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Evo-devo eco-evo-devo Evo-devo and eco-evo-devo: Two extended syntheses in evolutionary biology Yoshinari YOSHIDA, Katsuya TAKAO, Hisashi NAKAO 1 30 80 e.g., Amundson 2005 90 evolutionary developmental biology evo-devo e.g., Hall 1992, [1998]2001 ecological developmental biology eco-devo Gilbert and Epel [2009]2012 Amundson 2005; Gilbert and Epel [2009]2012; 2009 Arthur 2011 Gilbert and Epel [2009]2012 yoshinariyoshida@gmail.com katsta07@ma.com hisashinakao@gmail.com

68 7 2013 2 Hall (1992) 20 Arthur 2011 Hall 1992 evo-devo 20 Arthur evo-devo evo-devo p. x Hall 1992 Arthur 2011 2.1 Arthur 2011 developmental repatterning p. 9 1 evo-devo 2 developmental repatterning 3 4 Arthur evo-devo evo-devo 1970 1980 p. 315 Evo-devo p. 28 heterochrony 6 heterotopy 7 heterometry 8

Evo-devo eco-evo-devo 67 77 69 heterotypy 9 4 2 Arthur 3 12 Arthur 13 14 16 co-option pp. 264 269 17 Arthur evo-devo p. 273 p. 324 evo-devo 19 Arthur

70 7 2013 p. 320 Evo-devo evo-devo 2.2 Evo-devo? Arthur 2011 evo-devo Hall 1992 20 evo-devo 20 evo-devo Arthur e.g., Wagner 1986 evo-devo 12 BMP4 pp.186 190 4 p. 55

Evo-devo eco-evo-devo 67 77 71 4.6 19 Arthur evo-devo p. 324 evo-devo pp. 324 325 p. 325 Gould and Lewontin 1978 e.g., Mayr and Provine 1980 3 Eco-(evo-)devo Gilbert and Epel [2009]2012 Integrating epigenetics, medicine, and evolution evo-devo evo-devo

72 7 2013 3.1? ecological developmental biology; eco-devo p. v, [2009] p. xii Gilbert and Epel [2009]2012 4 1 p. v, [2009] p. xiii 4 Gilbert and Epel eco-devo 1 eco-devo p. 7, [2009] p. 9 eco-devo Gilbert and Epel eco-devo 1 2 2 3 2 eco-devo eco-devo 1

Evo-devo eco-evo-devo 67 77 73 pp. 206 208, [2009] pp. 256 258 3 Gilbert and Epel evo-devo (ecological evolutionary developmental biology; eco-evo-devo) 3 eco-evo-devo heritable genetic p. 370, [2009] p. 298 eco-evo-devo Ibid. eco-evo-devo Ibid. eco-evo-devo p. 396, [2009] p. 322 eco-devo 2 3 eco-devo 3.2? 3

74 7 2013 3 eco-evo-devo evo-devo Gilbert and Epel ecoevo-devo evo-devo eco-evo-devo evo-devo Hall 1992 pp. 204 208 Hall [1998]2001 19 Gilbert and Epel eco-evo-devo evo-devo Gilbert and Epel [2009]2012 DNA DNA

Evo-devo eco-evo-devo 67 77 75 Smocovitis 1996 20 2 e.g., Cosmides et al. 1992 evolutionary medicine e.g., Nesse and Williams [1996]2001 2 Cosmides et al. 1992 conceptual integration

76 7 2013 4 Arthur 2011 Gilbert and Epel [2009]2012 evo-devo eco-(evo-)devo Arthur 2011 20 evo-devo eco-(evo-)devo evo-devo Amundson, Ron. 2005. The changing role of the embryo in evolutionary thought : Roots of evo-devo Cambridge: Cambridge University Press. Arthur, Wallace. 2011. Evolution : A developmental approach. Chichester: Wiley- Blackwell. Cosmides, Leda, John Tooby, and Jerome Barkow. 1992. Introduction: Evolutionary psychology and conceptual integration. In The adapted mind: Evolutionary psychology and the generation of culture, eds. J. Barkow, L. Cosmides, and J. Tooby, pp. 3 15. New York: Oxford University Press. Francis, Richard C. [2011]2011. [ Epigenetics : The ultimate mystery of inheritance. (New York: W.W. Norton 2011)] Gilbert, Scott F. and Epel, David. [2009]2012 - - [ Ecological developmental biology : Integrating epigenetics, medicine, and evolution. (Sunderland, Massachusetts: Sinauer 2009)]. Gould, Stephen Jay, and Richard C. Lewontin 1979. The spandrels of San Marco and the Panglossian paradigm: A critique of the adaptationist programme. Proceedings

Evo-devo eco-evo-devo 67 77 77 of the Royal Society of London B 205(1161): 581 598. Hall, Brian K. 1992. Evolutionary developmental biology, 1st ed. London: Chapman and Hall.. [1998]2001. [ Evolutionary developmental biology, 2nd ed. (London: Chapman and Hall 1998)]. Mayr, Ernst and William Provine, ed. 1980. The evolutionary synthesis: Perspectives on the unification of biology. Cambridge, MA: Harvard University Press. Nesse, Randolph, M., and George C. Williams. [1996]2001 [ Why we get sick: The new science of Darwinian medicine New York: Vintage, 1996)] Smocovitis, Vassiliki B. 1996. Unifying biology. Princeton, NJ: Princeton University Press. Wagner, Gunter, P. 1986. The systems approach: An interface between development and population genetic aspects of evolution. In Patterns and processes in the history of life, eds. D. M. Raup and D. Jablonski, pp. 149 165. Berlin: Springer-Verlag. 2009 37(5): 235 249.