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1 ISSN Bulletin of Ibaraki Nature Museum No. 14 November, 2011 Bando, Ibaraki, Japan

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3 Bull. Ibaraki Nat. Mus.,14: 1-13, pls Outcrops of the Unconformity between the Cambrian and Carboniferous Formations with 150 My Gap in the Hitachi Paleozoic Terrane Michio TAGIRI and Wataru KOIKE Accepted November 13, 2011 Abstract The unconformity between the Cambrian Akazawa Formation and the Carboniferous Daioin Formation is traced from the northeast to the southwest in the Hitachi Paleozoic Terrane. We newly describe the geological features of the unconformity in four outcrops. Basal conglomerates occur above the unconformity in the northeastern part, but not in the southwestern part. In the latter part, blocks of volcanic rock of the Akazawa Formation and pebbles of meta-granitoids are intercalated in the Daioin Formation. Key words: Hitachi Paleozoic Terrane, Cambrian, Carboniferous, unconformity, basal conglomerate, Great Hiatus. 2010; Tagiri et al., Sakashima et al., A 1B SHRIMP 500 MaTagiri et al., C Tagiri, * Hitachi City Museum, Miyata-cho, Hitachi, Ibaraki , Japan. ** Ibaraki Nature Museum, 700 Osaki, Bando, Ibaraki , Japan.

4 2 1 4 : N40W 55 E 2 3 m20 m N50W30N : 3A N40E m 3B : 3C N40E 70E N35E 90 Fig. 1. Geological map of the Hitachi Paleozoic Terrane and geologic ages. Fig. 2. Stratigraphical divisions of the Hitachi Paleozoic Terrane.

5 1.5 3 :: : : Fig. 3. Localities of four unconformity outcrops. 1: Daioin, 2: Fudodaki Fall of the Miyata River, 3: Takidaira, 4: Upper reaches of the Ayu River. 3D : 4A EW 90 4B : N30E30E N50E35 S 1 Fig. 4. Lithofacies distribution near the unconformity outcrop between the Cambrian Akazawa Formation and the Carboniferous Daioin Formation in Daioin. 500 Ma 2010; Tagiri et al., ; Minato, Ma 1 5 Great Hiatus, 1979; Kim,

6 4 Ehiro, 2001; Ishiwatari and Tsujimori, 2003;, 2004;, 2010; Isozaki et al., Tagiri et al.2010, SHRIMP D.J. Dunkley 2010 Ehiro, M Origins and drift histories of some microcontinents distributed in the eastern margin of Asian Continent. Earth Sci.Chikyu Kagaku, : : Ishiwatari, A. and T. Tsujimori Paleozoic ophiolites and blue schists in Japan and Russian Primorye in the tectonic framework of East Asia: A synthesis. Island Arc, : Isozaki, Y., K. Aoki, T. Nakamura and S. Yanai New insight into a subduction-related orogen: A reappraisal of the geotectonic framework and evolution of the Japanese Islands. Gondwana Res., : Kim, O.J Tectonic Evolution. In: Lee, D.S.ed., Geology of Korea, Chapter 11, pp , Geological Society of Korea, Kyohaku-Sa, Seoul Ma : pp ,. Minato, M Japanese Carboniferous and Permian corals. J. Fac. Sci., Hokkaido Univ. Series 4, Geology and Mineralogy, : Sakashima, T., K. Terada, T. Takeshita and Y. Sano Large-scale displacement along the Median Tectonic Line, Japan: evidence from SHRIMP zircon U-Pb dating of granites and gneisses from the South Kitakami and paleo-ryoke belts. J. Asian Earth Sci., : Tagiri, M Metamorphism of Paleozoic rocks in the Hitachi district, southern Abukuma Plateau, Japan. Sci. Rep. Tohoku University, 1: D.J. Dunkley2010 SHRIMP : Tagiri, M., D.J. Dunkley, T. Adachi and Y. Hiroi Great hiatus in the Cambrian Hitachi metamorphic terrane comparable to the North China Craton. 117 O-60p. 65 Tagiri, M., D.J. Dunkley, T. Adachi, Y. Hiroi and C.M. Fanning SHRIMP dating of magmatism in the Hitachi metamorphic terrane, Abukuma belt, Japan: Evidence for a Cambrian island arc. Island Arc, : : :

7 4 Plates and Explanations 4 plates

8 A: B: C: Outcrop of the unconformity at the upper stream of Fudodaki Fall, Torentsu river. A: Meta-conglomerate and metagranite contact with the boundary of the unconformity, B: Boulder of meta-granite in the meta-conglomerate, C: Biotite schistmatrixin the meta-conglomerate.

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10 Outcrop of the unconformity at a steep slope of the hillside in DaioinOutcrop 1.

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12 A: B: C: D: Outcrops of the unconformity at Fudodaki Fall of the Miyata RiverOutcrop 2and TakidairaOutcrop 3. A: the boundary of the unconformity at the side of Fudodaki FallOutcrop 2, B: Granitic pebbles included in phyllite in Outcrop 2, C: the boundary of the unconformity at the road-side of TakidairaOutcrop 3, D: Granules included in phyllite in Outcrop 3.

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14 A: B: Outcrop of the unconformity at the upper reaches of the Ayu RiverOutcrop 4. A: Loosely curved boundary of the unconformity between amphiboliteakazawa F.and phylliteoaioin F., B: An amphibolite block embedded in phyllite.

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16 Bull. Ibaraki Nat. Mus.,14: Species Composition of Wild Bees at the Okadaira Shell Mound, Ibaraki Prefecture, Central Japan Masaki HISAMATSU Accepted May 14, 2011 Abstract The species composition of wild bees was surveyed from March to November in 2007 at the Okadaira Shell Mound in Miho, Ibaraki Prefecture. A total of 1464 individuals belonging to 52 species in five families were collected. The most predominant family in the number of species was Halictidae15 spp., 271 individuals, followed by Apidae14 spp., 478 individuals. With respect to the number of individuals, Apidae was the largest family and Colletidaethree spp., 302 individualswas the second-largest one. The largest number of individuals207was collected for Colletes patellatus, and the second largest172 was for Apis cerana japonicaapidae. Ten of the 52 species were regarded as dominant species. The large number of individuals in Co. patellatus and Co. collaris might have resulted from an abundance of composite plants blooming in autumn, on which these bees foraged for pollen. The results were compared with those of previous studies performed in six areas in Ibaraki Prefecture. The number of species and the values for species evenness at the Okadaira site were ranked high among areas that have so far been studied in Ibaraki Prefecture. This area seems to have a richer bee fauna than the other studied areas. Lasioglossum primavera, Amegilla florea florea and Thyreus decorus, which were also regarded as dominant species in the Okadaira area, have not been recorded in large numbers in other areas of Ibaraki Prefecture. The species composition of wild bees at the Okadaira site is similar to Sugao and Mito, which are cultivated and/or urbanized. Key words: Apiformes, Hymenoptera, Bee community, Similarity coefficient, Okadaira Shell Mound, Ibaraki Prefecture. Hymenoptera Apoidea Apiformes Hisamatsu and Yamane, 2006;, 2010, , 1992; 2010, 2011; 1997; Matsumura et al., 1974; 1980; Sakagami and Fukuda, Hisamatsu2010 * Ibaraki Nature Museum, 700 Osaki, Bando, Ibaraki , Japan

17 Hisamatsu and Yamane ha m ha 2.6 km 1-B Veronica persica Poir. Lamium purpureum L. Taraxacum officinale Weber ex F. H. Wigg. Corydalis incisathunb.pers. Erigeron philadelphicus L. Vicia sativa L. subsp. nigral.ehrh. Trifolium repens L.Pollia japonica Thunb.Leonurus japonicus Houtt. Oxalis corniculata L. Solidago canadensis var. scabra L A: 6 B: Fig. 1. A: Location of the Okadaira Shell Mound and six other points surveyed previously in Ibaraki Prefecture, B: magnified map showing the surveyed area. The double line indicates the survey route.

18 : 0012 : : m2008: m 1985: 33 m 1992: m 2011: mhisamatsu 2010: mhisamatsu and Yamane A N ns n nnn N N 3 1 S SNSimpson 1 D Simpson 1/D4 Krebs, CC Krebs1999: 2n i1 n i2 C 2 N N 2 n i1n i1 N 1N 1 =D n i1 n i2 1 2 i N1= n i1 N2= n i21 2 : LasioglossumEvylaeus hirashimae Ebmer et SakagamiLasioglossumEvylaeus sp.7ebmer and Sakagami, 1985a, Lasioglossum Evylaeus allodalum Ebmer et Sakagami LasioglossumEvylaeussp.9Ebmer and Sakagami, 1985b LasoglossumEvylaeuslongifacies Sakagami et Tadauchi LasioglossumEvylaeussp.5 LasioglossumEvylaeuszunaga Sakagami et Tadauchi LasioglossumEvylaeussp.4 LasioglossumEvylaeuspumilum Sakagami et Tadauchi=LasioglossumEvylaeussp.3Sakagami and Tadauchi, 1995a LasioglossumLasioglossumnipponicola Sakagami et Tadauchi LasioglossumLasioglossumsp.3 LasioglossumLasioglossum ebmerianum Sakagami et Tadauchi Lasioglossum Lasioglossumsp.2Sakagami and Tadauchi, 1995 b LasioglossumEvylaeus pallilomumstrand LasioglossumEvylaeussp.18 Takahashi and Sakagami, 1993 Nomada okubira TsunekiMitai and Tadauchi, 2006Coelioxys

19 18 Boreocoelioxyshiroba NagaseNagase, 2006 Nomada sheppardanacoelioxys acuminata ,464 1 Halictidae 15 Apidae 13 Megachilidae 11 Andrenidae 10 Colletidae ColletesColletespatellatus Pérez % Apis cerana japonica Radoszkowski % 10% % 95% 1.9% Megachile tsurugensis Cockerell1147.8% AndrenaSimandrenayamato Tadauchi et Hirashima976.6% ColletesColletescollaris Dours94 6.4%Eucera nipponensis Pérez 745.1% LasioglossumLasioglossummutilumVachal54 3.7% Lasioglossum Lasioglossumprimavera Sakagami et Maeta50 3.4%3-A Amegilla florea floreasmith50 3.4% 3-B Thyreus decorussmith42 2.9%3-C Fig. 2. Relative and cumulative frequencies of 12 species collected at the Okadaira Shell Mound. Ten species, for which the lower end of the 95% confidence limit exceeded 1.9% (an average relative frequency), were regarded as dominant species.

20 19 Table 1. Species and number of wild bees collected at the Okadaira Shell Mound. Family and species name Japanese name Number of individuals Females Males Total % Colletidae ColletesColletescollaris Dours ColletesColletespatellatus Pérez HylaeusNesoprosopisfloralisSmith Halictidae HalictusSeladoniaaerarius Smith LasioglossumEvylaeusaffineSmith LasioglossumEvylaeusduplexDalla Rorre LasioglossumEvylaeusjaponicumDalla Torre LasioglossumEvylaeusohei Hirashima et Sakagami LasioglossumEvylaeuspallilomumStrand LasioglossumEvylaeustaeniolellumVachal LasioglossumEvylaeusvulsumVachal LasioglossumLasioglossummutilumVachal LasioglossumLasioglossumoccidensSmith LasioglossumLasioglossumprimavera Sakagami et Maeta LasioglossumLasioglossumproximatumSmith LasioglossumLasioglossumscitulumSmith NomiaHoplonomiapunctulata Dalla Torre Sphecodes japonicus Cockerell Andrenidae AndrenaChlorandrenaknuthi Alfken AndrenaEuandrenahebes Pérez AndrenaEuandrenalaridiloma Strand AndrenaHoplandrenadentata Smith AndrenaMelandrenawatasei Cockerell AndrenaMicrandrenahikosana Hiraahima AndrenaMicrandrenakaguya Hirashima AndrenaMicrandrenaminutulaKirby AndrenaMicrandrenasemirugosa brassicae Hirashima AndrenaSimandrenayamato Tadauchi et Hirashima Megachilidae CoelioxysBoreocoelioxyshiroba Nagase CoelioxysBoreocoelioxysyanonis Matsumura Euaspis basalisritsema Megachile humilis Smith Megachile kyotensis Alfken Megachile nipponica Cockerell Megachile pseudomonticola Hedicke Megachile sculpturalis Smith Megachile tsurugensis Cockerell Osmia cornifronsradoszkowski Osmia taurus Smith Apidae Amegilla florea floreasmith Apis cerana japonica Radoszkowski BombusDiversobombusdiversus diversus Smith BombusPyrobombusardens ardens Smith Epeolus japonicus Bishoff Eucera nipponensispérez Eucera spurcatipes Pérez Nomada ginran Tsuneki Nomada japonica Smith Nomada shirakii Yasumatsu et Hirashima Nomada sp. Thyreus decorussmith Xylocopa appendiculata circumvolans Smith Total 1, ,

21 20 A:, B:, C:. Fig. 3. Some wild bees collected at the Okadaira Shell Mound. A: Lasioglossum primavera, B: Amegilla florea florea, C: Thyreus decorus , D, 1/D, 1 D=0.94 1/D=16.42 C C ,464 1, Colletes 2 Hisamatsu and Yamane2006 Solidago altissima L. Lactuca indica L. Aster yomenakitam.honda var. dentatus Kitam.H.HaraBidens frondosa L. 2010

22 21 : ;, 2008;, 1985;, 1992;, 2011;, Hisamatsu2010;, Hisamatsu and Yamane2006. Fig. 4. Relative numbers of bee species for each family collected at the Okadaira Shell Mound. Data sources: Okadaira, present study; Yamizo, Hisamatsu and Yamane2008; Gozenyama, Iho and Yamane1985; Mito, Saito et al.1992; Ajigaura, Hisamatsu2011; Tsukuba, Hisamatsu2010; and Sugao, Hisamatsu and Yamane2006. :, ;, 2008;, 1985;, 1992;, 2011;, Hisamatsu2010;, Hisamatsu and Yamane2006. Fig. 5. Relative numbers of individual bees for each family collected at the Okadaira Shell Mound. Data sources: Okadaira, present study; Yamizo, Hisamatsu and Yamane2008; Gozenyama, Iho and Yamane1985; Mito, Saito et al.1992; Ajigaura, Hisamatsu2011; Tsukuba, Hisamatsu2010; and Sugao, Hisamatsu and Yamane2006.

23 22 7 1SN 1 D1/D. Table 2. Number of wild bee species collected over 1 years, total number of individuals collectedn, Simpson's diversity1å D, and inverse of Simpson's index1/dat seven localities in Ibaraki Prefecture. Okadaira Surveyed localities Yamizo Gozenyama Mito Ajigaura Tsukuba Sugao Number of speciess Number of individualsn Simpson s diversity1-d Inverse of Simpson's index1/d Data sources: Okadaira, present study;yamizo, Hisamatsu and Yamane2008; Gozenyama, Iho and Yamane1985; Mito, Saito et al.1992; Ajigaura,Hisamatsu, 2011; Tsukuba, Hisamatsu2010; and Sugao, Hisamatsu and Yamane C. Table 3. Number of common species and Morishita s index of similaritycbetween the Okadaira Shell Moundpresent studyand six other localities in Ibaraki Prefecture. Surveyed localities Yamizo Gozenyama Mito Ajigaura Tsukuba Sugao Number of common species Morisita s Cindex Data sources: Yamizo, Hisamatsu and Yamane2008; Gozenyama, Iho and Yamane1985; Mito, Saito et al.1992; Ajigaura, Hisamatsu2011; Tsukuba, Hisamatsu2010; and Sugao, Hisamatsu and Yamane Table 4. Number of Amegilla florea florea and Thyreus decorus individuals that were collected on each collection date. Species names Dates of collection 7-Aug 16-Aug 26-Aug 5-Sep 14-Sep 24-Sep Amegilla florea florea female male Thyreus decorus female male C

24 23 4 LasioglossumEvylaeusohei Hirashima et Sakagami32 Osmia taurus Smith18 Euaspis basalisritsema16 Megachile kyotensis Alfken6 Megachile pseudomonticola Hedicke1 4 1 D 1/D Bombus Diversobombusdiversus diversus Smith LasioglossumEvylaeusapristumVachal C 0.64C C ; 2008; 1985; 1992; 2011; Hisamatsu2010; Hisamatsu and Yamane2006. Fig. 6. Dendrogram showing the intercommunity similarities among seven points surveyed in Ibaraki Prefecture, based on Morishita s index of similarityc. The clustering was made using an average linkage clustering method. Data sources: Okadaira, present study; Yamizo, Hisamatsu and Yamane2008; Tsukuba, Hisamatsu2010; Gozenyama, Iho and Yamane 1985; Ajigaura, Hisamatsu2011; Mito, Saito et al.1992; and Sugao, Hisamatsu and Yamane2006.

25 24 Ebmer, A. W. and S. F. Sakagami. 1985a. Taxonomic notes on the palaearctic species of the Lasioglossum nitidiusculum group, with a description of L. allodalum sp. nov. Hymenoptera, Halictidae. Konchû, : Ebmer, A. W. and S. F. Sakagami. 1985b. Lasioglossum Evylaeus hirashimae n. sp., ein Vertreter einer paläotropischen Artgruppe in Japan Hymenoptera, Apoidea. Nachrichtenblatt der bayerischen Entomologen, : : : : Hisamatsu, M Species diversity and composition of wild bees observed at Mt. Tsukuba, Ibaraki Prefecture, eastern Japan. Japanese Journal of Environmental Entomology and Zoology, : : : Hisamatsu, M. and Sô. Yamane Faunal makeup of wild bees and their flower utilization in a semi-urbanized area in central Japan. Entomological Science, : : : , 27: Japanese Journal of Entomology, : Krebs, C. J Ecological methodology, 2nd edn. 624 pp., Addison Wesley Longman, Menlo Park, New Jersey, USA. Mitai, K. and O. Tadauchi Taxonomic notes on Japanese species of the Nomda furva species grouphymenoptera: Apidae. Entomological Science, : Matsumura, M., S. F. Sakagami and H. Fukuda A wild bee survey in KibiWakayama Pref., southern Japan. Journal of Faculty of Science, Hokkaido University, Series VI, Zoology, : Nagase, H Synopsis of the bee genus Coelioxys Latreille Hymenoptera: Megachilidaeof Japan, with a description of a new species. Entomological Science, : : pp., : Sakagami, S. F. and H. Fukuda Wild bee survey at the campus of Hokkaido University. Journal of Faculty of Science, Hokkaido University, Series VI, Zoology, : : Sakagami, S. F. and O. Tadauchi. 1995a. Taxonomic studies on the halictine bees of LasioglossumEvylaeuslucidulum subgroup in Japan with comparative notes on some palaearctic specieshymenoptera, Apoidea. Esakia,35: Sakagami, S. F. and O. Tadauchi. 1995b. Three new halictine bees from JapanHymenoptera, Apoidea. Esakia,35: pp., pp., Takahashi, H. and S. F. Sakagami Notes on the Halictinae Hymenoptera, Apoideaof the Izu Islands: Lasioglossum kuroshio sp. nov., life cycle in Hachijo-jima Is., and a preliminary list of the species in the Izu Islands. Japanese Journal of Entomology, :

26 , Colletes patellatus 207 Apis cerana japonica Co. collaris Lasioglossum primavera Amegilla florea florea Thyreus decorus

27 Bull. Ibaraki Nat. Mus.,14: h Effectiveness of a Modified Cage Trap for Raccoon to Avoid Mis-capturing of Native Species in Ibaraki Prefecture, Central Japan Koji YAMAZAKI, Midori SAEKI, Masanori HIROHARA and Tsutomu SAKAMOTO Accepted April 7, 2011 Abstract The effectiveness of a modified cage trap, which was equipped with a specialized trigger for raccoon, was tested and compared with two types of commercial cage traps in Ibaraki prefecture. Camera traps were also set close to the modified traps to monitor approaching animals and to know how the triggers were activated. No differences were detected in capturing rates between the modified trap and the commercial traps ( vs raccoons/tn). On the other hand, mis-capturing of native mammals by the modified trap was very low, showing that use of this type of trap can minimize adverse effects on the native ecosystem, avoid accidental capture of pet animals, and decrease the laborious work of releasing non-target species. Widespread use of this trap in raccoon removal operations is recommended. Key words: cage trap, Ibaraki Prefecture, invasive alien species, Procyon lotor, raccoon, removal Procyon lotor h2009; h 2010 h 2010 * Ibaraki Nature Museum, 700 Osaki, Bando, Ibaraki , Japan. ** National Agricultural Research Center, Kannondai, Tsukuba, Ibaraki , Japan. *** Kasumigaura City Hall, Owada, Kasumigaura, Ibaraki **** Ibaraki Prefecture Kennan-Kenmin Center, Manabe, Tsuchiura, Ibaraki

28 28 h 2010 Table 1. Results of raccoon trapping by modified cage traps, and commercial cage traps in Kasumigaura City and Bando City, Ibaraki in 2010.!!!! " "! a h 2010: : Havahart R #1081, Woodstream Corp., Lititz, Pennsylvania, USA: 381 mm 381 mm 1,066 mm b : No.281L: 370 mm 370 mm 800 mm c : Havahart R #1089, Woodstream Corp., Lititz, Pennsylvania, USA: 315 mm 265 mm 815 mm a b c a b c a b h c h

29 a: 20:19b: 20:20. Fig. 1. An example of raccoon capture by a modified cage trap at a peanut field in the Saka area, Kasumigaura City on 2 October a: A raccoon approached the trap at 20:19 (above), b: A raccoon then trapped at 20:20. Wildview Xtream STC-TGL5IRWildview, Texas, USA 2GB SD high resolution Chi-square test, P = Fisher s exact test, P = Fisher s exact test, P = Fisher s exact test, P = Nyctereutes procyonoides 11 1 Paguma larvata 9 Felis silvestris catus Table 2. Results of raccoon trapping monitoring by a modified cage trap using a digital sensor camera in Kasumigaura City and Bando City, Ibaraki in 2010.

30 30 h 2 SD 29100% 1 Canis lupus familiaris Corvus coronec. macrorhynchos Garrulus glandarius %5 / % %1 / h L 45% h 2010

31 : h : h Procyon lotor

32 Bull. Ibaraki Nat. Mus.,14: A List of Coleoptera Collected at Mt. Tsukuba, Ibaraki Prefecture, Central Japan Sadahiro OHMOMO and Masaki HISAMATSU Accepted January 18, 2011 Abstract Based on records published by previous authors, a list of Coleoptera at Mt. Tsukuba is presented. This updated list includes a total of 720 species in 76 families. Key words: List of Coleoptera, Mt. Tsukuba. 877 m , , ; , * Sumiyoshi, Ami, Inashiki, Ibaraki , Japan ** Ibaraki Nature Museum, 700 Osaki, Bando, Ibaraki , Japan

33 Fig. 1. The map of surveyed area at Mt. Tsukuba.

34 35 Table 1. The number of coleopterous species for each family recorded at Mt. Tsukuba. 1Cicindelidae 3 2Paussidae 1 3Carabidae 76 4Brachinidae 1 5Dytiscidae 8 6Gyrinidae 3 7Helophoridae 1 8Hydrophilidae 1 9Histeridae 4 10Leiodidae 1 11Silphidae 3 12Staphylinidae 38 13Scydmaenidae 2 14Pselaphidae 1 15Scaphidiidae 4 16Helodidae 3 17Rhipiceridae 1 18Lucanidae 4 19Scarabaeidae 31 20Ptilodactylidae 2 21Psephenidae 1 22Elmidae 6 23Buprestidae 25 24Elateridae 30 25Throscidae 1 26Eucnemidae 3 27Lycidae 8 28Cantharidae 13 29Omethidae 1 30Lampyridae 5 31Dermestidae 2 32Anobiidae 3 33Trogossitidae 2 34Cleridae 4 35Melyridae 4 36Nitidulidae 22 37Phalacridae 3 38Sphindidae 1 39Laemophloedae 1 40Silvanidae 3 41Cryptophagidae 2 42Helotidae 1 43Byturidae 1 44Biphyllidae 3 45Languriidae 4 46Erotylidae 13 47Corylophidae 1 48Endomychidae 7 49Coccinellidae 21 50Lathridiidae 1 51Ciidae 5 52Mycetophagidae 3 53Tetratomidae 6 54Zopheridae 5 55Tenebrionidae 18 56Lagriidae 3 57Alleculidae 6 58Salpingidae 1 59Cephaloidae 1 60Pyrochroidae 2 61Melandryidae 10 62Rhipiphoridae 1 63Mordellidae 7 64Scraptiidae 1 65Oedemeridae 6 66Meloidae 1 67Anthicidae 2 68Disteniidae 1 69Cerambycidae 91 70Chrysomelidae 73 71Anthribidae 12 72Attelabidae 19 73Apionidae 5 74Curculionidae 55 75Rhynchophoridae 3 76Scolytidae 3

35 36 Cicindelidae 1 Cicindela gemmata Falderermann 1999; Cicindela japana Motschulsky 1969; ; Cicindela specularis Chaudoir 1999; 2000 Paussidae 1 Eustra japonica Bates, 1990;, 2000 Carabidae 1 Carabus albrechti esakianusnakane, ; 1999; ; ; Carabus insulicola insulicola Chaudoir ; ; 1990; Damaster blaptoides oxuroides Schaum 1969; 1990; ; Hemicarabus tuberculosusdejean et Boisduval 1999; Leptocarabus arboreus parexilisnakane 1989; 1998; 1990; Leptocarabus proceruluscaudoir 1989; 1990; Nebria lewisi Bates Notiophilus impressifrons Morawitz ; Kurasawatrechus quadraticollis S. Ueno 1990; Asaphidion semilucidum Motschulsky 1999; ; Bembidion morawitzi Csiki 1999; Patrobus flavipes Motschulsky 1969; 1990; Pterostichus leptis Bates 1999; Pterostichus microcephalus Motschulsky 1990; ; Pterostichus mirificus Bates 1986; Pterostichus nishiyamai Kasahara 1986; ; ; Pterostichus rhanis rhanis

36 37 Tschitscherine 1986; Pterostichus takaosanus Habu ; ; Pterostichus tsukubasanus Kasahara Kasahara, 1988; ; Pterostichus yoritomus Bates 1998; 1986; ; ; Pterostichus fuligineus Morawitz 1990; Pterostichus spiculifer spiculifer Bates 1990; Stomis prognathus Bates 1986; ; Trigonognatha cuprescens Motschulsky Colpodes aequatus Jedliaka 1990; Colpodes atricomes Bates 1999; ; ; Colpodes aurelius aurelius Bates 1999; ; Colpodes buchanani Hope 1999; ; Colpodes elainus elainus Bates 1999; Colpodes japonicus Motschulsky 1999; Colpodes modestior Bates 1990; Colpodes speculator Halord 1999; ; Colpodes sylphis stichai Jedliaka ; Dicranoncus femoralis Chaudoir 1999; ; ; Dicranoncus femoralis Chaudoir Parabroscus crassipalpus Bates ; Platynus magnusbates 1987b; 2000

37 38 38 Platynus protensusmorawitz 1999; ; Platynus subovatus Putzeys 1999; Pristosia aeneolabates ; ; ; Synuchus cycloderus Bates ; Synuchus nitidus Motschulsky ; Synuchus silvester Habu 1999; Synuchus melantho Bates Amara chalcites Dejean 1999; ; Anisodactylus sadoensis Schauberger 1990; Anisodactylus punctatipennis Morawitz Harpalus eous Tschitscherine 1999; Harpalus griseuspanzer ; Harpalus pseudophonoides Schauberger 1999; Harpalus tridens Morawitz 1990; Oxycentrus argutoroidesbates ; ; Trichotichnus congruus Motschulsky 1990; ; ; Trichotichnus lewisi Schauberger 1999; Trichotichnus nipponicus Habu 1990; Bradycellus grandiceps Bates 1999; Bradycellus laeticolor Bates 1987b; Stenolophus propinquus Morawitz 1987b; Chlaenius micans Fabricius 1999; Chlaenius micansfabricius

38 ; Chlaenius posticalis Motschulsky 1969; 1990; Chlaenius variicornis Morawitz 1999; Chlaenius virgulifer Chaudoir 1999; Haplochlaenius costoger Chaudoir 1999; ; Coptodera subapicalis Putzeys 1999; Dromius prolixus Bates 1990; ; ; ; Dromius quadraticollis Morawitz 1990; Lebia bifenestrata Morawitz 1987b; ; ; ; Lebia duplex Bates 1999; Lebia retrofasciata Motschulsky 1969; ; ; ; Lebia viridis Say 1999; Lebidia bioculata Morawitz 1990; ; Lebidia octoguttata Morawitz 1999; ; Lachnolebia cribricollis Morawitz 1987b; Parena monostigma Bates 1987b; ; Parena nigrolineata nipponensis Habu 1999; ; 2000 Brachinidae 1 Brachinus scotomedes Redtenbacher 1999; 2000 Dytiscidae 1 Hyphydrus japonicus Sharp 1987a; Agabus conspicuus Sharp

39 a; a; Agabus japonicus Sharp 1987a; a; Platambus stygius Reginbart Rhantus erraticus Sharp 1986; 1987a; Rhantus pulverosusstephens 1987a; a; ; Cybister japonicus Sharp 1987a; Hydaticus grammicusgermar 1987a; 2000 Gyrinidae 1 Dineutus orientalismodeer 1987a; Gyrinus curtus Motschulsky 1987a; Gyrinus japonicus Sharp 1987a; a; 2000 Helophoridae 1 Helophorus auriculatus Sharp 1993; ; 2000 Hydrophilidae 1 Peratogomus reversus Sharp 2005 Histeridae 1 Carcinops pumilioerichson 1993; Notodoma fungorum Lewis 1999; Margarinotus weymarni Wenzel 1999; Platysoma celatum Lewis 1993; ; 2000 Leiodidae 1 Pseudoliodes strigosulus Portevin 2008 Silphidae 1 Nicrophorus maculifrons Kraatz 1999; Nicrophorus quadripunctatus Kraatz 1999; Necrodes asiaticus Portevin 1999; 2000 Staphylinidae 1 Siagonium vittatum Fauvel 1999; Megarthrus japonicus Sharp Megarthrus convexus Sharp

40 Lesteva plagiata Sharp 1993; Olophrum arrowi Scheerpeltzarp 1999; Olophrum vicinum Sharp Osorinus angustulus Sharp Stenus santira Naomi Oxyporus japonicus Sharp 1997; Oxyporus parcus Sharp 1999; ; Megalopaederus lewisi Cameron 1999; ; Paederus fuscipes Curtis 1999; ; Stilicoderus signatus Sharp Rugilus longipennis Sharp Nudobius apicipennis Sharp 1999; Algon grandicollis Sharp 1999; ; Anisolius elegans Sharp ; Eucibdelus japonicus Sharp 999; ; Creophilus maxillosus Linnaeus Philonthus cyanipennis Fabricius 1999; Gabrius sharpianus Cameron Quedius planatus Sharp Mycetoporus duplicatus Sharp Bolitobius prolongatus Sharp 1999; Bolitobius setigersharp ; Carphacis striatus Oliver 1999; Lordithon bicolor Gravenhorst 1999; Lordithon irregularis Weise 1998

41 ; Lordithon semirufus Sharp 1999; Sepedophilus bipustulatusgravenhorst 1999; Sepedophilus fimbriatus Sharp 1999; ; Sepedophilus tibialissharp 1999; ; ; Sepedophilus varicornis Sharp Tachinus cpunctiventris Sharp 1999; Phymatura oligotinulasharp 1997; Tachyporus celatus Sharp Tachyporus suavis Sharp Aleochara parens Sharp 2004; 2005 Scydmaenidae 1 Cephennodes vafer Kurbatou alt mjaloszynski et Hoshina, Paranesenthia paradoxak.sawada alt. 850 mjaloszynski et Hoshina, Batriscenaulax furuhatai Kurbata 1997; 2000 Scaphidiidae 1 Ascaphium tibiale Lewis 1999; Ascaphium apicale Lewis 2005; Scaphium japonum Reitter 1999; ; Scaphisoma rufum Achard 1999; 2000 Helodidae 1 Cyphon ishiharai K.Sasagawa 1997; ; Cyphon japonicola Nakane 1997; Sacodes protectus Harold 1999; ; 2000 Rhipiceridae 1 Sandalus segnis Lewis 1999; 2000 Lucanidae

42 43 1 Lucanus maculifemoratus Motschulsky 1969; ; ; ; Macrodorcas rectusmotschulsky 1999; Macrodorcas striatipennis Motschulsky 1969; ; Nipponodorcus rubrofemoratus S. van Vollenhoven 1999; 2000 Scarabaeidae 1 Geotrupes laevistriatus Motschulsky 1999; Heptophylla picea Motschulsky 1969; ; ; ; Hoplia communis Waterhouse 1999; Hoplia moerens Waterhouse Maladera japonicamotschulsky 1969; Maladera castaneaarrow 1999; ; Maladera renardiballion 1997; Nipponoserica peregrina Chapin 1999; Paraserica gricea Motschulsky 1969; ; ; ; ; Serica boops Waterhouse 1999; ; ; ; Sericania lewisi Arrow 1976; ; Sericania ohtakei Sawada 1999; Adoretus tenuimaculatus Waterhouse 1969; ; ; ; Anomala daimiana Harold 1999; ; ; Blitopertha orientaliswaterhouse 1999; Mimela costatahope 1999; Mimela flavilabriswaterhouse 1999;

43 Mimela testaceipesmotschulsky 1999; ; Popillia japonica Newmann 1969; ; Dasyvalgus tuberculatus Lewis 1969; Nipponovalgus angusticollis Waterhouse 1999; ; ; Lasiotrichius succinctuspallas 1969; Trichius japonicus Janson 1969; Eucetonia piliferamotschulsky 1969; Eucetonia roelofsiharold 1969; Glycyphana fulvistemma Motschulsky 1969; ; Oxycetonia jucundafaldermann 1969; Protaetia orientalis submarmorea Burmeister 1969; Rhomborrhina japonica Hope 1969; Rhomborrhina polita Waterhouse 1969; Allomyrina dichotoma Linnaeus 1969; Epilichas flabellatus Kiesenwetter 1993; ; Epilichas monticola Nakane 1999; ; ; 2000 Psephenidae 1 Eubrianax pellucidus Lewis 1997; 2000 Elmidae 1 Dryopomorphus nakanei Nomura 1996; 1997; 1999; ; ; Optioservus nitidus Nomura 1 ex., 12.IX Grouvellinus nitidus Nomura Zaitzevia nitida Nomura 1998; 2000; 1 ex., 12.IX Zaitzeviaria gotoi Nomura 1998; Zaitzeviaria ovatanomura 1996; 2000; ; ; ; 2000; 1 ex., 12.IX.1998 Ptilodactylidae Buprestidae

44 45 1 Chrysochroa fulgidissima fulgidissima Schönherr 1969; 1979a; ; Chalcophora japonica japonicagory 1969; 1979a; ; Buprestis haemorrhoidalis japanensis Saunders 1979a; ; Anthaxia proteus Saunders 1982; ; Chrysobothris succedanea Saunders 1982; Coroebus quadriundulatus Motschulsky 1979b; ; Agrilus adelphinus Kerremans 2005; Agrilus daimio Obenberger 1999; Agrilus moerens Saunders 1982; Agrilus ribbei Kiesenwetter 1980; Agrilus tempestivus Lewis 1982; ; Agrilus trinotatus Saunders 1982; Agrilus viridiobscurus Saunders 1980; ; Habroloma subbicorne Motschulsky 1982; Habroloma yuasai Y.Kurosawa 2004; Trachys auricollis Saunders 1979a; ; ; ; ; Trachys broussonetiae Y.Kurosawa 1980; Trachys cupricolor Saunders alt. 260m Trachys griseofasciata Saunders 1999; Trachys inconspicua Saunders 1980; Trachys minuta salicislewis 1979a; 1980; Trachys reitteri Obenberger 1980; Trachys tsushimae Obenberger 1980;

45 ; Trachys variolaris Saunders 1980; Trachys yanoi Y.Kurosawa 1999; 2000 Elateridae 1 Pectocera fotunei Candeze 1999; ; ; ; Agrypnus binodulusmotschulsky 1969; ; ; ; Agrypnus cordicolliscandeze 1999; ; ; ; Paracalais beruscandeze 1973d; Tetrigus lewisi Candeze 1999; ; Actenicerus giganteus Kishii 1993; Actenicerus orientalis Candeze 1969; Actenicerus pruinosusmotschulsky 1969; ; Corymbitoides gratus Lewis 1999; Denticollis nipponensis Ohira 1999; Eanoides puneriliscandeze 1999; ; Limoniscus vittatuscandeze 1999; Hemicrepidius desertor Candeze 1997; Selatosomus onerosuslewis 1999; Stenagostus umbratilislewis 1999; ; ; Ampedus japonicus Silfverberg 1999; ; Ampedus vestituslewis 1999; Dolerosomus graciliscandeze 1999; ; Ectinus sericeuscandeze 1999;

46 ; ; Hayekpenthes pallidus Lewis 1999; ; Ischnodes sanguinicollis mikado W.Suzuki 1999; Neotrichophorus juniorcandeze 1999; Melanotus cete Candeze 1999; ; Melanotus japonicus Ohira 1997; Melanotus koikei Kishii et Ohira 1999; ; Melanotus legatus Candeze 1999; ; ; ; Melanotus restrictus Candeze 1999; Melanotus senilis Candeze 1999; Paracardiophorus opacus Lewis 2004; Paracardiophorus pullatuscandeze 1999; 2000 Throscidae 1 Aulonothroscus longulus Weise 1996; 2000 Eucnemidae 1 Fornax victor Fleutiaux 2004; Dromaeolus nipponensis Fleutiaux 2004; Dromaeolus brevipes Fleutiaux 2004; 2005 Lycidae 1 Benibotarus spinicoxis Kiesenwetter 1993; Lyponia delicatulakiesenwetter 1999; Lyponia quadricollis Kiesenwetter 1999; Macrolycus aemulus Barovskij 1993; Macrolycus flabellatus Motschulsky 1997; Macrolycus montanus

47 48 Nakane 1993; Plateros coracinuskiesenwetter 1993; Pyropterus nigroruberdegeer 1999; 2000 Cantharidae 1 Lycocerus insulsusharold 1999; Lycocerus aegrotus Kiesenwetter 1999; Lycocerus attristatuskiesenwetter 1993; Lycocerus suturellusmotschulsky 1969; ; ; ; Podabrus heydeni Kiesenwetter 1999; ; ; ; Podabrus lictorius Lewis 1993; ; ; Podabrus malthinoides Kiesenwetter 1999; Podabrus temporalis Harold 1999; ; Prothemus reinii Kiesenwetter 1969; ; ; ; Stenothemus badius Kiesenwetter 1999; Themus cyanipennis Motschulsky 1999; ; ; Themus niisatoi M. Sato et K. Takahashi 1997; ; Malthodes minutopygus Wittmer 2005 Omethidae 1 Omethes rugicepslewis 1996; ; 2000 Lampyridae 1 Hotaria parvulakiesenwetter 1933; 1980; 1993; Luciola cruciata Motschulsky 1980; ; Luciola lateralis Motschulsky 1980;

48 Lucidina biplagiatamotschulsky 1999; ; ; Lychnuris fumosagorham 1999; 2000 Dermestidae 1 Thaumaglossa hilleri Reitter 1997; Anthrenus japonicus N.Ohbayashi 1999; 2000 Anobiidae 1 Priobium cylindricumnakane 1999; Lasioderma serricornefabricius alt. 60 m ; Dorcatoma shigaensis N.Hayashi Opilo carinatus Lewis 1999; Opilo niponicus Lewis 1999; ; ; Stigmatium nakanei Iga 1999; 2000 Melyridae 1 Dasytes japonicus Kiesenwetter 1999; ; Dasytes vulgaris Nakane 1999; Attalus elongatulus Lewis 1999; Malachius prolongatus Motschulsky 1969; ; 2000 Trogossitidae 1 Trogossita japonica Reitter 1997; Latolaeva japonicareitter 1999; 2000 Cleridae 1 Tillus igarashii Kono 2004; Nitidulidae 1 Carpophilus chalybeus Murray 1999; ; Aethina inconspicua Nakane Amphicrossus lewisi Reitter 1999; 2000

49 50 4 Atarphia fasciculata Reitter 2004; Cyllodes aterherbst 1999; ; Cyllodes dubiusreitter 1999; Epuraea durareitter 2004; ; Epuraea adumbrata Mannerheim 2004; Epuraea commutata Grouvelle 1999; Epuraea mandibularis Reitter 1999; Haptoncurina paulula Reitter 1997; Ipidia variolosa Reitter 1999; ; ; Pallodes umbratilis Reitter alt. 260 m Neopallodes inermis Reitter 1996; 2000alt m Omosita colonlinnaeus 1999; Physoronia explanata Reitter 1999; Pocadites dilatimanus Reitter 1999; ; Pocadites japonusreitter 2004; Soronia lewisi Reitter 1999; Stelidota multiguttata Reitter 1999; 2000 alt. 60 m Meligethes flavicollis Reitter Librodor japonicus Motschulsky 1969; 2000 Phalacridae 1 Heterolitus coronatus Flach 1999; Heterolitus nipponicus Hisamatsu 1999; Stilbus avunculus Flach 1997; 2000 Sphindidae 1 Aspidophorus japonicus Reitter

50 ; 2005alt. 660 m 2005 Laemophloedae 1 Laemophloeus submonilis Reitter 2004; 2005 Silvanidae 1 Silvanus bidentatus Fabricius 1996; Silvanoprus inermisreitter alt. 60 m Psammoecus fasciatus Reitter 1999; 2000 Cryptophagidae 1 Cryptophagus callosipennis Grauvelle 1999; Cryptophagus latangulus Reitter 1999; 2000 Helotidae 1 Helota gemmata Gorham 1969; 2000 Byturidae 1 Byturus affinis Reitter 1999; ; 2000 Biphyllidae 1 Biphyllus humeralisreitter 1993; Biphyllus rufopictus Wollaston alt. 60 m Biphyllus throscoides Wollaston 2004; 2005 Languriidae 1 Toramus glisonothoides Reitter alt. 60 m Henoticonus triphylloides Reitter alt. 60 m Anadastus praeustus Crotch 2004; Languriomorpha lewisi Crotch 1999; ; ; ; 2000 Erotylidae 1 Aulacochilus japonicus Crotch 1999; Aulacochilus sibiricus Reitter 1993; Encaustes praenobilis Lewis 1999; Neotriplax atrata Lewis 1999; ; ;

51 Neotriplax lewisicrotch 1999; Triplax japonica Crotch 1999; Triplax sibirica connectens Lewis 1999; Tritoma latifasciata Lewis 1999; ; Tritoma niponensislewis 1999; ; ; Tritoma pallidicincta Lewis 1996; ; ; Dacne zonalia Lewis 1999; Episcapha gorhami Lewis 1999; ; Episcapha morawitzisolsky 1969; 2000 Corylophidae 1 Arthrolips oblongus Matthews 1997; 2000 Endomychidae 1 Danae orientalis Gorham 1985; Ectomychus basalis Gorham 1985; Ectomychus musculus Gorham 1985; Ancylopus pictus asiaticus Strohecker 1990; Mycetina amabilis Gorham 1983; Mycetina ancoriger Gorham 1990; Endomychus gorhamilewis 1983; ; ; ; ; 2000 Coccinellidae 1 Serangium japonicum Chapin 1999; Hyperaspis japonicacrotch 1991; ; ; Pseudoscymnus harejaweise

52 ; ; Scymnus kawamuraiohta 1991; ; Stethorus japonicus H.Kamiya 1999; ; ; ; Chilocorus kuwanae Silvesti 1999; Phymatosternus lewisiicrotch 1991; Telsimia nigraweise 1999; Rodolia concolorlewis 1999; ; Aiolocaria hexaspilotahope 1999; ; Anatis halonis Lewis 1969; ; ; Calvia quatuordecimguttata Linnaeus 1999; ; ; Calvia muiritimberlake 1999; Coccinella septempunctata Linnaeus 1969; ; ; ; Harmonia axyridispallas 1969; ; ; ; ; Illeis koebelei koebelei Timberlake 1991; ; ; ; Oenopia scalaristimberlake 1999; Propylea japonicathunberg 1969; ; ; ; Sospita oblongoguttata Linnaeus 1999; ; Vibidia duodecimguttatapoda 1999; ; Epilachna admirabilis Crotch

53 ; ; 2000 Lathridiidae 1 Corticaria japonica Reitter 1996; 2000 Ciidae 1 Cis boleti polypori Chujo 1999; ; Cis seriatopilosus Motschulsky 1999; Cis nipponicus Chujo 2004; Nipponocis ashuensis Nobuchi 1999; ; Orthocis shizophylli Nakane et Nobuchi 1996; ; ; 2000 Mycetophagidae 1 Mycetophagus antennatus Reitter 1999; ; Mycetophagus aterreitter 1999; Mycetophagus pustulosus Reitter Tetratomidae 1 Pisenus rufitarsisreitter 1999; Abstrulia japonica Miyatake 1996; Tetratoma nobuchii Nakane 1996; Orchesia elegantula Lewis 1999; ; Orchesia imitans Lewis 1999; ; Orchesia ocularis Lewis 1993; ; ; 2000 Zopheridae 1 Pycnomerus vilis Sharp Glyphocryptus brevicollis Sharp Acolophus debilis Sharp Cicones niveus Sharp 2000; Endophloeus serratus Sharp 2004; 2005

54 55 Tenebrionidae 1 Boletoxenus bellicosus Lewis 1999; Parabolitophagus felix Lewis 1996; Ceropria induta Wiedemann 1999; Ceropria sulcifrons Harold 1999; Platydema nigroaeneum Motschulsky 1999; ; ; Derispia maculipennismarseul 1999; Leiochrodes convexus Lewis 1999; 2000alt. 260 m Uloma latimanus Kolbe 1999; ; Uloma marseuli marseuli Nakane 1999; Encyalesthus violaceipennis Marseul 1999; Toxicum tricornutum Waterhouse 1969; ; ; Addia scatebrae Lewis 1987; ; Tarpela elegantula Lewis Simalura coerulea Lewis 1999; Plesiophthalmus nigrocyaneus nigrocyaneus Motschulsky 1969; ; ; Ainu tenuicornis Lewis 1987; ; ; Strongylium impigrum Lewis 1999; ; ; ; Strongylium niponicum Lewis 1987; ; 2000 Lagriidae

55 56 1 Arthromacra decoramarseul 1969; ; ; ; Lagria nigricollis Hope 1969; ; ; Macrolagria rufobrunnea Marseul 1999; ; ; ; ; 2000 Alleculidae 1 Alleculta cryptomeriae Lewis 2004; Alleculta melanaria Maklin 1999; ; Alleculta simiola Lewis 1999; ; ; Borboresthes acicularis Marseul 1999; ; ; Hymenalia unicolor Nakane 1999; Isomira oculatamarseul 1999; 2000 Salpingidae 1 Lissodema dentatum Lewis 1996; 2000 Cephaloidae 1 Stolius vagepictus Lewis 2000; Pyrochroidae 1 Pseudopyrochroa peculiaris Lewis 1999; Pseudopyrochroa brevitarsis Lewis 2002; 2005 Melandryidae 1 Lederia latalewis Sasaji1987; Lederia japonica Reitter Anisoxya conicicollis Champion 1999; Enchodes crepusculus Lewis 1999; Melandrya gloriosa Lewis 1993; ; 2000

56 ; Phloeotrya obscuralewis 1999; Phloeotrya rugicollis Marseul 1993; ; ; ; ; Serropalpus niponicus Lewis 1999; Symphora atra Nomura 1993; Symphora miyakei Nomura et Hayashi 1993; 2000 Rhipiphoridae 1 Pelecotomoides tokejii Nomura et Nakane 1997; ; ; ; 2000 Mordellidae 1 Falsomordellina takaosana Kono 1996; Glipostenoda resseola Marseul 1999; Hoshihananomia perlatasulzer 1999; Mordellina yamamotoi Nomura et Kato Mordellaria arakiinakane et Nomura 1993; Mordellistena tokejii Nomura 1997; Mordellistenoda akakono 1997; 2000 Scraptiidae 1 Anaspis marseuli Csiki 1999; ; ; 2000 Oedemeridae 1 Asclera konoi Nakane 1997; Oedemerinia lucidicollis Motschulsky 1999; ; ; Oedemerinia manicata Lewis 1999; ; Nacerdes deformislewis 1997; ;

57 ; Nacerdes katoikono 1997; Nacerdes luteipennis katoi Marseul 1999; ; Nacerdes waterhouseiharold 1999; ; 2000 Meloidae 1 Zonitis japonica Pic 1999; 2000 Anthicidae 1 Sapintus cohaereslewis 1999; Anthicomorphus niponicus Lewis alt. 160 m 2005 Disteniidae 1 Distenia gracilis gracilisblessig 1989; ; 1990; ; ; 2000 Cerambycidae 1 Megopis sinica sinica White 1977; ; 1990; ; Prionus insularis insularis Motschulsky 1999; Spondylis buprestoides Linnaeus 1989; Arhopalus coreanussharp 1987; 1989; Asemum striatum Linnaeus 1999; Megasemum quadricostulatum Kraatz 1999; Dinoptera minutagebler 1990; ; 1990; ; ; ; ; Lemula decipiens Bates 1982; 1990; Toxotinus reiniheyden 1999; Pidonia aegrota aegrota Bates 1982; 1990; ; ;

58 59 11 Pidonia puziloisolsky 1989; ; 1990; ; ; Pidonia signiferabates 1989; ; 1990; ; ; ; Anastrangalia scotodes Bates 1982; 1990; ; 1990; ; 1990; 2000; ; Aredolpona succedanealewis Leptura dimorpha Bates 1999; ; ; Leptura modicenotata Pic 1969; ; 1989; ; 1990; Leptura ochraceofasciata ochraceofasciata Motschulsky 1969; ; ; 1990; ; ; ; Bellamira regalisbates Parastrangalis nymphula Bates 1982; 1990; ; ; ; Stenodryas clavigeta clavigeta Bates 1999; ; ; Stenhomalus cleroides Bates 1999; Stenhomalus lighti Gressitt 1999; Stenhomalus takaosanus Ohbayashi 1999; Glaphyra gracilis Hayashi 1982; 1990; 2000

59 ; 1990; Glaphyra kojimai Matsushita 1999; ; Pyrestes nipponicus Hayashi 1990; Chloridolum viridethomson 1989; Aeolesthes chrysothrix Bates Callidiellum rufipenne Motschulsky 1982; 1990; ; Semanotus japonicuslacordaire Phymatodes albicinctus Bates Phymatodes testaceus Linnaeus Dere thoracica White 1969; ; 1989; ; Purpuricenns temminckiig.-meneville 1999; Chlorophorus japonicus Chevrolat ; 2000; Demonax transilis Bates 1982; 1990; ; ; Rhaphuma diminuta Bates 1982; 1990; ; Rhaphuma xeniscabates Grammographus notabilis notabilispascoe Cyrtoclytus caproides caproides Bates Paraclytus excultus Bates ; 1990; ; ; ; Mesosa myops myopsdalman 1999; Mesosa hirsuta hirsuta Bates Mesosa longipennis Bates Asaperda agapanthina Bates ; 1990;

60 ; ; ; Atimura japonica Bates 1999; Microlera ptinoides Bates ; 1990; ; ; ; Cleptometopus bimaculatus Bates 1979; 1982; 1990; ; Egesina bifasciana bifasciana Matsushita Mesosella simiola Bates 1982; 1990; ; 1990; 2000; ; ; Pterolophia angustabates 1999; Pterolophia caudata Bates ; ; Pterolophia granulata Motschulsky ; ; Pterolophia jugosa jugosa Bates ; 1990; ; Pterolophia leiopodinabates 1999; ; ; ; Pterolophia tsugianamatsushita 1999; ; Mecynippus pubicornis Bates 1999; Acalolepta fraudatrix fraudatrix Bates ; 1990; ; ; Acalolepta sejuncta sejuncta Bates Acalolepta luxuriosa luxuriosa Bates 1969; ; 1990; Aoplophora malasiacathomson

61 ; ; 1990; ; Dolichoprosopus yokoyamaigressitt 1973; 1973; 1973; 1984; 1989; 2000; Monochamus grandis Waterhouse 1973; ; Monochamus subfasciatus subfasciatusbates Psacothea hilaris hilarispascoe 1999; Uraecha bimaculata bimaculata Thomson ; ; ; Batocera lineolata Chevrolat 1969; Palimna lituratabates 1969; ; 1973; 1982; 1990; 2000; Rhodopina lewisii lewisiibates ; ; ; 1987; Rhopaloscelis unifasciatus Blessig b; 1990; ; ; Miccolamia cleroides Bates ; ; ; Eupogoniopsis tenuicornis Bates 1990; 2000; Pogonocherus seminiveus Bates 1982; 1990; Rondibilis saperdinabates ; 1990; ; Rondibilis sapporensis Matsushita Exocentrus galloisi Matsushita 1984b; 1990; ; Exocentrus lineatus Bates ; ; ; Exocentrus testudineus Matsushita ; 1990; ;

62 ; ; Exocentrus fasciolatusbates 1999; Exocentrus guttulatus Bates Sciades tonsusbates 1999; Epiglenea comes Bates ; Glenea relicata relicata Pascoe 1969; ; ; Pareutetrapha simulans Bates ; 1990; ; 1982; 1990; Phytoecia rufiventris Gautier 1982; 1990; ; 1990; Nupserha marginella Bates ; 1990; Oberea hebescens Bates 1982; 1990; ; Oberea japonicathunberg 1969; ; Oberea nigriventris Bates ; ; Oberea shirahatai Ohbayashi 1989; Oberea sobosana Ohbayashi 2000; 2005 Chrysomelidae 1 Donacia nitidiornakane c; Plateumaris constricticollis Jacoby c 1987d; ; Plateumaris sericealinnaeus 1987; 1987c; Lema adamsii Baly 1986; ; Lema delicatula Baly 1986; Lema diversa Baly 1986; ;

63 Lema honorata Baly 1999; ; ; Lilioceris rugatabaly 1999; ; ; Lilioceris subpolita Motschulsky 1999; ; Smaragdina auritalinnaeus ; ; Adiscus lewisiibaly 1986; Cryptocephalus aeneoblitus Takizawa 1986; Cryptocephalus approximatus Baly 1969; ; ; ; ; Cryptocephalus fortunatus Baly 1986; ; Cryptocephalus nobilis Kraatz 1999; Cryptocephalus parvulus Muller 1999; Cryptocephalus scitulus Baly 1999; ; ; Chlamisus spilotusbaly 1999; Oomorphoides cupreatusbaly 1969; ; ; ; Acrothinium gaschkevitchii gaschkevitchiimotschulsky 1969; ; ; ; Demotina fasciculata Baly 1999; Syneta adamsi Baly 1973; ; ; Chrysolina aurichalcea Mannerheim 1969; Chrysolina exanthematicawiedemann 1997;

64 65 25 Chrysolina virgatamotschulsky 1986; Chrysomela populi Linnaeus 1986; Chrysomela vigintipunctatascopoli 1969; Gastrilina depressa Baly 1986; Gastrilinnoides japonicusharold 1999; ; ; Gonioctena rubripennis Baly 1986; ; ; ; Agelasa nigriceps Motschulsky 1999; ; ; Agelastica coerulea Baly 1986; Arthrotus niger Motschulsky 1986; ; Atrachya menetriesifaldermann 1986; ; Autacophora femoralismotschulsky 1969; ; ; Autacophora nigripennis Motschulsky 1969; ; Calomicrus cyaneusjacoby 1993; ; ; Chujoa uetsukichujo 1995; ; Clerotilia fiavomarginata Jacoby 1997; ; Exosoma flaviventremotschulsky 1999; Fleutiauxia armatabaly 1969; ; ; Galerucella nipponensis Laboissiere 1987c; Lochmaea capreaelinnaeus 1986; Paridea angulicollismotschulsky 1973; ; ; ; ; ;

65 Paridea quadriplagiatabaly 1986; Pyrrhalta annulicornisbaly 1986; Pyrrhalta semifulvajacoby 1999; ; ; Altica caerulescensbaly 1986; Altica circaeae Ohno 1986; Altica latericostajacoby 1999; ; Altica viridicyaneabaly 1969; Altica oleracealinnaeus 1986; Aphthona perminuta Baly 1999; ; Argopus balyi Harold 1986; ; Argopus clypeatus Baly 1999; Hemipyxis flavipennisbaly 1969; ; ; ; Hemipyxis plagioderoides Motschulsky 1986; ; ; ; Luperomorpha funestabaly 1999; Nonarthra cyanea Baly 1969; ; Phygasia fulvipennisbaly 1986; Pseudodera xanthospila Baly 1969; ; ; ; Sphaeroderma akebia Ohno 1986; Sphaeroderma apicale Baly 1999; Sphaeroderma unicolor Kimoto 1999; Dactylispa masonii Gestro 1969; ; ; Dactylispa subquadratabaly

66 ; ; Rhadinosa nigrocyanea Motschulsky 1999; Aspidomorpha indica Boheman 1969; Aspidomorpha transparipennis Motschulsky 1969; Cassida japana Baly 1986; Cassida versicolorboheman 1986; ; ; Cassida vesperina Boheman 1999; Thlaspida cribrosa Boheman 1986; ; ; ; Thlaspida lewisiibaly 1999; ; 2000 Anthribidae 1 Araecerus tarsalis Sharp 1999; Choragus anobiodes Sharp 1997; Euparius oculatus oculatussharp 1999; ; Ozotomerus japonicus Sharp 1999; Bastiropis nitidicutis Jekel 1999; Sphinctotropis laxus Sharp 1999; Tropideres naevulus Faust 1999; Autotropis basipennis Sharp 1999; Exechesops leucopisjordan 1999; ; Gibber nodulosussharp 1999; Phaulimia aberranssharp 1999; Uncifer truncatus Sharp alt. 450 m2005 Attelabidae 1 Apoderus jekelii Roelofs 1969; 2000

67 ; Apoderus erythrogaster S. van Vollenhoven 1969; ; ; ; Apoderus balteatus Roelofs 1999; ; Apoderus rubidus Motschulsky 1999; ; ; Cycnotrachelus roelofsi Harold 1999; ; ; Paracentrocorynus nigricollis Roelofs 1969; ; Paroplapoderus pardalis S. van Vollenhoven 1969; ; ; Paratrachelophorus longicornis Roelofs 1999; Euops splendidus Voss 1999; ; Byctiscus puberulus puberulus Motschulsky 1999; Aderorhinus crioceroides Roelofs 1999; Involvulus amabilisroelofs 1999; Involvulus pilosusroelofs 1997; Involvulus placidussharp 1997; Neocoenorrhinus assimilis Roelofs 1999; Chonostropheus chujoi Voss 1999; Deporaus unicolor Roelofs 1999; Deporaus mannerheimi Humel 1999; Lasiorhynchites brevirostris Roelofs 2005 Apionidae 1 Cyanapion protractum Sharp alt. 60 m Holotrichapion semisericeumwagner

68 Microconapion collare Schilsky 1999; Microconapion pallidirostreroelofs Pseudopirapion placidumfaust alt. 450 m ; 2000alt. 260 m Curculionidae 1 Phyllobius incomptus Sharp 1999; Phyllobius mundus Sharp 1996; Phyllobius longicornis Roelofs 1969; Phyllobius picipes Motschulsky 1969; ; ; ; Phyllobius annectens Sharp 1997; Phyllobius armatus Roelofs 1969; ; Phyllobius intrusus Kono 1999; Phyllobius subnudus Kono 1999; Cyphicerus viridulus Roelofs 1997; Cyrtepistomus castaneus Roelofs 1997; ; ; ; Macrocorynus elegantulus Roelofs 1999; Macrocorynus griseoides Zumpt 1999; Myllocerus griseus Roelofs 1999; ; ; Episomus turritusgyllenhal 1999; Dermatoxenus caesticollis Gyllenhal 1969; ; ; Eugnathus distinctus Roelofs 1969; ; 2000

69 ; ; Lixus maculatus Roelofs 1999; Mecysolobus erro Pascoe 1999; Mesalcidodes trifidus Pascoe 1969; ; ; Dorytomus maculipennis Roelofs Dorytomus hiripennis Bedel 1999; ; Lissorhoptrus oryzophilus Kuschel 1999; Miarus flavoscutellatus Morimoto 1999; Cionus tamazo Kono 1997; Orchestoides decipiens Roelofs 1997; Orchestoides nipponicus Morimoto 1997; Rhynchaenus sanguinipes Roelofs 1999; Ochyromera suturalis Kojima et Morimoto Anthonomus yuasai Kono Demimaea circula Roelofs alt. 260 m Curculio distinguendusroelofs 2000; Curculio albovittatus Kono 1999; Curculio convexusroelofs 1999; Curculio flavoscutellatus Roelofs 1997; ; Curculio sikkimensisheller 1999; Curculio styracisroelofs 1999; ; ; ; Paracythopeus melancholicus Roelofs 1999; Mecysmoderes ater Hustache 1996; 2000

70 71 39 Mecysmoderes fulvus Roelofs 1997; ; Ceutorhynchus shaowuensis Voss 1999; Wagnerinus costatus Hustache 1999; Wagnerinus harmandi Hustache 1999; Sphinxus crypticus Kojima et Morimoto Metialma cordata Marshall 1999; Kumozo ishikawai Morimoto Carcilia strigicollis Roelofs 1999; Carcilia tenuistriata Heller 1997; Acicnemis dorsonigrita Voss 1999; ; Acicnemis suturalis Roelofs 1999; Kobuzo rectirostrisroelofs 2000; Dyscerus elongatusroelofs 1999; Catagmatus japonicus Roelofs alt. 450 m Cryptorhynchus electus Roelofs 1999; Rhadinopus confinis Voss 1999; Xenomimetes destructor Wollaston 2005 Rhynchophoridae 1 Dryophthorus konishii Morimoto Konishi1963; Dryophthorus japonicus Konishi alt. 560 m Sipalinus gigasfabricius 1999; ; ; 2000 Scolytidae 1 Xyleborus validus Eichhoff 1999; Phloeosinus perlatus Champion Scotytoplatypus tycon Blandford 1999; 2000

71 Pterosticus tsukubasanus Kasahara 12 mm 2 Rhantus erraticus Sharp mm 1986 et Nakane mm 6 Dolichoprosopus yokoyamai Gressitt mm 7 Plateumaris constricticollis Jacoby mm babai 3 Holophorus auriculatus Sharp 4 6 mm Psammoecus sp. 3 mm Psammoecus triguttatus Reitter 5 Pelecotomoides tokejii Nomura : pp pp : Cerambycidae 11: : : : : 43-63

72 pp pp , , : : : : 21 Jaloszynski P. and H. Hoshina Notes on the distribution of Cephennodes vafer Kurbator, 1955 Coleoptera, Scydmaenidaein the mainland of Japan. Elytra, 1: Jaloszynski P. and H. Hoshina Revision of Japanese species of Paraneseuthia FranzColeoptera, Scydmaenidae. Jpn. J. Syst. Ent., 1: : : a1: d1: : Donaciist,1: Konishi M On the Japanese species of the genus Dryophthorus Schonherr, with description of two new species. Insecta Matsumurana, 2: pp / : : : : : : : : a 1 44: b 2 48: Buprestidae9: Buprestidae11: Endomychidae 12: a 166: b Cerambycidae 13: : Chrysomelidae 15: a 5 Adephaga 16: b. 6 Carabidae1 16: c 4 Chrysomelidae 16: d1986 Donaciist1: : : : : : pp : Sasaji H Contribution to the taxonomy of the superfamily Cucujoidea of Japan and her adjacent districts III. Mem. Fac. Edu. Fukui Univ., Ser IINat. Sci., 3: :

73 74 pp : : : : : : :

74 Bull. Ibaraki Nat. Mus.,14: : A List of Coleoptera Recorded in Ukishima Area of Lake Kasumigaura, Inashiki, Ibaraki Prefecture, Central Japan Sadahiro OHMOMO, Keiichi TAKAHASHI and Akira NISHIYAMA Accepted November 20, 2010 Abstract A list of Coleoptera containing 335 species in 65 families is prepared, based on specimens collected in the Ukishima area, which is one of the valuable reed fields left around Lake Kasumigaura in Ibaraki Prefecture. Among these, 32 species in 16 families have so far been recorded only from this area. Key words: Coleoptera, Reed field, Lake Kasumigaura, Ibaraki Prefecture km 2 4 m 7 m Cybister chinensis Motschulsky Cybister tripunctatus orientalis Gschwendtner 1980 * Sumiyoshi, Ami, Inashiki, Ibaraki , Japan ** Shimone-cho, Ushiku, Ibaraki , Japan *** Minami-cho, Warabi, Saitama , Japan

75 ha a: 6 b: 3 c: 4 Fig. 1. Sights of the Ukishima area, in early Junea, mid Marchb, and wooden-path in mid Aprilc Anadastus pulchelloides Nakane %

76 77 a: 7.3 mmb: 2.0 mmc: 3.0 mmd: 4.2 mme: 6.0 mm Fig. 2. Some selected species collected in the Ukishima area. In Ibaraki Prefecture, these species have so far been found in the Ukishima area. a: Bledius salsus MiyatakeBody length 7.3 mm, b: Scymnus nakaikemensis SasajiBody length 2.0 mm, c: Mimemodes emmerichi MaderBody length 3.0 mm, d: Curimopsis japonica NakaneBody length 4.2 mm, e: Anadastus pulchelloides NakaneBody length 6.0 mm. 1 2 Elaphrus punctatus Motschulsky Coelambus chinensissharp Hydraena miyatakei M.SatoTelmatophilus orientalis Sasaji Chrysoline virgatamotschulsky Limnobaris babai Chujo et Morimoto

77 Table 1. Number of species according to families collected in the Ukishima area of Lake Kasumigaura in Ibaraki Prefecture56 families 335 species. 1.Carabidae 44 2.Brachinidae 1 3.Haliplidae 1 4.Dytiscidae 7 5.Hydrophilidae 10 6.Hydraenidae 1 7.Leiodidae 1 8.Silphidae 6 9.Staphylinidae Lucanidae 1 11.Trogidae 1 12.Scarabaeidae Eucinetidae 1 14.Heteroceridae 1 15.Scirtidae 4 16.Byrrhtidae 2 17.Buprestidae 4 18.Throscidae 1 19.Elateridae Demestidae 4 21.Melyridae 3 22.Cleridae 1 23.Anobiidae 2 24.Lampyridae 1 25.Kateretoidae 1 26.Nitidulidae Monotomidae 6 28.Laemophloeidae 1 29.Silvanidae 8 30.Biphyllidae 2 31.Cryptophagidae 7 32.Erotylidae 5 33.Cerylonidae 1 34.Corylophidae 1 35.Endomychidae 6 36.Coccinellidae Phalacridae 2 38.Lathridiidae 5 39.Ciidae 2 40.Mycetophagidae 1 41.Tenebrionidae 7 42.Lagriidae 1 43.Pyrochroidae 3 44.Oedemeridae 3 45.Alleculidae 1 46.Mordellidae 1 47.Melandryidae 2 48.Anthicidae 5 49.Aderidae 3 50.Cerambycidae Chrysomelidae Anthribidae 2 53.Curculionidae Apionidae 4 55.Attelabidae 1 56.Scolytidae 2 Carabidae 1 Campalita chinensekirby 1 ex., 6. IX Elaphrus punctatus Motschulsky 1 ex., 4. IV exs., 3. IV. 2008, 3 Dyschirius batesi Andrewes 1 ex., 18. III. 2006, 4 Dyschirius ordinatus Bates 2 exs., 30. VI. 2006; 2 exs., 10. VII Clivina vulgivaga Boheman 1 ex., 30. VI Demetrias marginicollis Bates 10 exs., 18. III exs., 9. IV. 2006, ; 1 ex., 18. IV. 2009, 7 Bembidion inouyei Habu 1 ex., 18. IV exs., 10. VII Bembidion morawitzi Csiki 1 ex., 10. VII Tachyura fumicata Motschulsky 1 ex., 10. VII Tachyura laetifica Bates 42 exs., 10. VII Paratachys sericans Bates 1 ex., 6. IX ex., 10. VII. 2010

78 Table 2. List of species recorded from only the Ukishima area of Lake Kasumigaura in Ibaraki Prefecture16 families 32 species. 1Bledius salsus Miyatake2-a 2Falagria sapidasharp 3Astenus suffusussharp 4Olophrum vicinum Sharp 5Aphodius rugosostriatus Waterhouse 6Curimopsis japonica Nakane2-d 7Oedosthethus kanmiyai Ohira 8Europs temporis Reitter 9Mimemodes monstrosusreitter 10Mimemodes emmerichi Mader2-c 11Silvanoprus scuticolliswalker 12Silvanoprus cephalotesreitter 13Haptoncus ocularisfairmaire 14Meligethes mikado Reitter 15Anadastus pulchelloides Nakane2-e 16Holoparamecus contractus Wollaston 17Enicmus histrio Toy et Tomlin 18Nephus oshimensis Sasaji 19Scymnus nakaikemensis Sasaji2-b 20Corticaria fasciata Reitter 21Migneauxia lederi Reitter 22Eobia cinereipennismotschulsky 23Mordellina signatellamarseul 24Anthelephila cribricepsmarseul 25Orthocrepis adamsiibaly 26Longitarsus bimaculatusbaly 27Chaetocnema bicolorata Kimoto 28Mantura clavareaui Heikertinger 29Curculio aino Kono 30Tychius picirostrisfabricius 31Phytobius quadricornis Gyllenhal 32Rhinoncus perpendicularisreich 12 Elaphropus latissimusmotschulsky 11 exs., 20. III. 2010, 13 Trechus ephippiatus Bates 1 ex., 20. III. 2010, 14 Lesticus magnusmotschulsky 1 ex., 16. VIII Pterostichus sulcitarsis Morawitz 10 exs., 18. IV. 2009, 16 Platynus suavissimus Bates 20 exs., 10. VII Colpodes elainus Bates 4 exs., 18. IV. 2009, 18 Anisodactylus sadoensis Schauberger 13 exs., 10. VII Anisodactylus signatuspanzer 6 exs., 10. VII Amara chalcites Deiean 2 exs., 10. VII Amara nipponica Habu 1 1, 2. X. 2005, 22 Bradycellus subdituslewis 19 exs., 10. VII Harpalus capito Morawitz 1 ex., 6. IX Harpalus jurecekijedlicka

79 80 13 exs., 6. IX Harpalus eous Tschitscherine 1 ex., 7. VII Harpalus sinicus Hope 6 exs., 7. VII Stenolophus difficilishope 1 ex., 15. III. 2009, ; 6 exs., 7. VII Stenolophus propinquus Morawitz 1 ex., 15. III. 2009, ; 4 exs., 7. VII Stenolophus quinquepustulatuswiedemann 1 ex., 10. VII Acupalpus inornatus Bates 1 ex., 16. VIII. 2009; 2 exs., 18. IV exs., 6. IX exs., 7. VII Anoplogenius cyanescenshope 1 ex., 18. IV. 2009, 32 Platymetopus flavilabris Fabricius 1 ex., 6. IX Dischissus japonicus Andrewes 1 ex., 15 IV. 2007, 34 Chlaenius inops Chandoir 5 exs., 18. IV exs., 6. IX Chlaenius variicornis Morawitz 1 ex., 6. IX Chlaenius micans Fabricius 3 exs., 7. VII Lachnocrepis prolixabates 1 ex., 15. III. 2009, ; 4 exs., 18. IV exs., 7. VII Oodes helopioides tokyoensis Habu 1 ex., 15. III exs., 20. III. 2010, 39 Lebia viridis Say 1 ex., 18. III. 2006, 40 Calleida onoha Bates 1 ex., 24. VI. 2010, 41 Lachnolebia cribricollismorawitz 1 ex., 26. VI. 2010, 42 Drypta japonica Bates 1 ex., 10. VII Odacantha aegrota Bates 3 exs., 18. III. 2006, ; 1 ex., 18. IV ex., 10. VII Odacantha puziloi Solsky 1 ex., 18. III exs., 15. III exs., 20. III. 2010, ; 4 exs., 18. IV. 2009, Brachinidae 1 Pheropsophus jessoensis Morawitz 1 ex., 18. IV. 2009, Haliplidae 1 Peltodytes intermediussharp 1 ex., 30. VI. 2006, Dytiscidae 1 Hydrovatus subtilis Sharp 1 ex., 22. X. 2007, ; 1 ex., 10. VII Coperatus weymarni B.- Browne 1 1, 30. VI , 23. VIII ; 5 exs., 27. VIII ; 2 exs., 10. VII Coelambus chinensissharp 6 exs., 30. VI. 2006; 4 exs., 27. VIII. 2006; 2 exs., 10. VII Hydroglyphus japonicussharp 2 exs., 30. VI. 2006; 1 ex., 16. VIII. 2009; 5 exs., 10. VII. 2010

80 81 5 Rhantus pulverosusstephens 1 ex., 27. VIII ex., 16. VIII ; 2 exs., 6. IX ex., 10. VII Hydaticus grammicusgermar 2 exs., 27. VIII. 2006; 1 ex., 6. IX ex., 10. VII Eretes sticticuslinnaeus 2 exs., 10. VII Hydrophilidae 1 Coelostoma orbiculaefabricius 2 exs., 30. VI exs., 18. IV exs., 20. III. 2010, ; 2 exs., 18. IV. 2009, ; 3 exs., 10. VII Cercyon quisquiliuslinnaeus 6 exs., 10. VII Cercyon laminatus Sharp 18 exs., 10. VII Ampiops mater Sharp 1 ex., 3. IV. 2008, ; 1 ex., 31. IV. 2008, 5 Enochrus subsignatusharold 1 ex., 15. IV exs., 22. X. 2007, 6 Enochrus japonicussharp 4 exs., 22. X exs., 16. VIII ex., 20. III. 2010, ; 1 ex., 6. IX exs., 10. VII Sternolophus rufipesfabricius 6 exs., 27. VIII ex., 16. VIII ; 1 ex., 6. IX exs., 10. VII Hydrochara affinissharp 5 exs., 27. VIII ex., 16. VIII ; 4 exs., 10. VII Berosus signaticollis punctipennis Harold 10 exs., 27. VIII exs., 16. VIII ; 1 ex., 6. IX ex., 10. VII Berosus lewisius Sharp 10 exs., 27. VIII exs., 16. VIII ; 30 exs., 6. IX exs., 10. VII Hydraenidae 1 Hydraena miyatakei M. Sato 2 exs., 18. III ex., 9. IV ex., 20. III. 2010, ; 1 ex., 9. IV exs., 18. IV. 2009, Leiodidae 1 Anisotoma curtaportevin 1 ex., 1. VII. 2007, Silphidae 1 Nicrophorus concolor Kraatz 1 ex., 10. VII Necrodes asiaticus Portevin 1, 10. VII Necrodes nigricornis Harold 2, 10. VII Thanatophilus rugosus Linnaeus 56, 9. IV , 18. IV. 2009, 5 Thanatophilus sinuatus Fabricius 3 2, 9. IV , 18. IV. 2009, 6 Eusilpha japonicamotschulsky 1, 26. VI. 2010, ; 1, 24. VI. 2010, Staphylinidae 1 Megarthrus japonicus Sharp 4 exs., 7. X. 2005, 2 Bledius salsus Miyatake 2 6, 10. VII Bledius curvicornis Sharp 1 ex., 9. IV. 2006, 4 Creophilus maxillosus Linnaeus 3 exs., 9. IV. 2006, 5 Astenus suffusussharp 1 ex., 2. VI exs., 15. III. 2009, 6 Falagria sapida Sharp

81 82 1 ex., 9. IV. 2006, 7 Olophrum vicinum Sharp 1 ex., 15. III. 2009, 8 Paederus fuscipes Curtis 3 exs., 18. IV exs., 10. VII Lathrobium pallipes Sharp 2 exs., 15. III. 2009, 10 Ochthephilum densipenne Sharp 1 ex., 15. III. 2009, ; 1 ex., 18. IV exs., 6. IX Quedius simulans Sharp 4 exs., 6. IX exs., 10. VII Aleochara curtula Goeze 1 ex., 10. VII Lucanidae 1 Prosopocoilus inclinatus Motschulsky 1, 10. VII Trogidae 1 Trox niponensis Lewis 1 ex., 3. IV. 2008, Scarabaeidae 1 Aphodius rugosostriatus Waterhouse 1 ex., 4. VIII. 2005, 2 Aphodius nigrotesscellatus Motschulsky 1 ex., 9. IV. 2006, 3 Aphodius sublimbatus Motschulsky 11 exs., 10. VII Psammodius convexus Waterhouse 1 ex., 9. X ex., 30. VI Trichiorhyssemus asperulus Waterhouse 2 exs., 2. VI ex., 1. VII. 2008, 6 Melolontha frater Arrow 2, 10. VII Apogonia amida Lewis 1 ex., 10. VII Maladera castaneaarrow 4 exs., 6. IX ex., 10. VII Maladera japonicamotschulsky 2 exs., 6. IX Anomala puncticollis Harold 2 exs., 30. VI. 2006; 1 ex., 26. VI. 2010, ; 1 ex., 10. VII Anomala schoenfeldti Ohauss 4 exs., 30. VI Anomala cupreahope 1 ex., 26. VI. 2010, ; 1 ex., 10. VII Anomala lucens Ballion 2 exs., 26. VI. 2010, 14 Anomala rufocuprea Motschulsky 1 ex., 10. VII Mimela splendensgyllenhal 2 exs., 24. VI exs., 10. VII Adoretus tenuimaculatus Waterhouse 1 ex., 24. VI. 2010, 17 Popillia japonica Newmann 1 ex., 26. VI. 2010, ; 2 exs., 24. VI. 2010, 18 Blitopertha orientaliswaterhouse 1 ex., 26. VI. 2010, ; 1 ex., 24. VI ex., 10. VII Protaetia orientalis submarumoreaburmeister 1 ex., 26. VI. 2010, 20 Oxycetonia jucundafaldermann 1 ex., 6. IX ex., 26. VI. 2010, Eucinetidae

82 83 1 Eucinetus haemorrhoidalis Germar 1 ex., 16. VIII. 2009; 1 ex., 10. VII ex., 27. VIII exs., 18. IV exs., 26. VI. 2010, ; 2 exs., 20. VI. 2009, 4 Trachys reitteri Obenberger 1 ex., 4. X. 2009, Heteroceridae 1 Heterocerus fenestratus Thunberg 10 exs., 27. VIII. 2006; 1 ex., 18. IV ex., 6. IX exs., 10. VII Scirtidae 1 Cyphon consobrinus Nyholm Yoshitomi, Cyphon mizoro Nakane 10 exs., 18. III. 2006, 3 Cyphon puncticeps Kiesenwetter 10 exs., 6. IX exs., 10. VII Scirtes japonicus Kiesenwetter 1 ex., 6. IX exs., 10. VII Byrrhidae 1 Curimopsis japonica Nakane 1 ex., 27. VII. 2005, ; 1 ex., 10. VII Syncalypta japonica Nakane 1 ex., 27. VII. 2005, Buprestidae 1 Aphanisticus congener E. Saunders 1 ex., 15. X. 2006, ; 1 ex., 2. VI. 2008, 2 Trachys robusta E. Saunders 6 exs., 9. IV exs., 15. V exs., 22. X. 2007, 3 Trachys tsushimae Obenberger Throscidae 1 Trixagus turgidus Hisamatsu 1 ex., 18. III. 2006, Elateridae 1 Agrypnus binodulusmotschulsky 1 ex., 26. VI. 2010, ; 1 ex., 24. VI exs., 10. VII Agrypnus scrofacandeze 1 ex., 6. IX Aeoloderma agnatacandeze 3 exs., 9. X. 2005, ; 1 ex., 10. VII Lanecarus palustrislewis 1 ex., 9. X ex., 15. III. 2009, 5 Agriotes ograe Lewis 1 ex., 9. X ex., 15. III. 2009, 6 Neotrichophorus juniorcandeze 1, 1. VII. 2008, 7 Melanotus legatus Candeze 2 exs., 10. VII Oedesthethus kanmiyai Ohira 1, 1. VII. 2008, 9 Displatynychus adjutor Candeze 2, 1. VII. 2008, 10 Paracardiophorus pullatus Candeze 1 ex., 1. VII. 2008, Demestidae 1 Dermestes coarctatus Harold 1 ex., 15. V exs., 18. IV. 2009, 2 Thaumaglossa hilleri

83 84 Reitter 1, 15. VI. 2007, 3 Thaumaglossa rufopapillata Redtenbacher 1, 15. X , 2. VI , 1. VII , 6. IX , 26. VI. 2010, 4 Orphinus japonicus Arrow 1 ex., 26. VI. 2010, Melyridae 1 Haplomalachius ishiharai M. Sato et Wittmer 5 exs., 9. IV exs., 15. V exs., 15. IV ex., 12. IV exs., 18. IV. 2009, ; 16 exs., 18. IV. 2009, 2 Laius historio Kiesenwetter 3 2, 9. IV exs., 15. V , 15. IV , 22. X. 2007, ; 1, 23. VIII. 2009, 3 Laius pellegrini Pic 2, 2. VI , 1. VII , 26. VI. 2010, Cleridae 1 Neohydnus hozumii Nakane 7 exs., 27. VIII ex., 2. VI exs., 1. VII exs., 26. VI. 2010, ; 1 ex., 21. V ex., 2. VI. 2008, Anobiidae 1 Ernobius curticollis Pic exs., 2. VI exs., 18. IV exs., 20. III. 2010, 2 Ernobius mollislinnaeus 8 exs., 20. III. 2010, Lampyridae 1 Luciola lateralis Motschulsky 1 ex., 2. VI. 2008, Kateretidae 1 Heterhelus scutellaris Heer 12 exs., 15. IV. 2007, ; 3 exs., 20. III. 2010, Nitidulidae 1 Urophorus humeralis Fabricius 3 exs., 9. X. 2005, 2 Carpophilus chalybeus Murray 2 exs., 9. X ex., 6. IX exs., 26. VI. 2010, ; 1 ex., 24. VI. 2010, 3 Haptoncus ocularis Fairmaire 2 exs., 9. X. 2005, 4 Epuraea parilis Reitter 1 ex., 27. VII. 2005, 5 Meligethes mikado Reitter Meligethes scheklingi Reitter 5 exs., 2. VI exs., 26. VI. 2010, ; 4 exs., 20. VI. 2009, 7 Meligethes violaceus Reitter 1 ex., 15. VI. 2007, 8 Omosita discoidea Fabricius 1 ex., 9. IV. 2006, 9 Omosita colonlinnaeus 1 ex., 18. IV. 2009, 10 Omosita japonica Reitter 1 ex., 18. III. 2006, ; 23 exs., 9. IV exs., 2. VI ex., 18. IV. 2009, 11 Aethina maculicollis Reitter 2 exs., 15. V ex., 15. IV. 2007, Monotomidae 1 Europs temporis Reitter 1 ex., 15. X. 2006, 2 Mimemodes monstrosusreitter

84 85 2, 27. VIII , 22. X , 6. IX , 26. VI. 2010, 3 Mimemodes japonusreitter 21 ex., 22. X. 2007, 4 Mimemodes emmerichi Mader 1 ex., 15. X. 2006, 5 Rhizophagus japonicus Reitter 1 ex., 2. VI. 2008, 6 Monotoma picipes Herbst 1 ex., 27. VII. 2005, Laemophloeidae 1 Xylolestes hillerireitter 1 ex., 20. III. 2010, Silvanidae 1 Silvanus lewisi Reitter 1 ex., 27. VIII Silvanus bidentatus Fabricius 1 ex., 10. VII Silvanoprus scuticollis Walker 1 ex., 23. VIII. 2009, 4 Silvanoprus angusticollis Reitter 1 ex., 15. III. 2009, 5 Silvanoprus cephalotes Reitter 1 ex., 15. III. 2009, 6 Silvanolomus inermis Reitter 1 ex., 20. III. 2010, 7 Psammoecus triguttatus Reitter 3 exs., 18. III exs., 22. X exs., 15. III. 2009, ; 3 exs., 10. VII Psammoecus fasciatus Reitter 2 exs., 18. III. 2006, Biphyllidae 1 Biphyllus lewisireitter 1 ex., 15. V. 2006, 2 Biphyllus rufopictus Wollaston 1 ex., 15. IV. 2007, Cryptophagidae 1 Telmatophilus orientalis Sasaji 1 ex., 2. VI. 2008, ; 2 exs., 24. VI. 2010, 2 Cryptophagus acutangulus Gyllenhal 1 ex., 9. X. 2005, 3 Cryptophagus callosipennis Grouvelle 1 ex., 15. IV. 2007, ; 1 ex., 18. IV. 2009, 4 Atomaria lewisi Reitter 3 exs., 15. V exs., 30. VI exs., 22. X exs., 6. IX. 2009, 5 Atomaria horridula Reitter 1 ex., 9. X ex., 18. III exs., 27. VIII. 2006; 1 ex., 10. VII Henoticus japonicus Nakane et Hisamatsu 1 ex., 2.VI.2008, 7 Curelius japonicusreitter 2 exs., 9. X ex., 30. V I.2006, ; 3 exs., 10. VII Erotylidae 1 Anadastus pulchelloides Nakane 4 exs., 7. X ex., 2. VI exs., 15. III ex., 20. VI. 2009, ; 17 exs., 9. X ex., 18. III ex., 15. V exs., 15. X exs., 22. X exs., 1. VII exs., 15. III. 2009, 2 Anadastus atriceps Crotch 1 ex., 9. X. 2005, ; 7 exs., 20. VIII ex., 9. X ex., 9. IV ex., 15. V ex., 27. VIII. 2006,

85 86 3 Microlanguria jansonicrotch 1 ex., 9. IV ex., 15. V ex., 27. VIII ex., 22. X. 2007, 4 Aulacochilus japonicus Crotch 1 ex., 2. VI. 2008, 5 Episcapha gorhami Lewis 1 ex., 15. V. 2006, Cerylonidae 1 Cautomus hystriculus Sharp 1 ex., 2. VI. 2008, Corylophidae 1 Arthrolips lewisii Matthews 1 ex., 15. V exs., 2. VI. 2008, Endomychidae 1 Holoparamecus contractus Wollaston 1 ex., 9. IV exs., 15. IV. 2007, ; 2 exs., 22. X. 2007, 2 Saula japonica Gorham 1 ex., 16. V exs., 15. IV exs., 22. X exs., 12. IV. 2008, 3 Mycetina ancoriger Gorham 1 ex., 2. VI. 2008, 4 Ectomychus basalis Gorham 1 ex., 4. X. 2005, 5 Lycoperdina mandarinea Gerstaecker 1 ex., 10. VII Ancylopus pictus asiaticus Strohecker 1 ex., 15. III. 2009, ; 1 ex., 18. IV ex., 10. VII Coccinellidae 1 Stethorus japonicus H. Kamiya 1 ex., 15. IV. 2007, 2 Nephus oshimensis Sasaji 1, 9. IV. 2006, 3 Nephus patagiatuslewis 2 exs., 9. X ex., 18. III exs., 27. VIII exs., 15. IV exs., 1. VII ex., 18. IV exs., 6. IX exs., 20. III. 2010, ; 1 ex., 20. III. 2010, ; 1 ex., 18. IV ex., 24. VI. 2010, 4 Nephus phosphoruslewis 1 ex., 20. III. 2010, 5 Scymnus nakaikemensis Sasaji 1 ex., 3. IV. 2008, 6 Scymnus hoffmanni Weise 1 ex., 2. VI ex., 20. III. 2010, ; 5 exs., 15. V exs., 27. VIII exs., 15 IV exs., 15. VI ex., 20. III. 2010, 7 Scymnus kawamuraiohta 1 ex., 18. III exs., 27. VIII exs., 15. IV exs., 22. X ex., 12. IV. 2008, 8 Scymnus japonicus Weise 3 exs., 22. X. 2007, 9 Scymnus dorcatomoides Weise 2 exs., 15. IV ex., 22. X. 2007, 10 Scymnus babai Sasaji 1 ex., 19. X exs., 27. VIII exs., 1. VII exs., 27. VIII. 2008, 11 Scymnus kaguyahime H. Kamiya 1 ex., 24. VI exs., 10. VII Scymnus posticalis Sicard 1 ex., 24. VI. 2010, 13 Pseudoscymnus harejaweise 2 exs., 22. X IV. 2009, 14 Telsimia nigraweise 2 exs., 18. IV ex., 20. III. 2010, ; 3 exs., 4. X. 2009,

86 87 15 Cryptogonus orbiculus Gyllenhal 2 exs., 20. III. 2010, 16 Micraspis kurosai Miyatake 11 exs., 9. X. 2005, ; 5 exs., 9. X exs., 18. III exs., 15. V exs. 27. VIII exs., 15. VI exs., 27. VIII exs., 15. III exs., 18. IV ex., 6. IX exs., 20. III. 2010, ; 6 exs., 18. IV exs., 24. VI ex., 10. VII Hippodamia tredecimpunctatalinnaeus 4 exs., 9. X. 2005, ; 1 ex., 10. VII Anisosticta kobensis Lewis 3 exs., 18. IV ex., 10. VII Propylea japonica Thunberg 1 ex., 27. VIII. 2008; 1 ex., 4. X. 2009, 20 Chilocorus kuwanae Silvestri 1 ex., 24. VI. 2010, 21 Oenopia hirayamaiyuasa 1 ex., 24. VI. 2010, 22 Coccinella septempunctata Linnaeus 1 ex., 6. IX ex., 24. VI. 2010, 23 Harmonia axyridispallas 1 ex., 18. IV exs., 24. VI, exs., 10. VII Epilachna vigintioctomaculata Motschulsky 3 exs., 18. IV. 2009, Phalacridae 1 Heterolitus nipponicus Hisamatsu 2 exs., 26. VI. 2010, ; 1 ex., 10. VII Olibrus consanguineus Flach 16 exs., 10. VII Lathridiidae 1 Enicmus histrio Toy et Tomlin 4 exs., 9. IV ex., 27. VIII. 2006, 2 Corticaria ornata Reitter 6 exs., 18. III ex., 15. V exs., 15.I V ex., 18. IV. 2009, 3 Corticaria japonica Reitter 5 exs., 18. III exs., 15. IV ex., 15. III ex., 6. IX exs., 20. III ex., 26. VI. 2010, ; 1 ex., 10. VII Corticaria fasciata Reitter 2 exs., 20. III. 2010, 5 Migneauxia lederi Reitter 1 ex., 18. IV. 2009, Ciidae 1 Orthocis ornatus Reitter 8 exs., 9. IV ex., 15. IV. 2007, 2 Nipponocis ashuensis Nobuchi 1 ex., 9. X. 2005, Mycetophagidae 1 Mycetophagus antennatus Reitter 1 ex., 30. VI. 2006, Tenebrionidae 1 Gonocephalum japonum Motschulsky 3 exs., 6. IX Alphitobius diaperinus Panzer 1 ex., 9. X. 2005, 3 Palorus subdepressus Wollaston 1 ex., 9. X. 2005, 4 Leiochrinus satzumae Lewis 1 ex., 22. X exs., 18. IV. 2009, 5 Leiochrinus masidai Nakane

87 88 3 exs., 1. VII. 2008, 6 Leiochrinus convexus Lewis 1 ex., 1. VII. 2008, 7 Uloma bonzica Marseul 2 exs., 15. III. 2009, Lagriidae 1 Lagria nigricollis Hope 1 ex., 6. IX. 2009, Pyrochroidae 1 Pseudopyrochroa laticollis Lewis 1 2, 15. IV. 2007, 2 Pseudopyrochroa rufula Motschulsky 1, 15. IV. 2007, 3 Pseudopyrochroa vestiflualewis 1, 18. IV. 2009, Oedemeridae 1 Nacerdes luteipennis Marseul 1 ex., 26. VI. 2010, 2 Eobia cinereipennis Motschulsky 1 ex., 10. VII Oedemera lucidicollis Motschulsky 2 exs., 18. IV. 2009, Alleculidae 1 Borboresthes acicularis Marseul 1 ex., 26. VI. 2010, Mordellidae 1 Mordellina signatella Marseul 1 ex., 27. VIII. 2006, Melandryidae 1 Synstrophus macrophthalmus Reitter 1 ex., 2. VI. 2008, 2 Abdera trisignata Champion 4 exs., 2. VI. 2008, Anthicidae 1 Macratria japonica Harold 4 exs., 27. VIII ex., 26. VI. 2010, ; 1 ex., 18. IV ex., 10. VII Anthelephila cribricepsmarseul 3 exs., 9. IV ex., 6. IX. 2009, 3 Formicomus braminus coiffaiti Bonadona 3 exs., 20. III. 2010, 4 Anthicus watarasensis Sakai et Ohbayashi 1 ex., 18. III ex., 15. IV exs., 12. IV exs., 15. III exs., 18. IV exs., 20. III. 2010, ; 3 exs., 1. VII. 2008, 5 Stricticomus valgipes Lewis 1 ex., 18. III exs., 9. IV ex., 15. V ex., 27. VIII ex., 6. IX ex., 20. III. 2010, ; 1 ex., 18. IV ex., 10. VII Aderidae 1 Pseudoloterus distortus Champion 1, 22. X. 2007, 2 Aderus grouvellipic 1 ex., 18. III exs., 27. VIII ex., 22. X ex., 15. III. 2009, 3 Syzeton brunnidorsis Marseul 1 ex., 1. VII. 2008, Cerambycidae 1 Spondylis buprestoides Linnaeus 2 exs., 10. VII. 2001

88 89 2 Anoplophora malasiaca Thomson 1 ex., 27. VIII. 2006, 3 Batocera lineolata Chevrolat 1, 1. VII. 2008, 4 Ceresium sinicum White 3, 1. VII. 2008, ; 2, 1. VII. 2008, 5 Eupromus ruberdalman 1 2, 1. VII. 2008, 6 Sciades tonsusbates 1 ex., 1. VII exs., 26. VI. 2010, ; 1 ex., 24. VI. 2010, 7 Uraecha bimaculata Thomson 1, 1. VII. 2008, ; 1, 1. VII. 2008, 8 Asaperda agapanthina Bates 2 exs., 9. IV. 2006, 9 Atimura japonica Bates 1 ex., 9. IV ex., 27. VIII exs., 18. IV. 2009, 10 Miccolamia takakuwai Hasegawa et N. Ohbayashi 2 exs., 16. V. 2006, 11 Pterolophia angustabates 1 ex., 21. V. 2007, ; 1 ex., 10. VII Exocentrus lineatus Bates 4 exs., 26. VI. 2010, 13 Oberea shirahatai Ohbayashi 1, 1. VII. 2008, 14 Oberea japonicathunberg 20 exs., 2. VI. 2008, ; 3 exs., 24. VI. 2010, Chrysomelidae 1 Burchidius terreussharp 1 ex., 22. X. 2007, ; 1 ex., 2. VI. 2008, 2 Plateumaris sericealinnaeus 1 ex., 2. VI. 2008, 3 Lema diversa Baly 1 ex., 18. IV. 2009, 4 Lema coronata Baly 1 ex., 10. VII Oulema oryzaekuwayama 2 exs., 15. III ex., 6. IX ex., 26. VI. 2010, ; 1 ex., 18. IV exs., 10. VII Oulema tristisherbst 4 exs., 16. VIII ex., 6. IX Cryptocephalus exiguus Schneider 2 exs., 2. VI. 2008, 8 Cryptocephalus amiculus Baly 4 exs., 24. VI. 2010, 9 Pagria signatamotschulsky 2 exs., 9. X. 2005, ; 2 exs., 18. IV ex., 26. VI. 2010, ; 7 exs., 24. VI. 2010, 10 Scelodonta lewisii Baly 1 ex., 6. IX. 2009, 11 Basilepta fulvipesmotschulsky 2 exs., 26. VI. 2010, ; 4 exs., 24. VI exs., 10. VII Basilepta pallidulumbaly 2 exs., 24. VI. 2010, 13 Plagiodera versicolora Laicharting 2 exs., 18. IV ex., 24. VI. 2010, 14 Gonioctena nigroplagiata Baly 1 ex., 24. VI. 2010, 15 Chrysolina virgatamotschulsky 1 ex., 2. VI. 2008, ; 1 ex., 2. VI ex., 1. VII. 2008, 16 Galerucella nipponensis Laboissiere 1 ex., 2. VI. 2008, 17 Galerucella grisescensjoannis 2 exs., 15. III exs., 16. VIII. 2009, ; 5 exs., 18. IV. 2009, 18 Pyrrhalta humeralischen 1 ex., 26. VI. 2010,

89 90 19 Pyrrhalta semifulvajacoby 13 exs., 24. VI. 2010, 20 Ophraella communa LeSage 1 ex., 15. X. 2006, 21 Medythia nigrobilineata Motschulsky 1 ex., 9. X. 2005, ; 4 exs., 9. IV. 2006, ; 1 ex., 10. VII Atrachya menetriesifaldermann 2 exs., 24. VI. 2010, 23 Altica cyaneaweber 5 exs., 10. VII Lupermorpha funestabaly 1 ex., 9. IV. 2006, 25 Aphthona yuasai Ohno 1 ex., 2. VI. 2008, 26 Batophila acutangula Heikertinger 2 exs., 24. VI. 2010, 27 Longitarsus bimaculatusbaly 1 ex., 9. IV exs., 15. V exs., 18. IV. 2009, 28 Longitarsus holsaticus Linnaeus 1 ex., 10. VII Orthocrepis adamsiibaly 1 ex., 6. IX. 2009, 30 Sphaeroderma apicale Baly 1 ex., 18. IV2009, 31 Chaetocnema ingenuabaly 1 ex., 6. IX. 2009, 32 Chaetocnema bicolorata Kimoto 1 ex., 9.IV.2006, 33 Mantura clavareaui Heikertinger 2 exs., 2. VI. 2008, 34 Cassida japona Baly 1 ex., 10. VII Anthribidae 1 Rhaphitropis guttifer Sharp 1 ex., 16. V. 2006, 2 Uncifer truncatus Sharp 1 ex., 15. IV. 2007, Curculionidae 1 Myllocerus griseus Roelofs 1 ex., 22. X. 2007, 2 Myosides seriehispidius Roelofs 1 ex., 2. VI. 2008, ; 1 ex., 6. IX. 2009, 3 Scepticus griseusroelofs 1 ex., 24. VI. 2010, 4 Pseudocneorhinus bifasciatus Roelofs 1 ex., 24. VI. 2010, 5 Wagnerinus harmandi Hustacke 1 ex., 9. IV. 2006, 6 Lepyrus nebulosus Motschulsky 1 ex., 3. IV. 2008, 7 Curculio aino Kono 1 ex., 2. VI. 2008, 8 Curculio yanoi Morimoto 1 ex., 2. VI. 2008, 9 Limnobaris babai Chujo et Morimoto 2 exs., 18. III exs., 9. IV. 2006, ; 1 ex., 18. IV. 2009, 10 Limnobaris japonica Yoshihara et Morimoto 2 exs., 18. III exs., 9. IV. 2006, ; 2 exs., 21. V ex., 2. VI.2 008, 11 Limnobaris albosparsas Reitter 4 exs., 9. IV exs., 21. V. 2007, ; 1 ex., 21. V exs., 2. VI. 2008, ; 2 exs., 18. IV. 2009, 12 Baris ezoana Kono 1 ex., 20. III. 2010, 13 Baris deplanata Roelofs 1 ex., 26. VI. 2010, 14 Zacladus transversicollisfaust

90 91 1 ex., 9. X. 2005, 15 Lixus maculatus Roelofs 1 ex., 2. VI ex., 26. VI. 2010, ; 1 ex., 2. VI. 2008, 16 Anthonomus bisignifer Schenkling 1 ex., 9. IV exs., 2. VI ex., 26. VI. 2010, 17 Rhinoncomimus niger Chujo et Morimoto 1 ex., 19. X. 2005, 18 Rhinoncus perpendicularisreich 1 ex., 10.V II Phytobius quadricornis roelofsi Hustache 1 ex., 9. IV. 2006, 20 Tychius picirostris Fabricius 1 ex., 2. VI. 2008, 21 Dorytomus maculipennis Roelofs 3 exs., 18. III exs., 9. IV ex., 18. IV IX. 2009, 22 Dorytomus roelofsi Faust 1 ex., 18. IV. 2009, 23 Notaris oryzaeishida 1 ex., 20. III. 2010, 24 Lissorhoptrus oryzophilus Kuschel 2 exs., 9. IV. 2006, 25 Rhynchaenus japonicus Hustache 1 ex., 10. VII Orochlesis takaosanus Kono 1 ex., 22. X. 2007, 27 Stenoscelodes hayashii Kinoshita 1 ex., 2. VI. 2008, 28 Shirahoshizo rufescens Roelofs 1 ex., 26. VI. 2010, 29 Heterarthrus lewisii Wollaston 1 ex., 18. III ex., 9. IV exs., 15. IV. 2007, Apionidae 1 Apion griseopubescens Roelofs 3 exs., 2. VI. 2008, 2 Apion hilleri Schilsky 2 exs., 26. VI. 2010, 3 Apion protractum Sharp 1 ex., 27. VIII. 2006, 4 Nanophyes albovittatus Roelofs 1 ex., 18. III ex., 9. IV. 2006, Attelabidae 1 Auletobius uniformis Roelofs 4 exs., 18. III exs., 9. IV ex., 20. III exs., 26. VI. 2010, ; 1 ex., 18. IV exs., 24. VI. 2010, Scolytidae 1 Xyleborus validus Eichhoff 1 ex., 15. IV. 2007, 2 Cryphalus fulvus Niijima 2 exs., 18. IV. 2009, 2007 Monotomiidae Kanagawa-Chuho,158: : Coleoptera pp : : : Yoshitomi, H Systematic revision of the family Scirtidae of Japan, with phylogency, morphology and bionomics Insecta, Coleoptera, Scirtidae. Jpn. J. Sys. Ent., Monographic Ser.,3, 312 pp.

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92 Bull. Ibaraki Nat. Mus.,14: A Checklist of Bryophytes from Ibaraki Prefecture, Central Japan Koji SUGIMURA, Hisashi KONNO, Nobuya TAKANO, Mihoko UZAWA and Masanobu HIGUCHI (Accepted July 14, 2011) Abstract A checklist of bryophytes from Ibaraki Prefecture, central Japan is presented based on the literature as of the end of December Consequently, a total of 298 species, three subspecies, 14 varieties of mosses belonging to 144 genera and 45 families, 144 species, nine subspecies, seven varieties of hepatics belonging to 64 genera and 33 families, and five species of hornworts belonging to four genera and two families were recorded. Key words: Bryophytes, Ibaraki Prefecture, flora, checklist, literature Paris ; Amakawa and Hattori, 1954; Takaki, 1956; Ando, 1958; Inoue, 1958; Ochi, 1959; Tanegashima Division, Research Center for Medicinal Plant Resources, Matsubarayama, Noma, Nakatane, Kumage, Kagoshima , Japan Graduate School of Science and Engineering, Ibaraki University, Bunkyo, Mito, Ibaraki , Japan Japan Wool Products Inspection Institute Foundation, , Kagoya, Ichinomiya, Aichi , Japan Mito Dai-ichi High School, Sannomaru, Mito, Ibaraki , Japan Ibaraki Nature Museum, 700 Osaki, Bando, Ibaraki , Japan Department of Botany, National Museum of Nature and Science, Amakubo, Tsukuba, Ibaraki , Japan.

93 94 Osada, 1965; Noguchi, 1976; Iwatsuki and Suzuki, ; ; 2007; 2008; ; 2007a ; a2002b b 1995; ; Ahlburg 1879; 1891; 1894; ; 2000; 2009

94 Iwatsuki2004 Yamada and Iwatsuki a,b2009 INM. Sphagnaceae Sphagnum fimbriatum Wilson 2004 Sphagnum microporum Warnst. ex Cardot 2004;, 2010 Sphagnum palustre L. 1981; 2001; 2004; Iwama-cho Higuchi 2007; 2009 Sphagnum squarrosum Crome. 1981; 2004 Sphagnum subsecundum Nees var. junsaiensewarnst. H. A. Crum 2009 Tetraphidaceae Tetraphis pellucida Hedw. 1980; 1981; 2001; 2003; 2004; 2007; 2009 Buxbaumiaceae Buxbaumia minakatae S. Okam Diphysciaceae Diphyscium fulvifolium Mitt. 1980; 1981; 2001; 2002; 2002b; 2003; 2004; 2007; 2008; 2009; 2009; Diphyscium lorifoliumcardotmagombo 2002Theriotia lorifolia; 2002b; Makabe-choMagombo 2003; 2004; Polytrichaceae Atrichum crispulum Schimp. ex Besch Atrichum rhystophyllummüll. Hal.Paris 2003; 2004; 2007;

95 96 Atrichum undulatumhedw.p. Beauv ; 1981; 1988; 2001; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2010; Atrichum undulatum Hedw. P. Beauv. var. gracilisetum Besch. 2007; Atrichum yakushimensehorik.mizush. 1980; 1981; 2004; Pogonatum contortumbrid.lesq. Ibaraki-kenOsada1966; 1981; 2002; 2004; 2007; 2009; Pogonatum inflexumlindb.sande Lac. Ibaraki-kenOsada1966; 1980; 1981; 1988; 2001; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; Pogonatum neesiimüll. Hal.Dozy Ibaraki-kenOsada1966Pogonatum akitense; 1981; 2002; 2004; 2007; Pogonatum nipponicum Nog. et Osada 1980; 1981; 1988; 2004; 2007; 2009 Pogonatum otaruense Besch. 2001; 2002; 2007; Pogonatum spinulosum Mitt. Ibaraki-ken Osada1966; 1980; 1981; 2003; 2007; 2007; Pogonatum urnigerumhedw.p. Beauv. Mt. TsukubaOsada1965; 1981; 2009; Polytrichastrum formosumhedw.g. L. Sm. 1988Polytrichum formosum, Polytrichastrum formosum var. intersedens; 2002b; 2004; 2007; 2009 Polytrichum commune L. ex Hedw. 1981; 2001; 2002; 2004; 2007; 2009; Fissidentaceae Fissidens bryoides Hedw. var. lateralisbroth.z. Iwats et Tad. Suzuki 1981; Ibaraki-kenIwatsuki and Suzuki 1982Map II; 2001;

96 b; 2004; 2007; Fissidens bryoides Hedw. var. ramosissimus Thér Fissidens dubius Mitt. 1980; 1981; Ibaraki-kenIwatsuki and Suzuki 1982Map VIII Fissidens cristatus; 2001; 2002a; 2002b; 2003; 2004; 2007; 2008; 2009; 2009; Fissidens ganguleei Norkett et Gangulee 2004; 2007; Fissidens gardneri Mitt. Kitaibaragi-shi, Satone valleyiwatsuki and Suzuki 1982Fissidens microcladus; 2001Fissidens microcladus; 2002b; 2004; 2007; 2009; Fissidens geminiflorus Dozy et Molk. Takahagi-shi, TakahagiIwatsuki and Suzuki 1982; 2001; 2009; Fissidens geppii M. Fleisch. 2001; 2004 Fissidens grandifrons Brid. 2001; 2002b; 2004; 2007; 2007; 2008; 2009 Fissidens gymnogynus Besch. IbarakiParis1902; 1980; 1981; Takahagi-shi, Hananuki valleyiwatsuki and Suzuki1982; 1988; 2001; 2002; 2002a; 2004; 2007; 2009; Fissidens hyalinus Hook. et Wilson Fissidens involutus Wilson ex Mitt. Mt. Hanazono-yamaIwatsuki and Suzuki1982; 2007 Fissidens linearis Brid. var. obscuriretebroth. et Paris I. G. Stone 2001; 2009Fissidens obscurirete; Fissidens nobilis Griff. 2004; 2009 Fissidens protonemaecola Sakurai 2001; 2002b; 2007 Fissidens taxifolius Hedw. 1981; Ibaraki-kenIwatsukiand Suzuki 1982Map VII; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2009; 2009; Fissidens teysmannianus Dozy et Molk. Ibaraki-kenIwatsuki and Suzuki1982Map

97 98 VII, Fissidens adelphinus; 1980; 1981; 2001Fissidens adelphinus; 2002 Fissidens adelphinus; 2002b; 2003Fissidens adelphinus; 2004; 2005; 2007; 2007Fissidens adelphinus; 2008Fissidens adelphinus; 2009Fissidens adelphinus; 2009; Fissidens tosaensis Mitt. 2001; 2002b; 2004; 2007; 2009; Archidiaceae Archidium ohioense Schimp. ex Müll. Hal. Ditrichaceae Ceratodon purpureushedw.brid. 1988; 2001; 2002; 2004; 2007Tsukuba-shi AmakuboHiguchi 2009; Ditrichum linearesw.lindb. TakahagiMatsui and Iwatsuki1990; 2007 Ditrichum macrorhynchum Broth. ex Cardot 2001 Ditrichum pallidumhedw.hampe 1981; 2001; 2009; Ditrichum rhyncostegium Kindb. Pleuridium japonicum Deguchi, Matsui et Z. Iwats. Tomobe-cho Matsui and Iwatsuki1990; Bryoxiphiaceae Bryoxiphium norvegicumbrid.mitt ssp. japonicum Berggr.A.Löve et D.Löve 1980; 2003; 2004; 2007 Dicranaceae Brothera leanasull.müll. Hal. Mt. Tsukuba Takaki1968; 1981; 1988; 2001; 2002; 2004; 2007; Campylopus gemmiparus Z. Iwats. 2003; 2007 Campylopus japonicus Broth. IbarakiParis1902; 1988; 2001; 2002; 2007; Campylopus umbellatusschwägr. et Gaudid ex Arn. Paris Mt. KabaTakaki1967; 1988; 2001; 2002; 2007; Dichrodontium pellucidumhedw.schimp.

98 Dicranella heteromallahedw.schimp. IbarakiParis1902; 1980; 1981; 1988; 2001; 2002; 2004; 2007; 2007; 2008; 2009; 2010; Dicranodontium denudatumbrid.britt et R. S. Williams 1981; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2009; 2009; Dicranodontium uncinatumharv.a. Jaeger IbarakiParis1902 Dicranoloma cylindrotheciummitt.sakurai IbarakiParis1902Dicranum cylindrothecium; ShiobaraTakaki1966; 1981; 1988; 2001; 2002; 2002a; 2002b; 2004; 2007; 2009; Dicranum caesium Mitt Dicranum flagellare Hedw Dicranum hamulosum Mitt. Ibaraki Paris1902 Dicranum japonicum Mitt. 1981; 2001; 2002; : 2003; 2004,; 2007; 2008; 2009; Dicranum leiodontum Cardot 2002; Dicranum mayrii Broth Dicranum nipponense Besch. 2001; 2003; 2004; 2007; 2009; Dicranum polysetum Sw. 2003; 2007 Dicranum scoparium Hedw. 1981; 2007; 2009 Dicranum viridesull. et Lesq.Lindb. var. hakkodense CardotTakaki 1981; 1988 Oncophorus crispifoliusmitt.lindb. 1980; 1981; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; Mt. TsukubaHiguchi2006; 2007; 2007; 2008; 2009; 2009; 2009; 2010; Oncophorus virenshedw.brid Oncophorus wahlenbergii Brid. IbarakiParis1902

99 100 Rhabdoweisia crispatawith.lindb. 2004; 2007 Trematodon longicollis Michx. 2001; Leucobryaceae Leucobryum bowringii Mitt. 1980; 1981; 2004 Leucobryum glaucumhedw.angstr Leucobryum juniperoideumbrid.müll. Hal. 1980; 1981; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; Calymperaceae Syrrhopodon japonicusbesch.broth Pottiaceae Anoectangium stracheyanum Mitt Anoectangium thomsonii Mitt. 2001; 2002b; 2004; 2007; 2009 Barbula convoluta Hedw Barbula horrinervis K. Saito 2007 Barbula indicahook.spreng. 2001; 2004; 2009; Barbula unguiculata Hedw. Mt. Tsukuba Saito1975; 1981; 2004; 2005; 2007; 2009; Didymodon vinearisbrid.r. H. Zander Mt. MayumiSaito1975Didymodon constrictus; 1988Didymodon constrictus; 2001Didymodon constrictus; 2004Didimodon constrictus; Hyophila involutahook.a. Jaeger Mihara-mura, TsuodaSaito1975; 2007 Hyophila propagulifera Broth. Mt. Mayumi Saito1975; 1981; 1988; 2001; 2002; 2003; 2004; 2007; 2008; 2010; Leptodontium pergemmascens Broth. IwaiSaito1975 Leptophascum leptophyllummüll. Hal.J. Guerra et M. J. Cano 2007; Oxystegus tenuirostrishook. et TaylrA. J. E. Smith 2001Oxystegus cylindricus; 2003; 2004;

100 ; 2009; Scopelophila cataractaemitt.broth. 2001; 2004; 2007; Tortella japonicabesch.broth. 1980; 2004; 2010 Tortella tortuosahedw.limpr Trichostomum platyphyllumbroth. ex IisibaP. C. Chen Mt. MayumiSaito 1975 Tuerckheimia svihlaee. B. BartramR. H. Zander Weisiopsis anomala Broth. et Paris 2001; 2002; 2003; 2004; 2007; 2010; Weissia controversa Hedw. Mt. MayumiSaito1975; 1981; 2001; 2002; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2010; Weissia crispahedw.mitt. Mt. MayumiSaito1975; 2001; 2007; 2009; 2009; Weissia planifolia Dixon 2009 Grimmiaceae Campylostelium brachycarpumnog.z. Iwats 2001Campylostelium saxicola varbrachycarpum; 2004 Grimmia pilifera P. Beauv. Mt. TsukubaDeguchi1978; 1980; 1988; 2001; 2002; 2002b; 2004; 2007; 2007; 2008; 2009; 2009; Ptychomitrium dentatummitt.a. Jaeger 2004; 2007; Ptychomitrium fauriei Besch. SiioDixon 1942Holomitrium ebracteatum; Siino-mura Noguchi1954; 1988; 2003; 2004; 2007; 2009; 2009; Ptychomitrium linearifolium Reimers Mt. HanazonoNoguchi1954; 1980; 1981; 2001; 2002; 2002b; 2004; 2007; 2008; 2009; 2009; 2009; 2010; Ptychomitrium sinensemitt.a. Jaeger

101 ; Ptychomitrium wilsonii Sull. et Lesq. IbarakiParis1902 Racomitrium anomodontoides Cardot 1981; 1988 Racomitrium fasciculare varatroviride; 2002Racomitrium atroviride; 2004Racomitrium atroviride; 2007; 2009; Racomitrium barbuloides Cardot 1988; 2005; 2007; 2007; Racomitrium carinatum Cardot MtTsukubaNoguchi1974; 1981; 1988; 2001; 2002; 2002b; 2004; 2007; 2007; 2009; Racomitrium ericoidesf. Weber ex Brid.Brid. 1981; 1988Racomitriumcanescens varericoides; 2001 Rhacomitrium fascicularehedw.brid Racomitrium heterostichumhedw.brid Racomitrium japonicum Dozy et Molk. 1981; 2001; 2002; 2003; 2007; 2010; Racomitrium laetum Besch. et Cardot Ibaraki-kenNoguchi1974Racomitrium heterostichum vardiminutum; 1988Racomitrium diminutum; 2002; 2002b; 2003; 2007; 2009; Schistidium apocarpumhedw.bruch et Schimp. 1980; 1981 Schistidium liliputanummüll. Hal.Deguchi Mt. TsukubaDeguchi1978; 1988; 2009 Schistidium strictumturnerloeske ex O. Martensson 2001; 2002; 2002b; 2004; 2007; 2009; Erpodiaceae Erpodium sinense Venturi ex Rabh. 2001Venturiella sinensis; 2005; 2007; 2007Ventureiella sinensis; Glyphomitrium humillimummitt.cardot 2001; 2004; 2007; 2007; 2008; 2009; Ephemeraceae Ephemerum spinulosum Bruch et Schimp Micromitrium tenerumbruch et Schimp.Crosby 2001; 2007

102 103 Funariaceae Funaria hygrometrica Hedw. 1980; 1981; 2001; 2003; 2004; 2007; Physcomitrium eurystomum Sendtn. 1981; 2001; 2004; a 2005; 2007; 2010; Physcomitrium japonicumhedw.mitt. IbarakiParis1902Physcomitrium subacuminatum Physcomitrium sphaericumc. F. Ludw.Fürnr. 2007; 2010; Schistostegaceae Schistostega pennatahedw.f. Weber et D. Mohr 1984; 2004 Bryaceae Anomobryum filiformegriff.a. Jaeger 1988; 2001; 2004; 2007; 2008; Brachymenium exiledozy et Molk.Bosch. et Sande Lac. 1988; 2003; 2004; 2007; Brachymenium nepalense Hook. Mt. TsukubaOchi1959Brachymenium nepalense var. clavulum; 2003; 2007; 2009; Bryum argenteum Hedw. Ibaraki-kenOchi 1959; 1980; 1981; 1988; 2001; 2003; a 2005; 2007; 2007; 2008; 2009; Bryum caespiticium Hedw. Ibaraki-kenOchi1959Bryum caespiticium var. rubrolimbatum; 1980; 2003; 2007; Bryum cellulare Hook Bryum cyclophyllumschwägr.bruch et Schimp Bryum pseudotriquetrumhedw.gaertn. 2001; 2002; 2004; Leptobryum pyriformehedw.wilson Pohlia flexuosa Hook. Ibaraki-ken Ochi1959; 1981; 2002; 2004; 2007; 2008; Pohlia nutanshedw.lindb. 2003; 2004; 2005; 2007 Pohlia proligerakindb.lindb. ex Arn. 1981; 2001; 2007;

103 104 Pohlia wahlenbergiif. Weber et D. MohrA. L. Andrews 2007; Rhodobryum giganteumschwägr.paris 1981; 2004; 2009 Rosulabryum capillarehedw.j. R. Spence 1981; 2001Bryum capillare; 2004Bryum capillare; 2007; 2007 Bryum capillare; 2008Bryum capillare; 2009; Mniaceae Mnium heterophyliumhook.schwägr Mnium lycopodioideshook.schwägr. IbarakiParis1902Mnium wichrae; 1980; 2001Mnium laevinerve; 2002; 2002b; 2003; 2004; 2007; 2009; Plagiomnium acutumlindb.t. J. Kop. 1980; 1981; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2009; 2010; Plagiomnium confertidenslindb. et Arn.T. J. Kop. 1988; 2004; 2007; 2009 Plagiomnium maximovicziilindb.t. J. Kop. 1980; 1981; 1988; 2001; 2002a; 2002b; 2003; 2004; 2007; 2007; 2008; 2009; 2010; Plagiomnium succulentummitt.t. J. Kop Plagiomnium vesicatumbesch.t. J. Kop. 1980; 1981; 1988 Plagiomnium rostratum ssp. vesicatum; 2001; 2002b; 2003; 2004; 2007; 2007; 2008; 2009 Rhizomnium striatulummitt.t. J. Kop. 1980; 1981 Rhizomnium tuomikoskii T. J. Kop. Mt. YamizoKoponen1971; 1980; 2004; 2007 Trachycystis flagellarissull. et Lesq.Lindb. 1980; 1981; 2004 Trachycystis microphylladozy et Molk.Lindb. IbarakiParis1902 Rhizogonium microphyllum;

104 ; 1981; 1988; 2001; 2002; 2002b; 2003; 2004; 2007; 2007; 2009; 2009; 2010; Rhizogoniaceae Pyrrhobryum dozyanumsande Lac.Manuel 1980; 1981; 1988Rhizogonium dozyanum; 2001; 2002b; 2003; 2004; 2007; 2007; 2009; 2009 Pyrrhobryum spiniformehedw.mitt. var. badakense M.Fleisch.Manuel 2004 Bartramiaceae Bartramia pomiformis Hedw. 1980; 1981; 2003; 2004; 2007; 2007; 2008; 2009 Philonotis falcatahook.mitt. 2001; 2004; 2007; 2009; Philonotis lancifolium Mitt Philonotis thwaitesii Mitt Philonotis turnerianaschwägr.mitt. 2001Philonotis nitida; 2004; 2007; Philonotis yezoana Besch. et Cardot 2001; 2003; 2007 Orthotrichaceae Macromitrium ferriei Cardot et Thér Macromitrium gymnostomum Sull. et Lesq. IbarakiParis1902Dasymitrium gymnostomum; 2001; 2004; 2005; 2007; 2009 Macromitrium japonicum Dozy et Molk. 2001; 2004; 2007; 2009; Macromitrium prolongatum Mitt. Mt. TsukubaNoguchi1967; 2009 Orthotrichum consobrinum Cardot 2002a; 2004; 2007; 2009; Ulota crispahedw.brid. Mt. TsukubaIwatsuki1959; 1981; 2004; 2007 Ulota japonicasull. et Lesq.Mitt. IbarakiParis1902Ulota nipponensis Fontinalaceae Fontinalis hypnoides Hartm. 2004; 2004; 2008

105 106 Climaciaceae Climacium dendroideshedw.f. Weber et D. Mohr 2004 Climacium japonicum Lindb. 1981; 2001; 2002b; 2004; 2007; 2009; 2009 Hedwigiaceae Hedwigia ciliatahedw.ehrh. ex P. Beauv. 1980; 1988; 2002; 2007; 2009; Hedwigia ciliatahedw.ehrh. ex P. Beauv. var. viridis Bruch et Schimp IbarakiParis 1902 Cryphaeaceae Forsstroemia cryphaeoides Cardot 2007; Forsstroemia japonicabesch.paris 1981 Forsstroemia trichomitriahedw.lindb. 1981; 2004 Pilotrichopsis dentatamitt.besch. Ibaraki Paris1902Pilotrichopsis dentate var. filiformis; 1947; 2009 Leucodontaceae Dozya japonica Sande Lac. 1988; 2007 Leucodon atrovirens Nog. 2000; 2007; 2009 Leucodon luteus Besch Leucodon sapporensis Besch. 2004; 2007; 2009 Trachypodaceae Duthiella speciosissima Broth. ex Cardot 2007 Trachypus humilis Lindb. 2007; 2009 Pterobryaceae Eumyurium sinicummitt.nog. 1947Myurium sinicum; Mt. Tsukuba Iwatsuki1979Myurium sinicum; 2000; 2009 Palisadula chrysophyllacardottoyama Mt. TsukubaIwatsuki1979 Pterobryon arbuscula Mitt. 1981; 1988; 2002; 2002b; 2004; 2009; Meteoriaceae Barbella flagelliferacardotnog. 2001; 2002; 2004; 2007; 2009; 2009; Chrysocladium retrorsummitt.m. Fleisch. 1948; Mt. TsukubaNoguchi 1976; 2000; 2004; 2009 Meteorium buchananiibroth.broth. ssp. helminthocladulum CardotNog. 2000; 2007; 2009 Meteorium subpolytrichumbesch.broth Meteorium helminthocladum; Mt. TsukubaNoguchi1976; 2000; 2007; 2009

106 107 Neckeraceae Bissetia lingulatamitt.broth. 1988; 2007 Homalia trichomanoideshedw.bruch et Schimp. var. japonicabesch.s. He 2004 Homaliadelphus targionianusmitt.dixon et P. Vard Homaliodendron flabellatumsm.m. Fleisch. IbarakiParis1902Homalia scalpellifolia; 1988Homaliodendron scalpellifolium; 2000; 2009; 2009 Neckera fauriei Cardot 2009 Neckera flexiramea Cardot 2000; 2009 Neckera humilis Mitt. 2001; 2004; 2007; Neckera konoi Broth. ex Cardot 1988 Neckera yezoana Besch. 1981; 2000; 2009; 2009 Neckeropsis nitidulamitt.m. Fleisch. 2001; 2002; 2002a; 2002b; 2003; 2004; 2007; 2007; 2009; 2009; 2009; Thamnobryum plicatulumsande Lac.Z. Iwats. 2001; 2002a; 2002b; 2003; 2004; 2007; 2009; 2009 Thamnobryum subseriatummitt. ex Sande Lac.B. C. Tan 1980; Thamnobryum sandei; 2001; 2002; 2003; 2004; 2007; 2007; 2009; 2009; 2009; Lembophyllaceae Dolichomitra cymbifolialindb.broth. 2002b; 2007; 2009 Dolichomitra cymbifolia Lindb. Broth. var. subintegerrima S. Okam Dolichomitriopsis diversiformismitt.nog. 1980; 1981; 1988; 2002; 2007; 2009; Isothecium subdiversiforme Broth. 1988; 2001; Mt. TsukubaHiguchi2002; 2002; 2002a; 2002b; 2003; 2004; 2007; 2009; 2009; Hookeriaceae Distichophyllum maibarae Besch. 2001; 2004 Hookeria acutifolia Hook. et Grev. 2001; 2002b;

107 ; 2007; 2009; 2009 Theliaceae Fauriella tenuismitt.cardot 1981; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2007; 2007; 2009; 2010; Fabroniaceae Fabronia matsumurae Besch. 2007; 2009; Schwetschkeopsis fabroniaschwägr.broth. 1988; 2001; 2002; 2002a; 2003; 2004; 2007; 2008; 2009; 2010; Leskeaceae Leskeella pusillamitt.nog. Ibaraki-shi Noguchi1972; 2007; Okamuraea brachydictyoncardotnog Okamuraea hakoniensismitt.broth. 2004; 2007 Pseudoleskeopsis zippeliidozy et Molk.Broth. 1980; 2002; 2002b; 2004; 2007; 2009; 2009; Thuidiaceae Anomodon abbreviatus Mitt. 2004; 2007; 2009 Anomodon giraldii Müll. Hal. 1980; 1981; 2001; 2004; 2007; 2009; Anomodon minorhedw.lindb. IbarakiParis1902Anomodon ramulosus; 1981; 2003Anomodon minor ssp. integerrimus; 2004 Anomodon minor sspintegerrimus; 2007; 2009Anomodon minor ssp. integerrimus; Anomodon rugeliimüll. Hal.Keissl. 2003; 2004; 2007 Anomodon thraustus Müll. Hal. 2003; 2007 Anomodon viticulosushedw.hook et Tayl. 2007; 2009 Boulaya mitteniibroth.cardot 1981; 2002a; 2004; 2007; 2009; Claopodium aciculumbroth.broth. 2001; 2002; 2004; 2007; Claopodium gracillimumcardot et Thér.Nog Claopodium prionophyllummüll. Hal.Broth. 1980; 2004 Haplocladium angustifoliumhampe et Müll. Hal.

108 109 Broth. 1980; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2009; 2009; Haplocladium microphyllumhedw.broth. IbarakiParis1902Thuidium capillatum; 1988; 2001; 2003; 2005; 2007; 2007; 2008; 2009; 2009; 2010; Haplocladium perparaphyllum R. Watan. 2001; 2002; Haplocladium strictulumcardotreimers 2004; 2007; 2009 Haplohymenium longinervebroth.broth. 1981; 1988; 2001; 2002; 2002b; 2004; 2007; 2009; 2009; Haplohymenium pseudo-tristemüll. Hal.Broth ; 2002; 2002a; 2002b; 2004; 2007; 2008; 2009; 2009; 2009; Haplohymenium sieboldiidozy et Molk.Dozy et Molk Haplohymenium tristeces.kindb. 2001; 2004; 2007; 2009; 2009; 2010; Herpetineuron toccoaesull. et Lesq.Cardot 1981; 1988; 2001; 2002; 2003; 2004; 2007; 2009; 2009; Hylocomiopsis ovicarpabesch.cardot 2007 Miyabea fruticellamitt.broth. 1988; 2001; 2004; 2007; 2009 Pelekium versicolormüll. Hal.Touw IbarakiParis1902Thuidium fauriei; 1981; 2001Thuidium sparsifolium; 2002b; 2004 Thuidium sparsifolium; 2007; 2009;

109 110 Thuidium cymbifoliumdozy et Molk.Dozy et Molk. 1980; 2001; 2002; 2002b; 2004; 2007; 2008; 2009; 2010; 2009; Thuidium delicatulumhedw.schimp. 1980; 1981; 2004Thuidium recognitum var. delicatulum; 2007 Thuidium kanedae Sakurai 1980; 1981; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2009; 2009; 2010; Thuidium pristocalyxmüll. Hal.A. Jaeger 1980; 1981; 2001; 2002; 2002b; 2004; 2007; 2009; Thuidium subglaucinum Cardot 2007; 2009 Amblystegiaceae Amblystegium variumhedw.lindb. 1988Hygroamblystegium varium; 2010 Calliergonella cuspidatahedw.loeske 2004; 2007; 2009 Campyliadelphus chrysophyllusbrid.r. S. Chopra 2003; 2004; 2007; 2009 Cratoneuron filicinumhedw.spruce IbarakiParis1902Amblystegium filicinum; 2004; 2007; 2008; 2009; Drepanocladus polygamusbruch et Schimp.Hedenäs TakahagiKanda1975 Campyliadelphus polygamus Leptodictyum bandaiensetakakikanda NakaminatoIsozakiKanda 1975 Leptodictyum humilep. Beauv.H. A. Crum 2001 Leptodictyum ripariumhedw.warnst. 2005; 2009; Sasaokaea aomoriensispariskanda TakahagiDrepanocladus japonicusdixon 1936; TakahagiKanda1976; 2009; 2010 Brachytheciaceae Brachythecium buchananiihook.a. Jaeger 2002; 2003; 2005; 2007; 2007; 2008; 2009; Brachythecium buchananiihook.a. Jaeger var. gracillimum Dixon Mt. Tukuba

110 111 Dixon1932 Brachythecium coreanum Cardot 1981; 2003; 2005 Brachythecium garovaglioides Müll. Hal Brachythecium helminthocladum Broth. et Paris 2001; 2004; 2009; Brachythecium plumosumhedw.bruch et Schimp. 1980; 1981; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2007; 2008; 2009; 2009; 2009; Brachythecium populeumhedw.bruch et Schimp. 2001; 2002; 2003; 2007; 2009; Brachythecium rivulare Bruch et Schimp. 2004; 2007 Bryhnia novae-angliaesull. et Lesq.Grout 2001; 2003; 2004; 2007; Bryhnia tennerrima Broth. et Yasuda 2007; 2008; 2009 Homalothecium laevisetum Sande Lac. 1988; 2001; 2007 Kindbergia arbusculabroth.ochyra 2009 Kindbergia praelongahedw.ochyra 1988Stokesiella praelonga Myuroclada maximowicziiborszcz.steere et W. B. Schofield 1980; 1981; 2003; 2004; 2007; 2007; 2009; 2009; Oxyrrhynchium hianshedw.loeske 1988Eurhyncium hians; 2001 Eurhyncium hians; 2003Eurhyncium hians; 2004 Eurhyncium hians; 2005; 2007; 2007Eurhyncium hians; 2008Eurhyncium hians; 2009; 2009Eurhyncium hians; 2010Eurhyncium hians; Oxyrrhynchium savatierischimp. ex Besch.Broth. 2001Eurhynchium savatieri; 2002bEurhynchium savatieri; 2003Eurhynchium savatieri; 2004Eurhynchium savatieri; 2007; 2009; Palamocladium leskeoideshook.e. G. Britton 2001 Palamocladium nilgheriense; 2004 Rhynchostegium contractum Cardot

111 Rhynchostegium inclinatummitt.a. Jaeger 1980; 2001 Rhynchostegium pallidifoliummitt.a. Jaeger IbarakiParis1902; MitukaidoTakaki 1956; 1980; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2009; 2009; Rhynchostegium riparioideshedw.cardot 1981; 2001Platyhypnidium riparioides; 2002b; 2003; 2004; 2007; 2007; 2008; 2009; 2010; Entodontaceae Entodon challengeripariscardot 1980; 1981; 2001; 2002; 2003; 2004; 2005; 2007; 2007; 2008 ; 2009; Entodon diversinervis Cardot 2001; 2009 Entodon flavescenshook.a. Jaeger 1988Entodon rubicundus; 2004; 2007; 2009; Entodon luridusgriff.a. Jaeger Mito-shiOkamura1915Entodon andoi Entodon scabridens Lindb Entodon sullivantiimüll. Hal.Lindb. IbarakiParis1902Entodon attenuatus; 1988 Entodon sullivantii var. versicolor; 2001Entodon sullivantii varversicolor; 2002; 2002b; 2003; 2004; 2005; Hanazono-keikokuHiguchi2006; 2007; 2007; 2008; 2009; 2010; Plagiotheciaceae Plagiothecium cavifoliumbrid.z. Iwats. Berg TsukubaReimers and Sakurai1931; Mt. TsukubaIwatsuki1970Plagiothecium sakuraii; 1988; 2008; 2009 Plagiothecium euryphyllumcardot et Thér.Z. Iwats. Mt. TsukubaIwatsuki 1970Plagiothecium neckeroideummt. Tsukuba Sasaoka s Exsiccatano. 68, det. by Sakurai as P. neckeroideum; 1981;

112 ; 2001; 2002; 2003; 2004; 2007; 2007; 2008; 2009; Plagiothecium nemoralemitt.a. Jaeger 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2007; 2007; 2008; 2009; 2009 ; 2010; Sematophyllaceae Brotherella complanata Reimers et Sakurai 1981 Brotherella faurieicardotbroth. 2002; Brotherella henoniidubym. Fleisch 1980; 1981; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2010; Pylaisiadelpha tenuirostrisbruch et Schimp. ex Sull. W. R. Buck 1980; 1981; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2009; 2010; Pylaisiadelpha yokohamaebroth.w. R. Buck 1988Brotherella yokohamae; 2009 Sematophyllum subhumilemüll. Hal.M. Fleisch. 2003; 2007; Hypnaceae Breidleria erectiusculasull. et Lesq.Hedenäs 2001Hypnum erectiusculum; 2007 Callicladium haldanianumgrev.h. A. Crum 2001; 2002a; 2003; 2004; 2007; 2007; 2008 Ctenidium capillifoliummitt.broth. Tsukuba-sanNishimura1985; 2002b;

113 ; 2007; 2009; 2009 Ctenidium hastilemitt.lindb. Tsukuba-sanNishimura1985; 1988; a 2005 Ctenidium percrassum Sakurai 2001; 2002b; 2004; 2007; 2009 Eurohypnum leptothallummüll. Hal.Ando 2007 Glossadelphus ogatae Broth. et Yasuda 1988; 2002; 2007; 2008; 2009; Gollania ruginosamitt.broth Herzogiella perrobustabroth. ex CardotZ. Iwats. 1980; 2007; 2007 Homomallium connexumcardotbroth. Mt. TsukubaAndo1964 Hypnum densirameum Ando 2001 Hypnum fujiyamaebroth.paris 1988 Hypnum oldhamiimitt.a. Jaeger et Sauerb. Mt. TsukubaAndo1958; 1980; 1981; 2001; 2002; 2003; 2007; 2009; 2009; Hypnum plicatulumlindb.a. Jaeger et Sauerb 2002; Hypnum plumaeforme Wilson Mt. Tsukuba Ando1958; 1980; 1981; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2010; 2009; Hypnum plumaeforme Wilson var. minus Broth. ex Ando Mt. TsukubaAndo1958; 2001; 2007; Hypnum sakuraiisakuraiando IbarakiParis1902Stereodon rhynchothecius; Mt. TsukubaAndo1958Hypnum sakuraii var. venustum; 1988; 2001; 2002; 2004; 2007; Hypnum subimponens Lesq. subsp. ulophyllummüll. Hal.Ando Hypnum tristo-viridebroth.paris 1980; 1981; 1988; 2001; 2002; 2002b;

114 ; 2004; 2007; 2009; 2009; Pseudotaxiphyllum densumcardotz. Iwats. Pseudotaxiphyllum pohliaecarpumsull. et Lesq.Z. Iwats. 1980; 1988 Isopterygium pohliaecarpum; 2001; 2002; 2002b; 2003; 2004; 2005; 2007; 2007; 2008; 2009; 2009; Pylaisia brotheri Besch Pylaisia nana Mitt. Mt. Tsukubasan Higuchi1999 Pylaisia subcircinata Cardot 2001 Rhytidiadelphus japonicusreimerst. J. Kop. 2001; 2004; 2007 Taxiphyllum aomoriensebesch.z. Iwats Taxiphyllum alternanscardotz. Iwats. 2003; 2004; Taxiphyllum arcuatumbosch et Sande Lac.S. He 2004 Taxiphyllum taxirameummitt.m. Fleisch. 1980; 1988; 2001; 2002; 2002a; 2002b; 2003; 2004; 2005; 2007; 2008; 2009; 2010; Hylocomiaceae Hylocomium brevirostrebrid.bruch et Schimp. var. cavifoliumsande Lac.Nog. 1981; 2002b; 2003Loeskeobryum cavifolium; 2004Loeskeobryum cavifolium; 2007; 2009 Hepaticopsida Herbertaceae Herbertus aduncusdicksgray 1988; 1995; 1996; 2002; 2004; Pseudolepicoleaceae Blepharostoma minus Horik. 1995; 1996; 2002b; 2003; 2004; 2007; 2009;

115 116 Trichocoleaceae Trichocolea tomentellaehrh.dumort. 1996; 2004 Lepidoziaceae Bazzania denudatatorreytrevis. 1995; 1996; 2004; Bazzania pompeanasande Lac.Mitt. 1996; 2001; 2004; 2007; 2009; 2009 Bazzania tridensreinw., Blume et NeesTrevis. 1995; 1996; 2001; 2002; 2002a; 2002b; 2004; 2007; 2007; 2009; 2009; Kurzia makinoanasteph.grolle 1995; 1996; 2001; 2004; Lepidozia vitrea Steph. 1995; 1996; 1998; 2001; 2004; 2007; 2009; Calypogeiaceae Calypogeia arguta Nees et Mont. 1995; 1996; 2001; 2004; a2005; 2007; 2008; Calypogeia azurea Stotler et Crotz. 1995; 1996; 2001; 2004; 2007; Calypogeia japonica Steph. 1995; 2001; Calypogeia tosanasteph.steph. 1995; 1996; 2001; 2004; 2007; 2007; 2009; Metacalypogeia cordifoliasteph.inoue 1995; 1996; 2001; 2002; 2004; Kita-ibaraki-shiHiguchi2007; Cephaloziaceae Alobiellopsis parvifoliasteph.r. M. Schust. 2002; 2007; Cephalozia otaruensis Steph. 1995; 1996; 2001; 2002; 2003; 2004; 2007; 2008; 2009; 2009;

116 117 Nowellia curvifoliadicks.mitt. 1995; 1998; 2009; Odontoschisma denudatummart.dumort. 1998; 2004; 2004; 2007; 2009; 2009 Schiffneria hyalina Steph Cephaloziellaceae Cephaloziella kiaeriaustindouin 1988; 1995; 2007; Cephaloziella microphyllasteph.douin 1988; 1996; 2001; 2004; 2007; Cephaloziella spinicaulis Douin 1995; 1996; 2001; 2002a; 2004; 2007; 2008; 2009; Jungermanniaceae Anastrophyllum minutumschreb.r. M. Schust. 1995; 2001; Jamesoniella autumnalisdc.steph, 1988 Jungermannia atrovirens Dumort. 1996; 2004 Jungermannia exsertifolia Steph. 1996; 2004 Jungermannia fusiformissteph.steph. 1995; Jungermannia infuscamitt.steph. 1995; 1996; 2001; 2002b; 2003; 2004; 2007; 2007; 2009; Jungermannia rubripunctatas. Hatt.Amakawa Jungermannia subulata A. Evans 2002b; 2004; 2007; 2009 Jungermannia truncata Nees 2001; 2004; 2007; Jungermannia virgatamitt.steph. 1995; 1996; 2001; 2004; Nardia assamicamitt.amakawa 1995; 1996; 2001; 2003; 2004; 2007; Plicanthus birmensissteph.r. M. Schust. 1988; 1995; 2002; Marsupellaceae Marsupella emarginataehrh.dumort. ssp. tubulosa

117 118 Steph.N. Kitang. 1988; 1995Marsupella emarginata; 1996Marsupella emarginata; 1998; 2001Marsupella emarginata; 2002; 2002b; 2004; 2009; Marsupella emarginataehrh.dumort. var. patens N. Kitang Marsupella pseudofuncki S. Hatt. 1995; 1996; 2004; Marsupella revolutaneesdumort. 1995; Scapaniaceae Diplophyllum albicansl.dumort. 1988; 1996; 2002; 2004; Diplophyllum andrewsii A. Evans 1995; 1996; 2001; 2004; Diplophyllum serrulatummüll. Frib.Steph. 1995; 1996; 2001; 2004; 2010; 2007; Diplophyllum taxifoliumwahlenb.dumort. 1996; 2004 Scapania ampliata Steph. 1998; 2009 Scapania ciliata Sande Lac. Mt. TsukubaAmakawa and Hattori1954Scapanica spinosa; 2009 Scapania ligulata Steph. 1988; 1995; 1996; 2001; 2002; 2002b; 2004; 2007; 2008Scapania stephanii; Scapania parvitexta Steph 1996; 1998; 2004; 2009 Scapania undulatal.dumort 1996; 2004 Geocalycaceae Chiloscyphus polyanthosl.corda 1995; 1996; 2001; 2003; 2004; Iwama-cho Higuchi2005; 2007; 2007; 2009; Geocalyx lancistipulussteph.s. Hatt 1995; 1996; 2001; 2004; 2007; Heteroscyphus argutusreinw., Blume et Nees Schiffn 2001; 2004; 2005; 2007 Heteroscyphus coalitushook.schiffn 1995; 1996; 2001; 2003; 2004; 2007; 2008; 2009;

118 119 Heteroscyphus planusmitt.schiffn 1995; 1996; 2001; 2003; 2004; 2007; 2009; Lophocolea heterophyllaschrad.dumort 1995; 1996; 2001; 2003Chiloscyphus profundus; 2004Chiloscyphus profundus; 2005; 2007; 2010 Chiloscyphus profundus; Lophocolea minor Nees 1988; 1995; 1996; 2001; 2002Chiloscyphus minor; 2002a; 2002b; 2003Chiloscyphus minor; 2004Chiloscyphus minor; 2005; 2007; 2008; 2009; Plagiochilaceae Pedinophyllum truncatumsteph.inoue 2001; 2007 Plagiochila orbiculariss. Hatt.S. Hatt 1988; Mt. Tsukuba Higuchi2000 Plagiochila ovalifolia Mitt. 1995; 1996; 1998; 2002; 2002b; 2004; 2004; 2007; 2008; 2009; 2009; 2009; 2010; Plagiochila parvifolia Lindenb. Mt. TsukubaInoue1958Plagiochila yokogurensis; 1998; 2009; Plagiochila porelloidestorreylindenb 1995; 1996; 2002b; 2004; Plagiochila sciophila Nees 1995; 1996; 2001; 2002; 2002a; 2002b; 2003; 2004; 2007; 2007; 2008; 2009; 2009; 2009; 2010; Plagiochila semidecurrenslehm. et Lindenb. Lindenb 2009 Radulaceae Radula constricta Steph 1988; 1995;

119 ; 2001; 2002; 2002a; 2002b; 2004; 2004; 2005; 2007; 2008; 2009; Radula japonica Gottsche 1995; 1996; 2001; 2002; 2002a; 2002b; 2004; 2004; 2007; 2009; Radula kojana Steph 1996; 2004; 2007 Radula obtusiloba Steph 1996; 2004 Radula oyamensis Steph 2002; Radula perrottetii Gottsche 2004 Radula tokiensis Steph 1996; 2004; 2008 Lepidolaenaceae Trichocoleopsis sacculatamitt.s. Okamura 1995; 2001; 2002; 2004; Porellaceae Macvicaria ulophyllasteph.s. Hatt 1995; 1996; 1998; 2001; 2002; 2002a; 2002b; 2004; 2007; 2007; 2008; 2009; 2009; 2010; Porella acutifolialehm. et Lindenb.Trevis 2004 Porella acutifolialehm. et Lindenb.Trevisssp tosanasteph.s. Hatt 1988Porella campylophylla Porella caespitans S. HattvarcordifoliaSteph. S. Hatt 1995; 1996; 1998; 2002b; 2003; 2004; 2007; 2009; 2009; Porella caespitans S. Hattvarnipponica S. Hatt 1988 Porella densifoliasteph.s. Hatt. var. fallaxc. Massal.S. Hatt 1995; 1998; 2002b; 2007; 2009; 2009; Porella grandiloba Lindb 1996; 2001; 2002a; 2004; 2004; 2007; 2009; 2009; Porella japonicasande Lac.Mitt

120 ; 1998; 2002a; 2002b; 2003; 2004; 2004; 2007; 2009; 2009; 2009 Porella perrottetianamont.trevis 1996; 1998; 2004; 2004; 2007; 2009 Porella stephanianac. Massal.S. Hatt 2007 Porella vernicosa Lindb 1988; 1996; 2001; 2002b; 2004; 2004; 2007; 2009 Frullaniaceae Frullania cristata S. Hatt 1996; 2004 Frullania davurica Hampe 1988; 1995; 1998; 2007; 2009; Frullania diversitexta Steph 1995; 1996; 2001; 2004; 2007; Frullania ericoidesneesmont 1996; 1998; 2001; 2004; 2007; 2009; Frullania fauriana Steph 1995; 1996; 2004; Frullania hamatiloba Steph 1988; 1995; 1996; 2001; 2004; 2004; Frullania inflata Gottsche 1995; 1996; 2002a; 2004; 2007; Frullania kagoshimensis Steph 1995; 2001; 2003; 2004; 2007; Frullania monocerahook.fet TaylorTaylor 2007 Frullania muscicola Steph 1988; 1995; 1996; 1998; 2001; 2002a; 2004; 2004; 2005; 2007; 2007; 2008; 2009; 2009; 2010; Frullania muscicola StephvarinuenaSteph. Kamim 2007 Frullania osumiensiss. Hatt.S. Hatt

121 ; 2004 Frullania parvistipula Steph 1996; 2004; 2007; Frullania pedicellata Steph 1995; 2001; 2007; Frullania tamarisci sspobscuraverd.s. Hatt 1988; 1995; 1996; 1998; 2001; 2002a; 2004; 2004; 2007; 2009; 2009; Frullania taradakensis Steph 1996; 1998; 2009 Frullania usamiensis Steph 1988; 1996; 1998; 2004; 2004; 2009 Jubulaceae Jubula hutchinsiaehook.dumortsspjavanica Steph.Verd 1995; 1996; 2001; 2002b; 2004; 2004; 2007; 2008; 2009; 2009; 2010; Jubula japonica Steph. 1988; 2001; 2004; 2007; 2009 Lejeuneaceae Acrolejeunea pusillasteph.grolle et Gradst 1995; 1996; 2001; 2004; 2007; 2008; Cheilolejeunea khasianamitt.n. Kitag 1998; 2009 Cheilolejeunea nipponicas. Hatt.S. Hatt Cheilolejeunea obtusifolias. Hatt.S. Hatt 1996; 2004 Cheilolejeunea trapezianeesr. M. Schustet Kachroo 1996; 2004 Cololejeunea japonicaschiffn.s. Hatt 1995; 1996; Tsukuba-shi Amakubo Higuchi2001; 2001; 2002a; 2002b; 2004; 2005; 2007; 2009; 2009; 2010; Cololejeunea kodamae Kamim 1996; 2004 Cololejeunea minutissimasm.schiffn

122 ; 2004; 2007 Cololejeunea spinosa Pande et Misra 1996; 2004 Cololejeunea subkodamae Mizut 1995; 1996; 2001; 2002a; 2002b; 2004; 2007; 2009; Dicranolejeunea yoshinaganas. Hatt.Mizut 2007 Drepanolejeunea angustifoliamitt.grolle 2004 Lejeunea aquatica Horik 2001; 2007; 2008; 2009; 2010 Lejeunea compactasteph.steph 2002b; 2007 Lejeunea discreta Lindenb 1995; 1996; 1998; 2001; 2002a; 2004; 2007; 2009; Lejeunea flavasw.nees 1995 Lejeunea japonica Mitt. 1988; 1995; 1996; 1998; 2001; 2002; 2002a; 2002b; 2004; 2004; 2005; 2007; 2008; 2009; 2009; 2009; 2010; Lejeunea kodamae Ikegami et Inoue 1995; Lejeunea parvas. Hatt.Mizut 1995; 1996; 2002; 2002b; 2004; 2007; 2009; Lejeunea ulicinataylorgottsche 1995; 1996; 2001; 2002; 2002a; 2002b; 2004; 2004; 2007; Leptolejeunea ellipticalehmet Lindenb.Schiffn 2001 Leucolejeunea japonicahorik.verd 1995; 1996; 2001; 2002a; 2004; 2007; 2009; Nipponolejeunea piliferasteph.s. Hatt 1988; 1995; 1996; 1998; 2001;

123 ; 2002a; 2002b; 2004; 2004; 2007; 2009; Ptychanthus striatuslehm. et Lindenb.Nees 2007; 2009 Trocholejeunea sandvicensisgottschemizut 1998; 2001; 2005; 2007; 2009; Fossombroniaceae Fossombronia foveolata LindbvarcristulaAustin R. M. Schust Tsukuba-shi AmakuboHiguchi2001; 2003; 2007; Pelliaceae Pellia endiviifoliadicks.dumort 1988; 1995; 1996; 2001; 2002b; 2003; 2004; 2007; 2007; 2008; 2009; 2009; 2010; Allisoniaceae Calycularia crispula Mitt 1988; 1995; 1996; 2004; Makinoaceae Makinoa crispatasteph.miyake 1995; 1996; 2001; 2003; 2004; 2007; 2007; 2009; 2010;. Pallaviciniaceae Pallavicinia levieri Schiffn Pallavicinia subciliataaustinsteph 1996; 2003; 2004; 2007; 2007; 2008; Blasiaceae Blasia pusilla L 2001; Aneuraceae Riccardia glauca Furuki Riccardia multifidal.gray sspdecrescrnssteph. Furuki 2001; 2003; Metzgeriaceae Metzgeria conjugata Lindb 1995; 1996; Metzgeria furcatal.dumort 1995; 1996; 2001; 2002a

124 125 Metzgeria decipiens; 2002bMetzgeria decipiens; 2004; 2007; 2009; Metzgeria leptoneura Spruce 2004 Metzgeria lindbergii Schiffn 1988Metzgeria conjugata ssp japonica; 2002a; 2004; 2007; 2007; 2008; 2009; Metzgeria temperata Kuwah. 2001; 2002; 2002a; 2002b; 2004; 2004; 2007; 2009; Lunulariaceae Lunularia cruciatal.dumort. 1996; 2001; 2004; 2007; Wiesnerellaceae Dumortiera hirsutasw.nees 1995; 1996; 2001; 2003; 2004; 2005; 2007; 2009; 2009; Dumortiera hirsutasw.nees ssptatunoi Horik 2009 Wiesnerella denudatamitt.steph 1995; 1996; 2001; 2004; Conocephalaceae Conocephalum conicuml.dumort 1988; 1995; 1996; 2001; 2002; 2002b; 2003; 2004; ; 2007; 2008; 2009; 2009; 2009; 2010; Conocephalum japonicumthunb.grolle 1988; 1995; 1996; 2001; 2003; Tsukuba Botanical Garden Higuchi2004; 2004; 2005; 2007; 2007; 2008; 2009; Aytoniaceae Plagiochasma japonicumsteph.c. Massal Mannia fragransbalbisfrye et L. Clark

125 126 Reboulia hemisphaerical.raddi ssporientalis R.M. Schust 2001; 2004; 2005; 2007; Marchantiaceae Marchantia emarginata Reinw.Blume et Nees ssp tosanasteph.bischl 1996; 2004 Marchantia paleacea BertolsspdipteraNees et Mont.S. Hatt 1988; 2001; Marchantia polymorpha L 1996; 2001; 2004; 2005; 2007; 2009; Riccia glauca Lvarciliaris Warnst Riccia huebeneriana Lindenb 1995; 2001; 2004; 2007; Riccia lamellosa Raddi Riccia miyakeana Schiffn 2004 Riccia nigrella DC 2007; Riccia pubescens S. Hatt 2007; Ricciocarpos natansl.corda 2001; 2004; 2005; 2007; Joso-shi Kokai RiverHiguchi2009; Monosoleniaceae Monosolenium tenerum Griff Tsukuba Botanical GardenInoue1988; Tsukuba Botanical GardenHiguchi2000 Ricciaceae Riccia cavernosa Hoffm 2006 Riccia fluitans L 2001; 2003; 2005; 2007; Riccia glauca L 1995; 2001; 2004; 2007; Anthocerotopsida Notothyladaceae Notothylas orbicularisschwein.sull 2004; 2007; Anthocerotaceae Anthoceros punctatus L a2005; 2009; Anthoceros subtilis Steph 2004 Megaceros flagellarismitt.steph 2009 Phaeoceros carolinianusmichx.prosk 2003; 2007

126 127. Ahlburg, H Eine Reise nach dem Tsukuba von dem 1878 in Tokio verstorbenen Professor Ahlburg. Gartenflora, : Amakawa, T. and S. Hattori A revision of the Japanese species of Scapaniaceae. J. Hattori Bot. Lab., : Ando, H The Hypnum species restricted to Japan and adjacent areas3. J. Sci. Hiroshima Univ., ser. b, div. 2, : Ando, H A revision of the East-Asian species of Homomallium, a genus of Musci I. Hikobia, 1-2: Deguchi, H A revision of the genera Grimmia, Schistidium and CoscinodonMusciof Japan. J. Sci. Hiroshima Univ., ser. b, div. 2, 2: Dixon, H. N Contributions to japanese Bryology. Part 1. Brachyteciaceae. Rev. Bryol. Lich., : Dixon, H. N Mateiae ad Bryophytas Nipponicae. I. Bot. Mag. Tokyo, : Dixon, H. N Some new japanese mosses. Rev. Bryol. Lich., : pp , pp., , 2002., : Riccia cavernosa Hoffm.., : , : , : Higuchi, M A reappraisal of Pylaisia nanahypnaceae, Bryopsida. Bryobrothera, : Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXI, nos National Science Museum, Tokyo. Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXII, nos National Science Museum, Tokyo. Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXIII, nos National Science Museum, Tokyo. Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXV, nos National Museum of Nature and Science, Tokyo. Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXVI, nos National Museum of Nature and Science, Tokyo. Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXVII, nos National Museum of Nature and Science, Tokyo. Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXVIII, nos National Museum of Nature and Science, Tokyo. Higuchi, M Bryophyta Selecta Exsiccata. Fasc. XXX, nos National Museum of Nature and Science, Tokyo , : Inoue, H The family Plagiochilaceae of Japan and Formosa. II. J. Hattori Bot. Lab., : pp ,. Inoue, H Bryophyta Selecta Exsiccata. Fasc. XIX, nos National Science Museum, Tokyo. Iwatsuki, Z A revision of the Japanese species of the genus Ulota. J. Hattori Bot. Lab., : Iwatsuki, Z A revision of Plagiothecium and its related genera from Japan and her adjacent areas, I. J. Hattori Bot. Lab., : Iwatsuki, Z Re-examination of Myurium and its related genera from Japan and its adjacent areas. J. Hattori Bot. Lab., : pp.,. Iwatsuki, Z New Catalog of the Mosses of Japan. 182 pp., Hattori Botanical Laboratory, Nichinan. Iwatsuki, Z. and T. Suzuki A taxonomic revision of the Japanese species of Fissidens Musci. J. Hattori Bot. Lab., : , 2007., : Kanda, H A revision of the family Amblystegiaceae of Japan I. J. Sci. Hiroshima Univ., Ser. B, Div. 2, : Kanda, H A revision of the family Amblystegiaceae of Japan II. J. Sci. Hiroshima Univ., Ser. B, Div. 2, : ,.. : pp ,

127 128., : Koponen, T A report on RhizomniumMniaceaein Japan. J. Hattori Bot. Lab., : Magombo, Z. L. K Taxonomic revision of the moss family Diphysciaceae. J. Hattori Bot. Lab., : , : 348. Matsui, T. and Z. Iwatsuki A taxonomic revision of the family DitrichaceaeMusciof Japan, Korea and Taiwan. J. Hattori Bot. Lab., : pp , : , : , 349 pp., , 451 pp., , 490 pp., , 462 pp., pp , pp , , : Nishimura, N A revision of the genus Ctenidium Musci. J. Hattori Bot. Lab., : ,,., : ,,., : Noguchi, A Musci japonici. IV. The genus Ptychomitrium. J. Hattori Bot. Lab., : Noguchi, A Musci japonici. VII. The genus Macromitrium. J. Hattori Bot. Lab., : Noguchi, A Musci japonici. IX. The Leskeaceae. J. Hattori Bot. Lab., : Noguchi, A Musci japonici. X. The genus Racomitrium. J. Hattori Bot. Lab., : Noguchi, A A taxonomic revision of the family Meteoriaceae of Asia. J. Hattori Bot. Lab., : Ochi, H A revision of the Bryaceae in Japan and the adjacent regions. 124 pp. Biological Institute, Faculty of Liberal Arts, Tottori University, Tottori. Okamura, S Contributiones novae ad floram bryophyton japonicam. Pars Prima. J. Coll. Sci. Imp.Univ. Tokyo, : Osada, T Japanese Polytrichaceae. I. Introduction and the genus Pogonatum. J. Hattori Bot. Lab., : Osada, T Japanese Polytrichaceae. II. The genera Polytrichum, Oligotrichum, Bartramiopsis and Atrichum and phytogeography. J. Hattori Bot. Lab., : Paris, G Musci japonica. Bull. Herb. Boiss. sér.2, : Reimers, H. and K. Sakurai Beitraege zur Moosflora Japans. I. Bot. Jahrb., : Saito, K A monograph of Japanese PottiaceaeMusci. J. Hattori Bot. Lab., : , pp , ,5: ,6: , pp , , pp , ,12: 51-66, pls , a.., : b.,., : ,,., : a pp. 13, b pp. 27, , :

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129 Bull. Ibaraki Nat. Mus.,14: A List of Macromycetes on the Grounds of Ibaraki Nature Museum Hiroyoshi KITAZAWA, Kei IMAMURA, Norimasa SHINDO and Mihoko UZAWA Accepted September 29, 2011 Abstract Macromycete flora was investigated mainly in coppices on the grounds of Ibaraki Nature Museum located in southwest Ibaraki Prefecture. Surveys were carried out continuously from June 1999 to February 2011, and 95 species were newly added to a preexisting list. As a result, the number of macromycete species recorded on the grounds of this museum has reached 292. We also mention the seasonal appearance of main species observed on the grounds of Ibaraki Nature Museum. Key words: Ibaraki Nature Museum, macromycetes, coppice forest ha * Ibaraki Nature Museum Volunteer, 700 Osaki, Bando, Ibaraki , Japan ** Ibaraki Nature Museum, 700 Osaki, Bando, Ibaraki , Japan

130 INM I II a Fig. 1. The grounds of Ibaraki Nature Museum.

131 Table 1. Monthly observations on main species in the period from February, 1995 to February, 2011.

132 Kirk et al a1982b1983a1983b a1988b a I 4: b1 4: a1 15: b2 5: : : : a3 10: b4 211: pp.

133 pp pp pp pp pp pp : pp. 2004,7: pp I 328 pp II 315 pp : : 323 Kirk, P. M., P. F. Cannon, D. W. Minter and J. A. Stalpers Dictionary of the fungi. 10 th ed. 771 pp., CABI. 1999,2: pp. 1998,1: : Geastrum kotlabae1: : pp. a pp II pp

134 136 Appendix. A list of macromycetes collected on the grounds of Ibaraki Nature Museum. BASIDIOMYCOTA AGARICOMYCOTINA AGARICOMYCETES AGARICALES Agaricaceae 1. Agaricus abruptibulbus Peck Ts INM Agaricus arvensis Schaeff. : Fr. Tk INM Agaricus campestris L. : Fr. Kn INM Agaricus praeclaresquamosus Freeman Ts INM ; Dn INM ; Ya INM ; Ya INM ; Ya INM Agaricus subrutilescenskauffm.hotson and Stuntz Dn Bovista dermoxanthavittad.de Toni Mf Kasuya Calvatia craniiformisschw.fr. Dn INM ; Ya INM ; Ya INM ; Ha INM ; Kn INM Coprinus comatusmüller: Fr.Pers. Mf INM Cystoderma amianthinumscop. : Fr.Fayod Dn INM Lanopila nipponicakawam.y. Kobayasi Ht INM ; Ht INM Lepiota acutesquamosaweinm. : Fr.Gill. s. lat. Kn INM ; Kn INM ; Mf INM Lepiota clypeolariabull. : Fr.Kummer Dn M Leucoagaricus rubrotinctuspecksing. Km M-566; Ts M-371; Sb INM ; Ya INM Leucocoprinus brebissoniigod.locq. Km M Leucocoprinus bresadolaeschulz.s. Wasser Km M Lycoperdon perlatum Pers. Bt INM ; Dn INM Lycoperdon pusillum Batsch: Pers. Mf INM ; Ha INM Macrolepiota sp. Ya Phaeolepiota aureamatt. : Fr.Maire Bt INM ; Sg INM ; Km INM Amanitaceae 20. Amanita ceciliaeberk. and Br.Bas Ya INM Amanita citrinaschaeff.pers. var. citrina Ts INM ; Ts INM Amanita esculenta Hongo and Matsuda Dn INM ; Dn INM ; Dn INM ; Kn INM ; Kn INM ; Ya INM Amanita fuliginea Hongo Ts INM

135 Amanita gemmatafr.bertillon Dn INM Amanita hemibaphaberk. and Br.Sacc. subsp. javanica Corner and Bas Dn INM ; Dn INM ; Dn INM Amanita hongoi Bas Dn INM Amanita longistriata Imai Kn INM ; Sb INM ; Ts INM ; Ts INM ; Ts INM Amanita melleiceps Hongo Mf M Amanita neo ovoidea Hongo Mf Amanita pantherinadc. : Fr.Krombh. Mf Amanita pseudoporphyria Hongo Mf Amanita rubescens Pers. : Fr. Dn INM ; Dn INM Amanita sp. Mf INM Amanita spissacea Imai Dn INM ; Dn INM Amanita spretapecksacc. Kn Amanita sychnopyramis Corner and Bas f. subannulata Mf Amanita vaginatabull. : Fr.Vitt. var. punctata Cleland et CheelGilb. Dn ; Dn Amanita vaginatabull. : Fr.Vitt. var. vaginata Dn INM ; Dn INM ; Dn INM Amanita virgineoides Bas Dn M-568; Dn INM Amanita virosafr.bertillon Dn M-342; Dn INM ; Mf INM Amanita volvatapeckmartin Dn INM ; Dn INM ; Dn INM ; Dn INM Bolbitiaceae 42. Bolbitius reticulatuspers. : Fr.Ricken Ya INM Cortinariaceae 43. Cortinarius purpurascensfr.fr. Mf Cortinarius subarmillatus Hongo Km INM Entolomataceae 45. Entoloma ater Hongo Ts INM Entoloma kujuensehongohongo Km INM Entoloma murraiiberk. and Curt.Sacc. Dn INM ; Dn INM Entoloma murraiiberk. and Curt.Sing. f. albus HiroeHongo To INM Entoloma nidorosumfr.quél. Ya INM ; Dn INM Entoloma quadratumberk. and Curt.Horak Dn INM Entoloma rhodopoliumfr.kummer Dn INM ; Dn INM ; Dn INM ; Dn INM

136 Entoloma saepiumnoulet and Dass.Richon and Roze Mf INM ; Ba INM ; Ba INM ; Ba INM ; Ba INM ; Ba INM ; Mf INM ; Ha INM ; Ha INM Fistulinaceae 53. Porodisculus pendulusschw.murrill Kn INM Hydnangiaceae 54. Laccaria amethysteabull.murr. Bt INM Laccaria laccatascop. : Fr.Berk. and Br. Ts INM ; Ha INM Laccaria laccatascop. : Fr.Berk. and Br. f. minuta Imai Mf M Laccaria murina S. Imai Mf INM Laccaria vinaceoavellanea Hongo Ts INM ; Ts INM Hygrophoraceae 59. Hygrocybe conicascop. : Fr.Kummer Dn INM Inocybaceae 60. Crepidotus badiofloccosus Imai Mf INM ; Mf INM ; Ts INM ; Ya M Crepidotus malachiusberk. and Curt.Sacc. Dn INM Crepidotus mollisschaeff. : Fr.Kummer Dn INM Inocybe asterospora Quel. Me M Inocybe fastigiataschaeff.quél. Ts INM ; Ts INM Inocybe lacerafr.p. Kumm. Mf INM On the ground under Quercus spp. 66. Inocybe quercina Hongo Dn M Inocybe umbratica Quél. Km INM ; Ts INM ; Ts INM Phaeomarasmius erinaceellapecksing. Dn INM Lyophyllaceae 69. Asterophora lycoperdoidesbull.ditm. : Fr. Kn INM ; Kn INM ; Ts INM ; Ts INM ; Ts INM Calocybe ionidesbull. : Fr.Donk Kn INM Lyophyllum decastesfr. : Fr.Sing. Mf INM ; Bt INM ; Bt INM ; Me INM ; Mf INM ; Me INM Marasmiaceae 72. Gerronema fibulabull. : Fr.Sing.

137 139 Ya INM Gerronema nemorale epichysium Har. Takahashi Ts INM ; Dn INM Gymnopus confluenspers.antonin, Halling and Noordel. Mf INM Marasmiellus chamaecyparidishongohongo Ya INM Marasmius maximus Hongo Ts INM ; Ts INM ; Dn INM ; Ha INM ; Ya INM Marasmius pulcherripes Peck Dn INM ; Mf INM ; Bt INM ; Ts INM Marasmius scorodoniusfr.fr. Ya M Marasmius siccusschw.fr. Kn INM Mycenaceae 80. Mycena galericulatascop.gray Mf Mycena haematopodapers. : Fr.Kummer Km INM ; Ts INM Mycena purapers. : Fr.Kummer Ts INM ; Dn INM Mycena roridascop. : Fr.Quél. Dn INM Panellus stipticusbull.p. Karst. Mf Xeromphalina campanellabatsch: Fr.Maire Dn INM ; Ts INM ; Mf INM Physalacriaceae 86. Armillaria melleavahlp. Kumm. Kn INM ; Kn INM Mf INM ; Mf INM Armillaria nabsnona Volk and Burdsall Dn M Armillaria tabescensscop.emel Ts INM ; Mf INM Cyptotrama asprataberk.redhead and Ginns Sg INM ; Dn INM ; Dn INM Flammulina velutipescurt. : Fr.Sing. Dn INM ; Ya INM ; Ya INM ; Kn INM ; Km INM Megacollybia platyphyllapers. : Fr. Ts INM Xerula radicatarelhan: Fr.Sing. Kn INM ; Dn INM ; Ya INM ; Dn INM ; Dn INM ; Dn INM Pleurotaceae 93. Hohenbuehelia reniformisg. Mey.Singer Mf INM ; Kn INM Pleurotus djamorrumph. ex Fr.Boedijn Dn INM Pleurotus ostreatusjacq.p. Kumm.

138 140 Sg INM ; Kn INM ; Ts INM ; Ts INM ; Mf INM Pleurotus pulmonariusfr.quél. Dn INM Pluteaceae 97. Pluteus aurantiorugosustrogsacc. Sg INM ; Dn INM ; Mf INM On decayed wood of Quercus sp.; Ts INM Pluteus cervinusschaeff.p. Kumm. Dn INM ; Ts INM ; Ts INM ; Mf INM ; Ts INM Pluteus leoninusschaeff. : Fr.Kummer Tk INM Pluteus nanuspers. : Fr.Kummer f. nanus Mf INM Pluteus thomsoniiberk. and Br.Dennis Km M Volvariella speciosafr. : Fr.Sing. var. gloiocephala DC. : Fr.Sing. Ts INM Psathyrellaceae 103. Coprinellus disseminatuspers.j. E. Lange. Dn INM Coprinellus micaceusbull.vilgalys, Hopple and Jacq. Johnson Ts INM ; Ts INM ; Ts INM ; Dn INM ; Dn INM Coprinellus radiansdesm.vilgalys, Hopple and Jacq. Johnson Dn INM ; Dn INM ; Mf INM Coprinopsis atramentariabull.redhead, Vilgalys and Moncalvo Ka INM ; Mf INM Coprinus patouillardi Quél. Ya INM ; Dn INM ; Mf INM Coprinus rhizophorus Kawamura ex Hongo and K. Yokoyama Kn INM Psathyrella candolleanafr.maire Ts INM ; Dn INM ; Ts INM ; Ts INM ; Ts INM ; Mf INM On decayed wood of Quercus sp Psathyrella gracillisfr.quél. Km INM ; Dn INM Psathyrella obtusatapers.a. H. Sm. Ts INM Psathyrella piluliformisbull. : Fr.P. D. Orton Ts INM Psathyrella spadiceogriseaschaeff.maire Dn INM Psathyrella velutinapers.sing. Sg INM ; Sg INM ; Ts INM ; Sg INM ; Bt INM ; Mf M-186; Ha INM ; Ts INM Pterulaceae 115. Deflexula fascicularisberk. and Pat.Corner

139 141 Ts INM Pterula multifidachev.fr. Ts M-597 Schizophyllaceae 117. Schizophyllum commune Fr. Mf INM ; Mf INM ; Mf INM Strophariaceae 118. Agrocybe erebiafr.kühner ex Singer Dn INM Agrocybe praecoxpers. : Fr.Fayod Dn INM ; Dn INM ; Dn INM ; Dn INM ; Kn INM Gymnopilus aeruginosuspecksing. Sg INM ; Ts INM Gymnopilus liquiritiaepers.p. Karst. Mf Hypholoma fascicularehudson: Fr.Karst. Ts INM ; Dn INM ; Dn INM ; Dn INM ; Kn INM Hypholoma sublateritiumfr.karst. Dn INM Kuehneromyces mutabilisschaeff. : Fr.Sing. and A. H. Smith Ya INM Pholiota aurivellabatsch: Fr.Kummer Ya INM ; Ya M Pholiota highlandensispecka. H. Smith and Hesler Mf INM ; Dn INM Pholiota squarrosabatschp. Kumm. Mf INM On decayed wood of Quercus sp.; Bt INM On decayed wood of Quercus sp Pholiota terrestris Overh. Mf INM ; Km INM Stropharia rugosoannulata Farl. ex Murrill Sg INM ; Mf INM ; Bt INM Tricholomataceae 130. Clitocybe fragranswith. : Fr.Kummer Mf INM ; Km INM ; Mf INM Clitocybe nebularisbatsch: Fr.Kummer Dn INM ; Dn INM ; Dn INM ; Dn INM Clitocybe sp. Ya INM ; Dn INM ; Dn INM ; Ya INM Clitocybe trogioides Corner var. odorifera Har. Takahashi Ya M Lepista nudabull. : Fr.Cooke. Ha INM ; Kn INM ; Dn INM ; Dn INM ; Dn INM ; Mf INM Lepista sordidaschum. : Fr.Sing. Km INM

140 Leucopaxillus giganteussow. : Fr.Sing. Ya INM Melanoleuca melaleucapers. : Fr.Murr. Ts INM ; Km INM ; Mf INM On the ground 138. Phyllotopsis nidulanspers. : Fr.Sing. Ya INM ; Km M Resupinatus trichotispers.sing. To INM Tricholomopsis rutilansschaeff. : Fr.Sing. Ts INM Family incertae sedis 141. Plicatura crispafr.reid Mf AURICULARIALES Auriculariaceae 142. Auricularia auriculahook.underw. Tk INM ; Ya INM ; Mf INM On the ground under Quercus spp.; Mf INM Auricularia polytrichamont.sacc. Tk INM ; Dn INM ; Sg INM ; Kn INM ; To INM Exidia glandulosa Fr. Ts M Exidia uvapassa Lloyd Dn INM BOLETALES Boletaceae 146. Boletus aokii Hongo Dn Boletus fraternus Peck Kn INM Boletus pulverulentus Opat. Ts INM ; Ts INM ; Kn INM ; Ka INM Leccinum extremiorientalel. Vass.Sing. To INM Leccinum griseumquél.sing. Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Ts M Phylloporus bellusmass.corner Mf INM Strobilomyces confusus Sing. Mf INM Tylopilus fumosipespecka. H. Smith and Thiers Dn INM ; Dn INM Tylopilus neofelleus Hongo Mf Tylopilus virenschiuhongo To INM ; Dn INM ; Dn INM Xerocomus chrysenteronbull.quél. Dn INM ; Ts INM ; Ts INM ; Ts INM ; Kn INM Xerocomus nigromaculatus Hongo Ts INM

141 Xerocomus subtomentosusl. : Fr.Quél. Ts INM ; Ts INM ; Ha INM ; Kn INM Coniophoraceae 159. Gyrodontium versicolorberk. and Br.Mass G. Km INM Diplocystidiaceae 160. Astraeus hygrometricuspers.morgan Mf Gyroporaceae 161. Gyroporus castaneusbull. : Fr.Quél. Ya M-604 Paxillaceae 162. Paxillus curtisii Berk. in Berk. and Curt. Mf INM ; Ts INM Sclerodermataceae 163. Scleroderma reae Guzman Ts INM ; Mf INM Scleroderma verrucosumbull.pers. Ya INM CANTHARELLALES Cantharellaceae 165. Cantharellus cibarius Fr. Tk INM ; Dn INM ; Tk INM ; Kn INM Cantharellus minor Peck Km INM Craterellus cornucopioidesl. : Fr.Pers. Kn INM Clavulinaceae 168. Clavulina cristataholmsk. : Fr.Schroet. Ts INM GEASTRALES Geastraceae 169. Geastrum mirabilemont.fisch. Ya INM Sphaerobolus stellatus Tode: Pers. Km INM GLOEOPHYLLALES Gloeophyllaceae 171. Gloeophyllum abietinumfr.karst. Mf INM HYMENOCHAETALES Hymenochaetaceae 172. Cryptoderma substygiumberk. and Br.Imazeki Ts M Hymenochaete mougeotiifr.cooke Ts M Inonotus mikadoilloydimazeki Dn M Onnia scauralloyd Imazeki Dn INM Phellinus gilvusschw. : Fr.Pat. Ts INM ; Kn INM ; Mf M-609 POLYPORALES Fomitopsidaceae 177. Antrodia albidafr.donk To INM Antrodia sinuosafr.p. Karst. Mf M-610

142 144 Ganodermataceae 179. Ganoderma applanatumpers.pat. Dn M Ganoderma lucidumleyss. : Fr.Karst. Sg INM Meripilaceae 181. Grifola frondosadicks. : Fr.S. F. Gray Dn INM ; Dn INM Rigidoporus zonalisberk.imazeki Tk INM ; Tk INM ; Mf INM Meruliaceae 183. Bjerkandera adustawilld. : Fr.Karst. Ts M Bjerkandera fumosapers. : Fr.Karst Kn INM Gloeoporus dichrousfr.bresadola. Ya M Hydnophlebia chrysorhizatorr.parmasto Dn INM ; Dn M Junghuhnia nitidafr.ryv. Ya M Mycoacia copelandiipat.aoshima and Furukawa Dn INM ; Ya M-314 Phanerochaetaceae 189. Antrodiella fragransa. David and TorticA.David and Tortic 1 Mf M Antrodiella zonataberk.ryvarden Ts INM ; Dn INM ; Kn INM ; Mf INM Phanerochaete sordidakarst.j. Erikss. and Ryv. Ts Polyporaceae 192. Abundisporus pubertatislloydryvarden Sg INM Cerrena unicolorfr.murr. Dn INM ; Ts INM ; Kn M Coriolopsis strumosafr.ryv. Dn INM ; Dn INM ; Dn INM Cryptoporus volvatuspeckshear Ts INM Daedaleopsis styracinap. Henn. et ShiraiImazeki Dn INM ; Ya INM ; Mf INM Daedaleopsis tricolorbull. : Fr.Bond. and Sing. Ts INM Lentinus suavissimus Fr. Mf Lenzites betulinal. : Fr.Fr. Dn M Microporus subaffinislloyd.imaz. Ts M Microporus vernicipesberk.o. Kuntze Dn INM ; Dn INM ; Mf INM Neolentinus lepideusfr.redhead and Ginns Dn INM ; Dn INM-

143 ; Ts INM ; Mf INM ; Mf INM On decayed wood of Pinus densiflora 203. Oligoporus caesiusschrad. : Fr.Gilbn. et Ryv. Ts INM ; Ts INM ; Ya M Panus rudis Fr. Bt INM Panus tigrinusbull. : Fr.Sing. Sg INM Perenniporia fraxineabull.ryvarden Mf INM Perenniporia ochroleucaberk.ryvarden To INM Polyporus alveolariusdc. ex Fr.Boud. et Sing. Sg INM ; Km INM ; Mf INM Polyporus arcularius Batsch. : Fr. Ts INM ; Mf INM ; Ds INM ; Mf INM ; Ts INM Polyporus emerici Cooke Kn M Polyporus squamosus Fr. Km INM ; Kn INM ; Ts INM Polyporus tuberasterjacq.fr. Mf INM ; Ts INM Polyporus varius Pers. : Fr. Ts INM ; Ts INM ; Bt M Postia tephroleucafr.jülich Mf Pycnoporus coccineusfr.bond. et Sing. Tk INM ; Sg INM ; Kn INM ; Mf M-572 ; Ts INM Trametes hirsutawulfenlloyd Dn INM ; Mf M-612; Mf INM Trametes orientalisyasudaimazeki Ts INM ; Kn INM ; Ts M-285 ; Ya M Trametes pubescensschumach.pliȧt Mf Trametes versicolorl.lloyd Sg INM ; Ts INM ; Ts INM ; Dn INM ; Ts INM ; Kn INM ; Ts INM ; Mf INM Trichaptum abietinumdicks. : Fr.Ryvarden Ts M Trichaptum biformefr.ryvarden Mf Steccherinaceae 222. Steccherinum ochraceumpers.s.f.gray Ya M Steccherinum rhoisschw.banker Ts INM RUSSULALES Auriscalpiaceae 224. Lentinellus ursinusfr.kühner

144 146 Dn M-586 ; Ts M-587 Russulaceae 225. Lactarius chrysorrheus Fr. Dn INM Lactarius circellatus Fr. f. distantifolius Hongo Dn INM ; Dn INM ; Ts INM ; Ts INM ; Ya INM ; Ya M Lactarius subplinthogalus Coker Mf Lactarius subzonarius Hongo Kn INM ; Dn INM Lactarius volemusfr.fr. Dn INM Russula aeruginea Lindbl. apud Fr. Mf M Russula alboareolata Hongo Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM Russula castanopsidis Hongo Mf Russula cyanoxanthaschaeff.fr. Ts INM ; Dn INM ; Ts INM ; Ts INM ; Dn INM Russula delica Fr. Ts INM Russula emeticaschaeff.pers. Mf Russula foetens Pers. : Fr. Bt INM Russula gracillima J. Schaff. Ts INM Russula mariae Peck Dn INM ; Sg INM ; Ts INM ; Ts INM ; Ts INM Russula nigricansbull.fr. Dn INM ; Dn INM ; Dn INM ; Mf INM ; Dn INM ; Dn INM ; Dn INM ; Ts INM Russula ochroleucapers.fr. Ts INM Russula olivaceaschaeff.fr. Dn INM Russula pseudointegra Arnould and Goris Mf Russula sanguineabull.fr. Dn INM Russula senecis Imai Ts INM ; Ts INM Russula sororiafr.romell Ya M-588; Ya M Russula violeipes Quél. Ts INM Russula virescensschaeff.fr. Dn INM ; Ya INM ; Kn INM Stereaceae 248. Stereum gausapatum Fr. : Fr. Ya INM Stereum hirsutumwilld. : Fr.S. F. Gray Ya M-591

145 Stereum ostreabl. and NeesFr. Ya M Xylobolus spectabilisklotz.boidin Ts INM PHALLALES Phallaceae 252. Linderia bicolumnatalloydcunn. Ts INM Lysurus mokusinl. : Pers.Fr. Mf INM ; Mf INM ; Mf INM ; Dn INM ; Dn INM ; Dn INM ; Dn INM ; Mf INM Lysurus mokusinl. : Pers.Fr. f. sinensislloyd Kobayasi Dn INM Mutinus caninuspers.fr. Mf INM Phallus impudicus Pers. Ts INM ; Ts INM Phallus rugulosusfisch.o. Kuntze Mf NM ; Mf NM ; Ha INM ; Dn INM ; Ha NM Kobayasia nipponicay. KobayasiImai and Kawamura Dn INM THELEPHORALES Thelephoraceae 259. Thelephora aurantiotincta Corner Dn M-337; Mf INM On the ground under Quercus spp. Incertae sedis 260. Cotylidia diaphanaschw.lentz Ts INM DACRYMYCETES DACRYMYCETALES Dacrymycetaceae 261. Calocera comeeabatsch: Fr.Fr. Ya M Dacryopanax spathulariaschw.fr. Mf INM TREMELLOMYCETES TREMELLALES Tremellaceae 263. Tremella fimbriata Pers. : Fr. Kn INM ; Ts INM Tremella foliacea Pers. : Fr. Mf INM Tremella fuciformis Berk. Mf INM On decayed wood of Quercus sp. ; Sg INM ; Dn INM ; Dn INM ; Kn INM ASCOMYCOTA PEZIZOMYCOTINA LEOTIOMYCETES HELOTIALES Rutstroemiaceae 266. Dicephalospora rufocorneaberk. and Broome Spooner Ya M-606 Sclerotiniaceae 267. Ciborinia camelliae Kohn Bt INM ; Ya INM

146 148 Family incertae sedis 268. Cordierites frondosakobayasikorf Kn M Mollisia cinereabatsch: Fr.Karsten Ts M Pyrenopeziza sp. Mf INM ORBILIOMYCETES ORBILIALES Orbiliaceae 271. Orbilia sp. 1 Mf INM PEZIZOMYCETES PEZIZALES Caloscyphaceae 272. Caloscypha fulgenspers.boud. Ya INM Discinaceae 273. Discina parma J. Breitenb. et Maas Geest. Ya INM ; Ya INM ; Ya INM Helvellaceae 274. Helvella atra J. König Ya M Helvella crispascop.fr. Kn M-562 Morchellaceae 276. Morchella esculental.pers. var. esculenta Tk INM ; Mf INM ; Tk INM ; Sg INM ; Tk INM ; Tk INM ; Ya INM ; Ts INM ; Ya INM ; Ha INM ; Tk INM ; Bt INM ; Bt INM Pezizaceae 277. Peziza spp. 1 Ts INM ; Ya INM ; Ya M Peziza badia Pers. Mf INM Pyronemataceae 279. Aleuria aurantiafr.fuckel Km INM ; Mf INM Scutellinia erinaceusschwein.kuntze 1 Ts M Scutellinia scutellatal.lambotte Km INM Scutellinia sp. 1 Mf INM Sarcosomataceae 283. Galiella celebicap. Henn.Nannf. Ts INM ; Ts INM ; Ts INM ; Mf INM On decayed wood of Quercus sp. SORDARIOMYCETES HYPOCREALES Cordycipitaceae 284. Cordyceps tuberculatalebertmaire Mf INM Ophiocordyceps formicarumkobayasig. H. Sung, J. M. Sung, Hywel-Jones and Spatafora in Sung, Hywel-Jones, Sung, Luangsa-ard, Shrestha and Spatafora Mf INM Isaria japonica Yasuda

147 149 Km INM Hypocreaceae 287. Hypocrea sp. 1 Mf INM ; Mf INM Nectriaceae 288. Nectria sp. 1 Mf INM XYLARIALES Xylariaceae 289. Daldinia concentricaboltonces and de Not. Dn Hypoxylon spp. 1 Mf INM Hypoxylon truncatumschw.j. H. Miller Ts M Xylaria polymorphapers.grev. Mf INM Ts: Km: Dn: Kn: Ya: Ba: Me: Tk: Bt: To: Ka: Sb: Ht: Ha: Mf: Sg:

148 Bull. Ibaraki Nat. Mus.,14: Invited German Professor Hermann Ahlburg, the First Reporter on Plants from Mt. Tsukuba Minoru YAMAKAWA, Mihoko UZAWA and Kazuo OBATA Accepted April 26, 2011 Abstract The German scholar Hermann Ahlburg was invited by the Japanese Government to serve as a professor at the University of Tokyo Medical School in The next year, he traveled to Mt. Tsukuba via Sanuma Marsh in Ibaraki Prefecture. In 1878, he died of an infectious disease in Japan. An original paper on his trip to Sanuma March and Mt. Tsukuba, the first report on the plants of these areas, was published in 1879 and an article about his death appeared in the German journal Gartenflora. We translated these two articles from German into Japanese, traced his activity in Japan and clarified dates of his trip to Mt. Tsukuba. We also compared some plants described by Ahlburg with the present flora of these areas. Key words: Hermann Ahlburg, Meiji era, Mt. Tsukuba, plants, Sanuma Marsh. 877 m Gartenflora 1998 Eine Reise nach dem Tsukuba von dem in 1878 in Tokio verstorbenen Professor * National Institute of Agrobiological Sciences, 1-2 Owashi, Tsukuba, Ibaraki , Japan. ** Graduate School of Life and Environmental Sciences, University of Tsukuba, Tennodai, Tsukuba, Ibaraki , Japan. *** Ibaraki Nature Museum, 700 Osaki, Bando, Ibaraki , Japan.

149 152 Gartenflora * 50. Fig. 1. Scene of the exhibit on Professor Ahlburg s travels at the 50th special exhibition Vibrant Mt.Tsukuba! Discover beeches, toads, and boulders!. Ahlburg: 1878 Professor Ahlburg Professor Ahlburg Sanuma 1877 * *

150 153 Fukugawa* Raphanus sativus* Raphanus sativus L. Fukugawa Diospyrus Kaki* Diospyros kaki Thunb. Citrus* Citrus unshiuswingles. Marcov. 5 Fuhugawa* Fukugawa Hoschiba* 10 * 1 Shingawa* Rüenarten*

151 Quercus * Cryptomeria japonica* Castanea vesca* Tilia* Zelkowa kiaki* Zelkova serrata Aesculus turbinata* Rhus* Evonymus* EuonymusEuscaphys* Euscaphis 12Gentianeen* Cariceen* Oxalideen* Geraniaceen* Caryophyllaceen* 3 5 Sanuma* Nelumbo nucifera* Potamogeton natans* P. oxyphyllus * Sagittaria* Alocasia * Calocasia* Colocasia Leucocasia* Colocasia Conophyllus* Amorphophallus * 9 10 Meer* Rotunde* Yatsuashic8 Einlasskarten*

152 Rotunde* Ringer* Hejiga ike* Rokusho* Meer* Otayama* Tsukuba mikaye* Hirusawayama* 10 Arugawamura* 11 * Izanamino mikoto* 4 16 Miyas* Otto Volger * Gartenflora

153 Volger 1,000 Gartenflora E. R.* E. R. GartenfloraEduard Regel Niedersachsen Braunschweig WürzburgJulius von Sachs Franz M. Hilgendorf , Gartenflora Ahlburg Table 1. Chronological table of Professor Hermann Ahlburg. BraunschweigWürzburgSachs Eurotium oryzae Ahlburg Gartenflora

154 Fig. 2. Map of the route Professor Ahlburg traveled from Tokyo to Mt. Tsukuba. The river map of the early Meiji period is according to Ohkuma , Gartenflora. Fig. 3. Pictures of Gartenflora and the first page of the article on Professor Ahlburg s travelstop, and a page where plants from Mt. Tsukuba and Sanuma are describedbottom