (Fabricius, 1798), (, 1995). 83. Nannophya pygmaea Rambur, 1842, 1, , ; 1, , ;, 2, , ;, 2 1, , ;, 1, , ;, 1, , ; 1, , ;, 1 1, , ; 2 1, , ;, 1, , Fujimura, T;, 3, , ; 1 1, , ; 1, , ; 1, , ;, 1, , ;, 2 1, , ;, 1, ,., (1 1, , ; 5, , ), 84. Crocothemis servilia (Drury, 1770), 1, , ;, 2, , ;, 2 1, , ;, 1 1, , ; 1, , ; 1, , ;, 1, , ;,, , ; 1, , ; 1, , ;, , ;, 3 3, , ; 3, , ; 3, , ; 3 2, , ; 2 2, , ; 4 2, , ; 1, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1 1, , ; 3 3, , ; 1 1, , ;, 1 2, , ; 1, , ; 1, , ; 1 1, , ;, 2, , ; 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, ,
23 Pantala flavescens (Fabricius, 1798), 1, , ;, 1 1, , ;, 1 1, , ;, 1, , ;, 2, , ; 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ; 1, , ; 1, , ; 1, , ;,, , ;, 1, , ; 1, , ; 1 2, , ; 1, , ;, 1, , ; 1 2, , ;, 1, , ;, 1 1, , ; 1 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 3 exs, , ;, 1, , ;, 1, ,.,, 86. Lyriothemis pachygastra (Selys, 1878), 1 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 3 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1 1, , ;, 1, , ;, 1, , ;, 1 1, , ; 1 1, , ;, 1, , ; 1, , ;, 1 1, , ;, 1, , ;, 2 1, , ;, 1, , ;, 1, , ;, 1 1, ,. ( ) A 3 ( 7). 図 7. 3 (, , ). Fig. 7. A rare case of coupling by two males and a female of Lyriothemis pachygastra (photographed by R. Futahashi on May 19, 2011 at Ajiki, Tsukuba City). 87. Orthetrum albistylum (Selys, 1848), 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1 1, , ;, 1, , ;, 1, , ;,, , ;, , ;, , ;, 1, , ;,, , ;,, , ;, 1 1, , ; 1, , ;,, , ;, 1, , ; 1 1, , ; 5 5, , ; 1, , ; 1 1, , ; 1, , ;, 1, , ; 1, , ;
24 35, 1, , ; 1, , ; 1, , ;, 1, , ; 1, , ; 1, , ;, 1, , ;, 1 1, , ; 1 1, , ; 1 1, , ; 1, , ; 1, , ; 1, , ; 2 2, , ; 3, , ;, 1 1, , ;, 1, , ;, 4 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ; 1, , ; 2 1, , ;, 1 1, , ; 1, , ;, 1 ex, , ; 1 ex, , ;, 1, , ; 1, , ;, 2, , ; 1 ex, , ; 1, , ; 2 exs, , ; 3 exs, , ;, 1, , ;, 1, , ; 1 1, , ;, 1, ,. 88. Orthetrum japonicum (Uhler, 1858), 1, , ; 2, , ; 1, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;,, , ; 1, , ;, , ;,, , ;, 1, , ;, 1, , ;, 1, , ;, 5, , ;, 1, , ; 1, , ; 1 1, , ; 1, , ; 1, , ; 1, , ; 1 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, ,. 89. Orthetrum melania (Selys, 1883),, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;,, , ;, 1, , ;, 1, , ;, 1 1, , ; 1, , ;, 1, , ;,
25 36 1, , ; 1, , ; 1, , ; 1, , ; 1, , ;, 1, , ;, 1, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1, , ;, 1 1, , ;, 1 ex, 1991,0626, ; 1, , ; 1 ex, , ;, 1, , ;, 1, ,. 図 8. (, , ). Fig. 8. A male of interspecific hybrid between Anax parthenope and Anax nigrofasciatus (photographed by R. Futahashi on July 9, 2011 at To-higashi, Tsukuba City). 90. Libellula quadrimaculata Linnaeus, 1758, 2, , ; 1, , ;, 1, , ;, 1, , ;, 1, , ; 1, , ;, 2, , ; 2, , ; 1, , ; 1 1, , ; 1, , ;, 1, , ; 1, ,.,. 91. Libellula angelina Selys, 1883,, (, 2011). 1. Anax parthenope Anax nigrofasciatus, 1, ,. (2012a).,, 1 (, 1981;, 1998c;, 2001). ( 8), DNA (, 2012a). 考察 91,,, 4,,, 3, 87,,,, 4,,,, 3,,. 80,. (2011), 60,.,,,,,, 2,,,,,,,,,
26 37,,,,,.,,,.,.,, 2011, (, 2012). (, 2012),,, (, 2012).,,, 4,,,,,,,,, 8,,,,,,,,, 3,,, (, 2009;, 2011;, 2012b)., 3., (2011),,, (, 1998;, 2004;, 2005;, 2007;, 2009;, 2010).,,. 引用文献 , 13: Asahina, S A revisional study of the genus Mnais (Odonata, Calopterygidae), VIII. A proposed taxonomy of Japanese Mnais. Tombo, 19(1-4): ( 2 ). Tombo, 24(1-4): (2011 ) pp., Tombo, 15(1-4): , 42(9): DNA (1). Tombo, 53: a. Aeschna, (48): b., 47(8): , 17: Hayashi, F., S. Dobata and R. Futahashi Macro- and microscale distribution patterns of two closely related Japanese Mnais species inferred from nuclear ribosomal DNA, ITS sequences and morphology (Zygoptera: Odonata). Odonatologica, 33: , (1). Aeschna, (41): , (40): , (67): ( 2 )., 14(6): , (19): pp., pp , pp. 2-15,.
27 ( ). 1. pp , ( ). 2. pp , ( ). 3. pp , ( ). 4. pp , , (39): , 77(1): pp., ( ). 216 pp., pp., , 38: a., (22): b., (22): c., (22): , (39): , (50): , (25): ( 10 ). 283 pp , : ( : )., (11): , 2000., 35(11): , (22): 30-31, pp.7-8, pp.7-8, pp.11-13, pp.11-12, pp.19-20, pp.,. ( 要旨 ) 二橋亮 山中武彦 植村好延 久松正樹. 茨城県におけるトンボ目の採集 撮影記録. 茨城県自然博物館研究報告第 15 号 (2012)pp , 91, 87 1.,, 4., Stylurus oculatus, Sympetrum uniforme, Libellula angelina Tholymis tillarga 4, 3., ( キーワード ):,,,.
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