9 F1 * * Breeding of new varieties and morphological characteristics of Rosaseae hybrids Mika TAKIYA* and Hirokazu KON* 9 1991 38 1965 1989 1995 1987 1978 1997 1997 1985 3 3 Hokkaido Forestry Research Institute, Bibai,Hokkaido 079-0198 39 14 3 Bulletin of the Hokkaido Forestry Research Institute, No.39 March, 2002 ( 12 )
9 F1 F1 1. Rosa rugosa THUNB. (R. r. f. alba WORE) (R. r. f. plena BYHOUWER) (R. r. f. alboplena REHD.) (R. acicularis LINDL.) (R. davurica PALLAS.) (R. multiflora THUNB.) (R. ruburifolia VILL.) Open 9 (-1) (forma) Cultivated variety 3 Open -1 (Rosa rugosa) (Rosa rugosa f. alba) ~8cm (Rosa rugosa f.alboplena) 9~10cm (Rosa rugosa f.alboplena) 7~8cm (Rosa acicularis) 4.5~5cm (Rosa davurica) 4.5~5cm (Rosa multifora) 2.5~3cm (Rosa rubriforia) 3~4cm Opne (Rosa rugosa f. alba Opne) 3.5~6.7cm 7.5~8.0cm 1) ~ 1)
2. 2 1991 9 52-3 5 15 54 116 3. 1991 38-4 1992 29 F1 535 20 1993 1994 29 F1 19 1995-3 9 (1991 ) % 15 0 0.0 13 9 69.2 10 7 70.0 10 8 80.0 10 8 80.0 10 4 40.0 16 13 81.3 15 8 53.3 5 4 80.0 Open 10 2 20.0 10 0 0.0 10 0 0.0 10 0 0.0 10 0 0.0 5 1 20.0 Open 10 4 40.0 12 11 91.7 10 0 0.0 5 0 0.0 10 6 80.0 Open 5 0 0.0 10 3 30.0 10 1 10.0 10 1 10.0 10 0 0.0 10 0 0.0 Open 10 6 60.0 2 0 0.0 10 8 80.0 10 10 100.0 10 0 0.0 10 9 90.0 10 8 80.0 5 4 80.0 Open 5 4 80.0 116 37 31.9 55 3 5.5 56 13 23.2 94 74 78.7 Open 54 2 3.7 5 4 80.0 5 5 100.0 5 3 60.0 5 4 80.0 Open 10 3 30.0 10 0 0.0 6 2 33.3 10 1 10.0 10 2 20.0 Open 5 0 0.0 Open 5 2 40.0 52 794 302
-2 22m 1997 17 148 1. 1991 1991 1992 () 1990 1991 (52 ) (38 ) 1992 1 1993 2 (29 ) ( ) 1994 3 1995 4 (19 165 ) 1996 5 1997 6 17 (19 148 ) 1998 7 1999 8 (8 8 ) 19 165 F1-4 1 1999 148 2. 19951999 F1 1 19971999 8 1997 1999 1 38 0 % 785 un1) un 482 127 26.3 704 312 44.3 582 200 34.4 279 21 7.5 1298 360 27.7 510 170 33.3 227 34 15.0 Open 133 22 16.5 88 40 45.5 Open 158 35 22.2 890 321 36.1 432 120 27.8 391 26 6.6 114 23 20.2 27 7 25.9 41 16 39.0 Open 294 54 18.4 423 75 17.7 388 53 13.7 225 0 0.0 366 21 5.7 100 8 8.0 Open 147 17 11.6 108 3 2.8 5 0 0.0 102 0 0.0 653 0 0.0 Open 3 0 0.0 35 3 8.6 35 0 0.0 51 0 0.0 127 0 0.0 Open 291 80 27.5 161 40 24.8 50 32 64.0 96 27 28.1 Open 215 0 0.0 1)
19981999 F1 RFA RFA = Af/Ac Af Af = (Df/2) 2 Nf Df Nf Ac Ac = Dc Lc, Dc Lc 1. (1) 52-3 9 23.2 60.0 3 90 4080 9 80 Open 20 0 40.081.3 6091.7 Open 60 040 (2) -4 6.664.0 5.717.7 0 8.6 2.8 0 2. (1) 8 F1-1a F1 F1 Open 235.7 213.0 94.6cm
1 2 3 4 5 6 7 8 9 Open (2) 8 F1-1b F1 F1 230.8 F1 158.4198.5 83.5 r=0.848, n=19, p<0.01 (3) 8 F1-1c F11.3 1.2 F1 0.70.9 0.91.1-1 -1 3. (1) -1 8 (1999 ) F1 4 8-2 F1 5
6 8 8 F1-3a F1 8 F1-3b (2) -2 4~8 (1995~1999 ) 8 F1-3c 7 F1 95.6mm F1 F1 F1 35.7mm F1 F1 F1 F1 F1 (3) RFA 8 F1 RFA -3d
-3 8 (1999 ) -1 RFA 0.665 0.0130.014 F1RFA 0.474 F1 F1 0.1040.2250.086 F1 0.0080.036 F1 RFA (4) RFA RFA -4 4 1 4 RFA -4a 4 1 RFA RFA F1-4b RFA -4a RFA -4a
-4 8 (1999 ) ) ) RFA F1 3 RFA F1 RFA F1-3 -1 (5) F1-5 28 F1 F1-1A,B F1-2A F1-3A,B F1-2A4A -3A,B -5A6A
-5 F1 1) n n code 6 6 6 9207 10 10 10 9207 2 2 2 9208 5 5 5 9208 19 19 19 8905 12 12 12 9205 10 10 10 9207 9 6 3 2 7 9213 3 3 3 9207 5 1 1 2 1 1 4 9204 9 9 9 9207 10 3 7 7 3 8906 8 8 8 9208 3 2 1 1 2 1 9508 3 3 3 9707 Open 20 16 4 1 19 9207 3 2 1 3 9207 8 1 7 8 4 9207 3 3 3 9206 19 148 32 24 69 22 1 3 24 7 114 5 F1-4B7B -1B5B6B 1-2B F1 F1
1. 9 Ueda et al. (1996) 104 3 3 R.iwara SIEB. 34cm ( 1989 1965 1995 ) 1994 1995 F1 F1 F1 1 F1 2. F1 (1) F1-3C
F1-5C6C F1 F1 8 3m -4C (2) F1 23 F1 F1 3 8 F1 F1-4C -4B7B F1-2A 3A4A -5A 6A F1-2A4A7A8A F1-6A -3A F1 (3) F1 2 F1 F1 2
F1 F1 F1 F1 9 38 8 1 3 2 1 1 1 2 1 2 1 1 1 8 8-6 -1-8 -6 (mm) n=10 () ()() ( ) ()() (cm) ~ ~ ~ 1) 1999 2)
F1 F1 F1 1996 RHS F1 F1 F1 1996 300159pp. 1995 43131-133 1994 Rosa rugosa Thunb. Ex Murray 62857-866 1997 250pp. 1965 1560pp. 1987 6717-22 1985 96387-388 1989 I321pp. 1995 303pp. Ueda, Y., Takeshita, D., and Ando, T. 1996 Pollination in Rosa rugosa Thunb. Ex Murray. Acta. Hort. 424:309-310 Summary Rosa rugosa is widely used as ornamental plant in Hokkaido. It is usually planted along seashore, because R. rugosa is the salt tolerant species. We have crossed nine species of Rosaseae each other to develop new cultivated varieties, mainly concerned with R. rugosa. R. rugosa, R. acicularis, R. davurica
and R. multiforia occur in Hokkaido. R. rubriforia is from Europe. R. rugosa f. alba, R. rugosa f. plena and R. rugosa f. alboplena are forma of R. rugosa. Hybrids from intraspecific breeding did not have notable characters (e.g. the colour of flowers, the tree form and number of flowers). However, on the other hand, those from interspecific breeding had worth characters as the ornamental plants. From the latter hybrids, we selected eight F1 individuals for the new cultivated varieties as follows. R. rugosa R. acicularis; that flowers had weakly waving and pinkish coloured petals. R. rugosa R. multifora; the flowers had waving petals. R. rugosa R. rubriforia; the shape of the flowers resembled to the flowers of R. rugoss, however, that size was smaller than that. R. rugosa f. plena R. multifora; the colour of the petals were pinkish red and the flowers were multi-centered. The shape of the tree was creeping. R. davurica R. multifora; this hybrid had large and semi-climbing tree form, and the rate of the total flower area in the crown surface area was high. R. davurica R. rubriforia; the colour of the flowers had vivid purple-red petals, and the shape of the tree was upright. R. multifora R. rugosa; the colour of the petals were pale pink, and the flowers number was high. R. rubriforia R. rugosa; the colour of the petals, the stems and the leaves were purple-red. The petals had eyed variegation on that center. Keywords: Rosa rugosa, Rosaseae, interspecific breeding, ornamental plants -1 8 (1992 ) RFA cm cm day days mm 6 85.8 113.3 1.3 13 73 49.4 0.008 10 140.6 159.3 1.1 74 126 63.8 0.036 2 130.0 155.0 1.2 27 70 95.6 0.023 5 160.8 146.5 0.9 84 19 72.0 0.042 19 173.9 179.2 1.0 168 27 55.6 0.041 12 139.7 128.8 0.9 458 27 38.1 0.107 10 150.4 108.8 0.7 216 27 52.1 0.095 9 119.2 126.7 1.1 16 82 61.9 0.009 3 120.0 132.5 1.1 37 103 66.9 0.024 5 94.6 83.5 0.9 163 23 44.3 0.104 9 172.9 190.8 1.1 116 24 55.8 0.031 10 196.2 198.5 1.0 2501 21 41.5 0.225 8 199.6 158.4 0.8 549 27 36.4 0.086 3 197.0 230.8 1.2 13730 26 35.7 0.474 3 213.0 192.5 0.9 341 29 40.9 0.049 Open 20 13.9 1124.6 1.1 35 83 64.9 0.028 3 125.3 127.5 1.0 43 93 65.7 0.029 8 110.4 123.1 1.1 37 101 62.9 0.026 3 53.7 2198.3 0.8 2139 29 36.7 0.209 19 148 5 118.0 106.0 0.9 16 57 74.9 0.019 2 103.5 93.8 0.9 20 103 70.8 0.033 5 207.6 189.0 1.1 58 85 98.9 0.018 2 104.0 57.5 0.5 18 131 70.8 0.046 3 87.7 65.8 0.9 5 14 46.3 0.013 2 243.5 168.8 0.7 160 27 43.8 0.029 2 175.5 150.0 0.9 4831 18 25.9 0.665 4 216.3 144.4 0.7 82 23 33.3 0.014