JourVND3_05PolyanskiiLO

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1 1305ｨｬｨｺ ｨｰ ｨｺ0504ｨｦ ｨｹｨｪ0000ｨｲ04ｨｦ05ｨｪｨ, 2005, , 6ｧ8 3, ｨC367 ｨｬ fl fl; ｹ ｦ003fl0300 ｨｦ07ｨｺ0204ｨ ｨｮ ｨｹ ｨｨ000205ｨｪｨ ｨｮ04ｨｦ05ｨｪｨｲ ｨｺｨｦ04ｨｦ05 ｨｺ02ｨ 00ｨ ｨｮｨ ｨｹ ｨ ｨｺｨ ｨｪ0000ｨｲ04ｨｦ09 01ｨｦｨｺ01 ｨ 07ｨ ｨｨｨｨｨｦ010202ｨｨ02 01ｨｺｨｦ00ｨ ｨｨｨｺｨｦ090005ｨｬｨｺ00 ODDBALL (0500ｨｬｨｮ ｨ ) ｨｨ0702fl , , , apple0ｸ90ｹ ｦ0 016ｦ00200 apple0ｸ8 6 50ｸ apple fl ｦ0apple apple ｦ ｦ0, [email protected] ｨｨ ｦ0 6 5 apple00 6ｦ0010ｷ3090ｸ ｨｨapple0900fl036ｦ ｹ06ｦ ｨ ｦ ｦ apple07006ｦ0 09apple apple0ｸ apple fl CA ｦ003086ｦ0 01apple ｦ fl03 08apple fl 086ｦ0apple6ｦ ｸ8 oddball ｦ ｷ ｸ apple6ｦ apple ｦ000000ｸ apple ｦ0apple ｦ0 08apple090300apple apple ｦ ｹ0000ｸ (34% 0107apple ｸ % ｦ003086ｦ ｸ803). ｨｬ 0ｹ apple apple ｦ00ｹ ｹ apple006ｦ0010ｷ apple- 6 56ｦ0026ｦ ｨC ｨC ｨC 02fl apple apple apple0ｸ ｨC ｨC 02fl apple ｦ003086ｦ0 ｨC 006ｦ0 apple ｦ000030ｸ apple6ｦ ｸ ｦ ｦ0 0ｹ06ｦ006030ｸ ｦ000 6ｦ0apple030ｸ ｸ ｦ ｹ009006ｦ0 006ｦ00ｹ06ｦ ｦ ｦ0 (40ｨC ｦ00ｹ06ｦ0026ｦ ｦ0) ｨC ｰapple6ｦ009applefl 6ｦ0 apple6ｦ ｹ009fl ｱ ｨC ｦ06 56ｦ0026ｦ ｦ ｦ ｦ000030ｸ ｦ000 6ｦ0apple030ｸ8 006ｦ apple0703fl ｸ8036ｦ0. ｨｨapple ｦ06 76ｦ fl, 0ｹ ｹ06ｦ apple ｦ0 apple ｦ ｸ ｸ8 ( apple6ｦ000090ｹ ｦ0fl 08026ｦ ｹ ) 0703apple6ｦ0086ｦ ｸ90ｹ ｦ apple apple ｹ apple ｦ ｸ90ｹ ｸ ｦ0: 086ｦ0apple6ｦ ｦ0 oddball, 01apple , 07apple apple ｹ0000ｸ806 apple , ｦ ｸ ｦ000 6ｦ0apple03-000ｸ ｸ8, apple07000ｸ8 apple6ｦ ｹ009fl 09apple apple0ｸ ｦ003086ｦ0. Limited Plasticity of Rabbit ｯs Visual Cortex and Hippocampal Neurons of Differences V. B. Polyansky, D. V. Evtikhin, E. N. Sokolov, A. V. Kryuchkova Departments of Higher Nervous Activity and Psychophysiology, M.V. Lomonosov State University, Moscow, [email protected] The activity of 41 visual cortex and 20 hippocampal neurons from field CA1 was registered in experiments using oddball-stimulation with different color stimuli varied in intensity. 34% cortical and 37% hippocampal neurons demonstrated plasticity reactions. The significant increase of latest phases of neuronal activity (200ｨC500 and 200ｨC1000 ms after stimulation for cortical neurons and 300ｨC550 ms for hippocampal neurons) was shown in responses to rare deviant stimuli, which had a less intensity than frequently standards. The quantity of the earliest neuronal phase of activity (40ｨC120 ms after stimulation) was stabilized in responses to deviants and standards during the experiment. We propose that such increase of the latest phases of neuronal activity (the limited plasticity) may reflect the mechanisms of orienting reaction. Key words: oddball, rabbit, orienting reflex, deviants and standards, visual cortex and hippocampal neurons of differences. ｨｨ ｦ ｦ0fl 096ｦ ｦ ｷ ｦ0 apple ｦ ｦ0apple apple apple00 01apple ｦ apple07000ｸ8, 006ｦ00ｹ06ｦ ｸ ｦ ｸ806 apple6ｦ009applefl apple0ｸ apple0706 0ｷ apple6ｦ ｹ009fl fl00030ｸ [5]. ｨｮ ｦ , ｦ06 502fl0ｸ90ｸ70903 apple6ｦ009applefl apple6ｦ ｹ009fl, fl096ｦ ｦ ｸ ｹ apple fl, apple0ｸ ｦ000000ｸ ｹ ｹ apple ｦ0006ｦ ｦ0 026ｦ009 09apple ｸ ｦ ｷ3096ｦ0-026ｦ0 (ｨ 05ｨｨ) apple076 56ｦ ｹ ｷ ｸ ｸ8 [3, 6]. ｨｨ apple , 0ｹ00307 apple6ｦ apple6ｦ009applefl 0ｸ8 (50ｨC ｦ ｸ ) apple apple6ｦ ｹ009fl apple fl0ｸ90306fl apple6ｦ ｸ apple fl fl00030ｸ , apple ｷ apple070603apple6ｦ ｦ ｦ003apple090ｷ ｦ ｸ803 apple6ｦ009applefl ｦ apple07006ｦ0 apple6ｦ ｹ009fl 006ｦ0 apple6ｦ ｸ800 0ｷ ｦ0. ｨｴ apple ｷ apple070603apple6ｦ , 07 apple6ｦ009040ｸ90ｸ apple060200apple ｹ000030ｸ8apple apple apple070603apple6ｦ , ｦ ｷ ｸ803 08apple070603apple6ｦ , ｹ ｦ ｷ ｸ apple apple000ｸ apple00000ｷ309apple [5]. ｨｺ6ｦ apple6ｦ009applefl 0ｸ apple apple6ｦ ｹ009fl 036ｦ0apple6ｦ apple ｦ ｸ ｦ ｸ apple6ｦ009applefl 0ｸ8 036ｦ ｦ ｸ ｹ06ｦ ｨｨapple09 360

2 13ｨｦ07ｨｺ0204ｨ ｨｮ ｨｹ ｨｨ000205ｨｪｨ ｨｮ04ｨｦ05ｨｪｨｲ ｨｺｨｦ04ｨｦ apple6ｦ ｦ apple6 56ｦ ｨC ｰapple6ｦ ｱ apple6ｦ009applefl, fl6 502flfl apple0706 apple6ｦ ｹ009fl apple0ｸ ｸ90ｸ , 036ｦ ｦ06 56ｦ fl 06036ｦ ｸ ｦ fl0ｸ76ｦ0fl apple6ｦ ｦ0 006ｦ008apple6ｦ ｦ0 006ｦ0 08apple apple ｦ apple6ｦ apple6ｦ009applefl ｸ8fl ｸ9 096ｦ apple ｦ00ｷ ｦ apple fl apple fl0ｷ ｨｺ006ｦ ｦ00ｷ309fl 0ｹ ｦ0 6ｦ00ｹ0 0ｸ8026ｦ ｦ oddball ｹ06ｦ006000ｸ ｦ ｦ000 6ｦ0apple03000ｸ , ｸ90ｸ ｸ apple07fl ｸ9 (90%) ｦ ｸ803, 006ｦ ｸ apple07fl ｸ9 (10%). ｨｨ6ｦ0apple6ｦ ｦ0 oddball ｦ fl0ｸ apple00-03fl 0409apple apple fl000306fl 016ｦ ｦ00009fl03 006ｦ0 020ｸ9 fl03 [2, 12ｨC17, 21, 22], 036ｦ ｦ ｸ803 [8ｨC11, 18ｨC20] ｦ0006ｦ apple040ｸ90306fl fl apple6ｦ ｸ ｦ ｸ ｦ000000ｸ ｷ3096ｦ ｨｬ06apple ｸ800 05ｨｨ 006ｦ ｦ000030ｸ8 (apple , ｹ06ｦ006000ｸ ｸ8) ｹ06ｦ00ｸ90306fl ｦ ｦ000 6ｦ0apple030ｸ , 0ｹ ｦ ｨｨ ｦ ｦ0fl ｦ0, ｹ ｦ0fl 006ｦ ｦ ｰ ｦ apple6ｦ ｦ ｦ00009fl ｱ ｨC ｰmismatch negativity ｱ [2, 10, 17, 22]. ｨｺ6ｦ0 0703, 6 50ｸ ｸ ｦ apple07006ｦ apple6ｦ003016ｦ003 08apple076 7apple6ｦ003030ｸ8 oddball, [19, 20] ｹ ｦ ｦ00300, 016ｦ001 apple6ｦ ｦ fl 08026ｦ ｹ ｦ0apple6ｦ oddball ｦ0 0603apple apple000ｸ ｷ36ｦ ｦ0 ｨC apple07006ｦ ｦ ｸ8036ｦ ｦ0 09apple apple ｦ ( CA1) 01apple ｦ ｸ apple ｦ apple ｷ ｦ0, 0007 apple6ｦ ｨｪｨｦ05ｨ 0102 ｨｦ080ｸ8030ｸ8 08apple ｸ8 006ｦ apple apple ｦ003 (Orictolagus cuniculus). ｨｦ0800apple6ｦ00ｷ3090ｸ9 08apple ｦ0 5ｨC ｸ ｦ ｸ ｦ0apple ( /016 7) ｦ (2%-000ｸ ｦ00900). 0502fl ｦ ｦ003030ｸ apple07036ｦ fl0307apple6ｦ0 006ｦ0 09apple apple0706 ( ｦ0apple0900) ｦ ( apple ｦ0apple0900) c ｸ7000ｸ9 02apple00090ｸ8 08apple07apple00096ｦ apple apple ｦ apple (010707apple ｦ0030ｸ8 0ｷ apple6ｦ0 02fl 09apple apple0ｸ8: AP = ｨC , L = , 02fl ｦ003086ｦ0: AP = ｨC , L = ) apple0ｸ86 56ｦ00206fl apple ｸ ｦ ｦ ｨｰ6ｦ003030ｸ ｹ000apple00086ｦ0 01apple apple ｸ ｦ003-0ｷ ｦ0 09 6ｦ001apple ｦ ｦ006060ｸ8. ｨ apple ｸ800 0ｹ apple07 0ｸ fl ｸ apple0003fl 08apple070ｷ300 04apple0ｸ8 0ｹ apple ｦ0 01apple ｸ76ｦ00206fl apple006 5fl00000ｸ ｦ , ｦ apple076 56ｦ0026ｦ ｸ7000ｸ ｹ apple ｦ fl ｹ06ｦ , ｦ0096ｦ ｸ apple0ｸ8030ｸ apple046ｦ ｸ ｸ apple ｸ76ｦ00206fl 6 5 0ｹ101apple6ｦ00009apple076 56ｦ ｸ apple076 56ｦ ｸ9 016ｦ00300apple04 006ｦ0 apple6ｦ fl ｹ101apple6ｦ0006ｦ0 0ｷ apple6ｦ0 SVGA (0209apple036ｦ0 TVM, ｨｪ6ｦ ｦ00000). ｨｺ apple6ｦ00ｷ309fl 6ｦ apple ｦ ｦ0apple09fl, 08apple apple6ｦ009 apple6ｦ0086ｦ ｦ ｨｴ010603apple6ｦ ｹ ｦ apple apple ｦ00109apple ｦ000000ｸ apple6ｦ ｸ apple070ｹ apple07 6ｦ ｦ apple ｹ009016ｦ0 1ｨC apple apple ｨC2 02ｨｦ03. ｨｴ apple ｦ ｦ0fl ｦ ｦ fl026ｦ apple apple076 56ｦ000000ｸ ｸ90300apple076 5: (IBM PC-286) fl ｦ0-0ｹ , apple (Pentium-133) ｨC 02fl 0408apple6ｦ fl 0ｹ apple apple6ｦ apple ｦ ｦ00009fl ｦ apple07 000ｸ ｸ ｸ ｹ101apple6ｦ0006ｦ apple6ｦ0, 096ｦ ｹ ｦ00ｸ9-0ｸ ｸ ｹ06ｦ ｦ0096ｦ0 01apple ｦ0. 07apple07036ｦ003090ｹ flapple ｦ00209 apple076 56ｦ ｸ7000ｸ apple apple6ｦ0-flapple apple6ｦ0 ｨｪ01ｨｹ (ｨｺ fl). ｨｦ080ｸ8030ｸ8 08apple fl ｹ ｦ0 6ｦ008036ｦ00ｷ ｨｴ apple ｸ fl apple ｸ , apple0ｸ ｦ00ｹ ｹ06ｦ006030ｸ ｦ000 6ｦ0apple03000ｸ803 ( apple07fl ｸ9 90%) 09 apple ｦ ｸ803 ( apple07fl ｸ9 10%) ｦ ｸ90ｸ ｦ00ｷ30909 apple6ｦ009 apple6ｦ : 6ｦ0) 06036ｦ000 6ｦ0apple030ｸ ｦ000030ｸ ｷ ｦ0, 0007 apple6ｦ : 06036ｦ000-6ｦ0apple030ｸ8 ｨC 8 01 /03 2, ｦ000030ｸ8 ｨC 4ｨC7 01 /03 2 ; ) 06036ｦ000 6ｦ0apple030ｸ ｦ000030ｸ ｷ ｦ ( apple070200). ｨｨapple076 7apple6ｦ003036ｦ fl0ｷ ｦ ｸ8036ｦ fl- 026ｦ : 06006ｦ00ｹ06ｦ0026ｦ0 08apple00 ｨBfl6 502fl ｦ ｨC ｹ0000ｸ ｦ000030ｸ8, 096ｦ ｨC ｹ0000ｸ ｦ000 6ｦ0apple030ｸ8. ｨｨ ｹ ｦ0026ｦ apple070ｷ300 04apple6ｦ0 ｰoddball ｱ: , apple0ｸ803 0ｸ ｦ000036ｦ00309 (10%), 6ｦ0 216 (90%) ｨC 06036ｦ000 6ｦ0apple036ｦ ｸ ｦ06 56ｦ ｹ06ｦ applefl ｷ apple ｦ ｦ06 56ｦ ｹ0000ｸ ｦ ｦ000 6ｦ0apple ｨC ｸ apple6 56ｦ0020ｸ8 ｨC 0.8ｨC ｨ 6ｦ ｦ apple07006ｦ0 08apple ｦ ｹ ｨC5 0ｹ0. ｨｨ07 apple ｦ0036ｦ003 0ｹ apple ｦ0 0603apple apple6ｦ00603apple0ｸ ｸ apple6ｦ003030ｸ apple ｦ ｸ ｦ06 502fl0ｸ90ｸ ｸ8036ｦ ｦ apple076 56ｦ apple ｹ ｦ ｸ90ｸ apple6 56ｦ0-026ｦ003 6ｦ0006ｦ ｦ0: 40ｨC120, 200ｨC500, 200ｨC ｨC fl 06036ｦ ｹ apple6ｦ apple ｦ ｦ0026ｦ apple076 7apple6ｦ003036ｦ0 ｰStatistica for Windows 5.0 ｱ ( ｰStatSoft ｱ, Cｨｰ02). 0502fl 070ｷ apple6ｦ ｹ ｸ8-07apple016ｦ003 08apple fl0206fl ｦ0apple6ｦ apple090ｹ ｦ0apple000ｸ806 01apple090300apple ｦ0. ｨｺ00ｨ ｨｬ00ｨｲｨｪ02ｨｪ01 ｨ ｨｦ050204ｨ 09 ｨ apple ｦ0fl 0107apple6ｦ ｸ ｦ0apple apple09apple076 56ｦ apple ｸ ｦ009000ｸ80309 apple006ｦ0010ｷ309fl0309, apple0ｸ ｸ apple07006ｦ003 ｰapple6ｦ ｹ009fl ｱ [5]. ｨｦ ｸ90ｹ06ｦ00209 apple6ｦ apple6ｦ009applefl (40ｨC ｦ00ｹ06ｦ0026ｦ ｦ0, 0307apple ｸ9 086ｦ , apple090ｹ0000ｸ806 apple6ｦ009applefl ｰ ｦ003070ｹ000040ｸ9 6ｦ ｱ ｨｨapple090300apple apple006ｦ0010ｷ ｦ apple apple ｦ ｦ0 apple

3 13362 ｨｨ0702fl apple ｦ000030ｸ ｦ000 6ｦ0apple030ｸ8 N = ｦ00ｹ06ｦ ｸ800 N = N = 24 Oddball N = N = ｹ0000ｸ800 N = ｨｺ ｨｦ ｸ apple07006ｦ0 6ｧ8 2 09apple apple0ｸ8 01apple ｦ0 006ｦ ｸ ｷ ｦ0. ｨ ｦ ｨC 5 01 /03 2, 06036ｦ000 6ｦ0apple ｨC 8 01 / ｦ0 016ｦ ｦ0 apple00 apple ｦ apple ｦ0096ｦ00007: N ｨC 0ｹ , 6ｦ apple ｸ8 apple6ｦ00603apple0ｸ ｸ apple6ｦ003030ｸ ｨｮ ｦ apple ｦ ｦ0 apple6ｦ0 ｨC 06apple ｹ ｦ apple6ｦ009000ｸ apple6 56ｦ0020ｸ8 6ｦ0006ｦ ｦ0: 250ｨC , 250ｨC apple apple ｦ0096ｦ ｦ00ｹ06ｦ fl0ｷ30909, 0603apple ｨC ｷ fl ｦ0. ｨｨ apple 09006ｦ ｦ0 06apple00 00flfl 0ｹ06ｦ ｦ ｦ , ｦ0 060ｷ ｨC 6 5apple0003fl apple006ｦ0010ｷ30909, ｦ ｦ ｦ00ｹ ｦ0 ｨC ｦ0fl 6ｦ , 0ｹ ｦ ｦ0fl 0ｹ06ｦ ｦ0 apple6ｦ009applefl apple6 56ｦ0026ｦ003 6ｦ0006ｦ ｦ0 200ｨC ｨC ｦ0 020ｸ9 6ｦ fl ｦ ｦ0 oddball ｦ000030ｸ8 ( apple6ｦ ｸ ｦ ｦ0 0ｹ06ｦ006030ｸ ｦ000 6ｦ0apple030ｸ ｹ0000ｸ ｦ000030ｸ8). 02 ｨC ｹ0000ｸ ｦ00ｹ06ｦ ｸ ｦ000030ｸ8, 03 ｨC 006ｦ00ｹ06ｦ ｸ ｹ0000ｸ ｦ000 6ｦ0apple030ｸ8, 05 ｨC ｦ000030ｸ apple076 7apple6ｦ oddball, 07 ｨC 06036ｦ000 6ｦ0apple030ｸ apple076 7apple6ｦ oddball, 05 ｨC ｹ0000ｸ ｹ0000ｸ ｦ000030ｸ8, 00 ｨC ｹ0000ｸ ｹ0000ｸ ｦ000 6ｦ0apple030ｸ8. 046ｦ0 apple ｦ0096ｦ0006ｦ ｦ ｦ apple07006ｦ0 6ｧ apple070ｷ300 04apple00 oddball. ｨ ｦ00ｹ ｦ000 6ｦ0apple ｦ ｸ ｸ /03 2, 6ｦ ｦ00ｹ ｦ ｨC ｸ8, ｸ9 ｨC 5 01 /03 2. ｨｨapple ｦ0096ｦ00300, 0ｹ ｦ0apple ｦ fl 08apple ｦ apple096 50ｸ8016ｦ00009fl ｦ0 (0603. apple6ｦ00603apple0ｸ8) 08apple apple : fl ｦ , fl 06036ｦ000 6ｦ0apple ; 036ｦ ｦ ｸ8fl ｦ ｸ

4 13ｨｦ07ｨｺ0204ｨ ｨｮ ｨｹ ｨｨ000205ｨｪｨ ｨｮ04ｨｦ05ｨｪｨｲ ｨｺｨｦ04ｨｦ ｨｪ6ｦ ｷ36ｦ apple ｹ009000ｸ8 0ｹ06ｦ ｸ ｦ ｦ apple apple apple0ｸ8 01apple ｦ0 006ｦ ｦ000030ｸ ｦ000 6ｦ0apple030ｸ ｸ8036ｦ ｦ ｷ ｦ0, 0007 apple6ｦ ｨｴ ｦ0006ｦ ｦ0, ｸ8 40ｨC ｨC ｨC1000 0ｨC ｦ000030ｸ ｹ0000ｸ ｦ00ｹ06ｦ ｸ8036ｦ ﾀ ﾀ 9.0** 11.6 ﾀ 6.3** 13.2 ﾀ 5.6* ｦ000030ｸ8 oddball 78.1 ﾀ ﾀ ﾀ ﾀ ｦ000030ｸ ｹ0000ｸ ｷ ｸ8036ｦ ﾀ ﾀ 8.0** 11.9 ﾀ ﾀ ｦ000 6ｦ0apple030ｸ ｹ0000ｸ ｦ00ｹ06ｦ ｸ8036ｦ ﾀ ﾀ 10.4* 11.8 ﾀ 6.3* 13.3 ﾀ 6.1* 05036ｦ000 6ｦ0apple030ｸ8 oddball 78.7 ﾀ ﾀ 8.4* 12.1 ﾀ 5.4* 13.9 ﾀ ｦ000 6ｦ0apple030ｸ ｹ0000ｸ ｷ ｸ8036ｦ ﾀ ﾀ ﾀ ﾀ 6.1 ｨｨapple ｹ06ｦ ｨｮ apple apple6ｦ ｨｮ apple0906 ｨC 71. ｨ ｹ0016ｦ ｹ apple ｹ00900 apple006ｦ0010ｷ ｦ0 oddball ｦ000030ｸ8: * ｨC apple < 0.05, ** ｨC p < fl apple6ｦ009applefl ｦ apple ｦ apple00 ｨBfl apple6ｦ009 apple6ｦ ｨｨapple09 6ｦ0006ｦ apple ｦ ｦ , 0ｹ00307, fl ｦ000030ｸ ｹ ｦ0apple6ｦ apple ｦ ｸ ｦ0-0ｸ800, 0ｹ ｦ000 6ｦ0apple030ｸ8, 6ｦ apple07006ｦ apple0ｸ apple0907 0ｸ ｦ0 006ｦ apple ｹ ｦ ｸ80400, 0ｹ ｦ ｦ ｦ000 6ｦ0apple030ｸ8. ｨ , , 06016ｦ0090ｸ86 56ｦ fl ｦ fl fl ｦ , apple ｦ00309, apple apple ｹ apple ｦ0. ｨｨapple apple6ｦ apple0903, 016ｦ fl000306fl 0800apple6 5090ｹ0000ｸ806 apple6ｦ009applefl, , 016ｦ ｸ ｦ ｦ00209 apple6ｦ [5], ｰapple6ｦ009applefl apple6ｦ ｹ009fl ｱ ｦ0 apple ｦ000030ｸ ｦ apple070ｷ300 04apple00 oddball (apple , 05) 06apple00 00flfl 0ｹ06ｦ ｦ ｦ ｨC /06. ｨｴ , 0ｹ ｦ ｦ0 006ｦ00ｹ06ｦ ｸ ｹ0000ｸ ｦ000030ｸ8 (02) ｦ000 6ｦ0apple030ｸ8 (03), , 0ｹ ｦ ｦ000 6ｦ0apple030ｸ apple070ｷ300 04apple00 oddball (07) ｦ ｦ ｹ0000ｸ ｹ0000ｸ ｦ000030ｸ8 (05) ｦ000 6ｦ0apple030ｸ8 (00). ｨｦ 006ｦ ｦ ｦ0apple6ｦ apple fl 6ｦ apple07006ｦ0. 016ｦ ｦ (036ｦ ), ｦ ｹ ｦ00ｹ apple6ｦ ｷ30ｸ8 02fl 0ｹ apple6ｦ009applefl 6ｦ ｦ ｦ0 oddball ｦ000030ｸ ｦ apple ｸ ｦ0apple6ｦ apple000ｸ ｦ apple07006ｦ0 fl6 502fl fl 0307, 0ｹ ｦ0090ｸ ｦ0 006ｦ0 apple oddball ｦ000030ｸ8 (200ｨC600, 200ｨC ) ｦ0090ｸ86 56ｦ00ｸ903 fl ｹ ｸ9 06apple00 000ｸ90ｸ9 0ｹ06ｦ apple6ｦ009applefl 6ｦ0, / / apple6ｦ ｸ ｸ90ｸ ｦ0096ｦ ｦ apple ｸ ｦ apple006ｦ0010ｷ30909, 016ｦ apple07006ｦ0 6ｧ8 2, ｦ0apple ｸ8 6ｦ ｦ0apple apple09apple076 56ｦ000000ｸ apple ｹ apple0909. ｨｮ0307 0ｸ ｸ apple ｦ0030ｸ8, 0ｸ802 08apple ｦ0006ｦ apple006ｦ0010ｷ ｦ000000ｸ ｦ ｸ apple apple apple0ｸ ｦ ｦ0000ｸ apple076ｦ0006ｦ apple076 56ｦ0000ｸ ｸ apple09fl ｸ apple6ｦ009000ｸ fl apple07006ｦ ｦ ｦ0096ｦ002, 0ｹ apple apple ｦ0apple fl apple ｸ ｦ0apple ｸ apple6ｦ ｷ30ｸ ｦ ｦ0 oddball ｦ000030ｸ ｦ ｦ apple apple6ｦ009 apple6ｦ ( ｹ ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8) fl apple6 56ｦ0026ｦ0 6ｦ0006ｦ ｦ0. ｨｨapple ｸ ｦ ｹ ｦ0006ｦ ｦ0apple apple090300apple090ｸ ｦ apple ｹ ｸ ｨｦ 006ｦ apple ｸ ｦ ｦ ｦ000000ｸ apple ｸ8 6 50ｸ , 0ｹ00009 apple006ｦ0010ｷ ｦ0 oddball ｦ000030ｸ ｸ fl 0ｸ apple- 6 56ｦ ｦ0006ｦ ｦ ｦ0096ｦ ｨｮ apple6ｦ ｸ ｦ ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8. ｨｪ6ｦ ｦ0-096ｦ , % ｦ000000ｸ apple ( apple09fl 07080ｸ ). ｨｺ ｦ0030ｸ8 6ｦ0006ｦ ｦ0 apple006ｦ001-0ｷ ｹ apple apple ｦ ｸ ｦ ｨｺ6ｦ apple ｦ ｦ0090ｸ86 56ｦ00003, 0ｹ fl apple6 56ｦ ｨC ｨC apple006ｦ0010ｷ ｦ0 oddball ｦ ｸ apple006ｦ0010ｷ ｦ0 oddball-06036ｦ000 6ｦ0apple- 030ｸ8, ｦ ｹ0000ｸ ｦ00ｹ06ｦ ｸ ｹ0000ｸ ｦ000030ｸ ｦ00ｹ06ｦ ｸ ｦ000 6ｦ0apple030ｸ8 ( apple6 56ｦ002 6ｦ0006ｦ ｦ0 200ｨC ) ｦ0 006ｦ00ｹ06ｦ ｸ ｹ0000ｸ ｦ000 6ｦ0apple030ｸ ｦ000030ｸ8 ( apple6 56ｦ002 6ｦ0006ｦ ｦ0 200ｨC ) ｹ apple08 6ｦ fl 6ｦ000000ｸ ｦ ｹ ｦ0006ｦ ｦ0 apple ｦ ｦ0apple apple090300apple090ｸ ｦ0 (06 04apple ｦ00ｹ ) , 0ｹ fl 0ｹ ｦ ｹ0000ｸ apple ｦ00ｹ06ｦ ｸ806 apple6ｦ009applefl ｦ0 oddball ｦ000030ｸ ｦ fl ｦ003070ｹ ｦ ｸ ｹ06ｦ fl 09006ｦ00ｹ009030ｸ ｹ ｦ apple ｸ , ｹ ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ fl 0ｹ ｸ8 0ｹ apple ｦ0 0ｸ apple ｸ apple ｸ ｸ8030ｸ8 006ｦ apple07006ｦ003, apple0ｸ ｦ000030ｸ ｦ000 6ｦ0apple030ｸ8 0ｸ ｦ ｸ8 (016ｦ ｷ , 036ｦ ). ｨｴ ｸ8030ｸ ｸ8fl ｦ apple6ｦ ｹ apple006ｦ0010ｷ309fl apple ｦ0 apple ｹ06ｦ006030ｸ ｸ ｦ ｸ ｦ apple fl CA ｦ apple ｦ003086ｦ ｦ0096ｦ002, 0ｹ ｦ0apple6ｦ apple ｸ90306fl 026ｦ009000ｸ803 apple6ｦ009applefl ｦ0-0ｹ06ｦ ｸ ｸ ｹ06ｦ ｸ , 0307apple ｦ apple6ｦ009applefl ｨｪ6ｦ apple6ｦ , ｸ apple apple ｦ0 6ｦ00ｸ apple006ｦ0010ｷ309fl apple

5 13364 ｨｨ0702fl apple ｦ000030ｸ ｦ000 6ｦ0apple030ｸ8 N = ｦ00ｹ06ｦ ｸ800 N = N = 24 Oddball N = N = ｹ0000ｸ800 N = ｨｺ ｨｦ ｸ apple07006ｦ0 6ｧ ｦ003086ｦ0 01apple ｦ0 006ｦ ｸ ｷ ｦ0. ｨ ｦ ｨC 7 01 /03 2, 06036ｦ000 6ｦ0apple ｨC 8 01 / , 0ｹ apple00 00flfl 0ｹ06ｦ ｦ ｦ apple6 56ｦ0026ｦ003 6ｦ0006ｦ ｦ0 250ｨC ｨC ｦ ｦ0 oddball ｦ000030ｸ , 0ｹ ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ ｹ0000ｸ ｦ000030ｸ8. ｨｬ 6ｦ apple07006ｦ ｸ8 006ｦ0 apple ｦ000030ｸ apple- 6 56ｦ ｦ0006ｦ ｦ0 40ｨC ｦ0 0ｹ06ｦ006030ｸ ｦ000 6ｦ0apple030ｸ ｹ0000ｸ ｦ000030ｸ8, ｹ10307 fl6 502fl000306fl apple ｸ90ｹ , 0ｹ apple6ｦ ｨｦ ｦ00ｹ fl 016ｦ ｦ0 apple fl CA ｦ003086ｦ0, ｦ000000ｸ80309 apple6ｦ ｨｦ apple6ｦ [23]. 006ｦ ｸ apple0907 0ｸ apple6ｦ009applefl apple6ｦ ｸ apple ｸ apple07006ｦ ｸ700000ｸ apple ｨｮ ｦ00ｹ (apple ) ｦ0apple apple09apple076 56ｦ000000ｸ ｸ ｸ apple07006ｦ003 apple6ｦ ｹ009fl, ｦ0 6ｦ00ｸ90ｸ apple090300apple apple apple ｦ apple6ｦ009applefl 6ｦ0, 0ｹ apple apple6ｦ ｹ009fl 09apple apple0ｸ apple07000ｸ8 apple6ｦ ｹ009fl ｦ003086ｦ ｦ0apple ｦ apple096 50ｸ8016ｦ00009fl. ｨｪ6ｦ001, apple076 56ｦ apple ｦ000 6ｦ0apple03000ｸ ｦ ｸ apple07-006ｦ0 6ｧ ｦ003086ｦ0 (apple ) apple apple6ｦ ｸ9 08apple096 50ｸ8016ｦ00009fl, 6ｦ0 apple006ｦ0010ｷ ｦ0 006ｦ00ｹ06ｦ ｸ apple076 56ｦ ｸ ｦ000 6ｦ0apple03000ｸ ｦ ｸ ｸ

6 13ｨｦ07ｨｺ0204ｨ ｨｮ ｨｹ ｨｨ000205ｨｪｨ ｨｮ04ｨｦ05ｨｪｨｲ ｨｺｨｦ04ｨｦ ｨｪ6ｦ ｷ36ｦ apple ｹ009000ｸ8 0ｹ06ｦ ｸ ｦ c ｦ apple fl 05A ｦ003086ｦ0 01apple ｦ0 006ｦ ｦ000030ｸ ｦ000 6ｦ0apple030ｸ ｸ8036ｦ ｦ ｷ ｦ0, 0007 apple6ｦ ｨｴ ｦ0006ｦ ｦ0, ｸ8 40ｨC ｨC ｨC1000 0ｨC ｦ000030ｸ ｹ0000ｸ ｦ00ｹ06ｦ ｸ8036ｦ ﾀ ﾀ 13.5** 15.0 ﾀ ﾀ ｦ000030ｸ8 oddball 35.9 ﾀ ﾀ ﾀ ﾀ ｦ000030ｸ ｹ0000ｸ ｷ ｸ8036ｦ ﾀ ﾀ 11.8** 14.3 ﾀ ﾀ ｦ000 6ｦ0apple030ｸ ｹ0000ｸ ｦ00ｹ06ｦ ｸ8036ｦ ﾀ ﾀ 15.2** 15.5 ﾀ ﾀ ｦ000 6ｦ0apple030ｸ8 oddball 36.2 ﾀ ﾀ 12.9* 16.1 ﾀ ﾀ 8.6 C036ｦ000 6ｦ0apple030ｸ ｹ0000ｸ ｷ ｸ8036ｦ ﾀ ﾀ ﾀ ﾀ 8.8 ｨｨapple ｹ06ｦ ｨｮ apple apple6ｦ ｨｮ apple0906 ｨC 28. ｨ ｹ0016ｦ ｹ apple ｹ00900 apple006ｦ0010ｷ ｦ0 oddball ｦ000030ｸ8: * ｨC apple < 0.01, ** ｨC p < ｹ06ｦ apple apple046 76ｦ0 (apple , 02, 03) apple ｦ000 6ｦ0apple ｦ fl03 08apple070ｷ300 04apple0ｸ8 oddball (apple , 07) 036ｦ apple006ｦ0010ｷ apple07006ｦ0. ｨｦ 006ｦ apple00 ｨBfl fl ｦ ｦ0 ( ｷ ｦ0, ) ｦ0apple6ｦ oddball (apple0906.2, 05) 08apple apple6ｦ006036ｦ ｸ9 apple006ｦ0010ｷ30909, ｦ apple090ｹ apple6ｦ009applefl 00 (250ｨC ). ｨｮ ｦ ｦ ｦ ｸ ｦ0apple6ｦ oddball apple ｸ apple076 56ｦ apple ｸ ｦ ｦ0 (apple , 02, 05). ｨ 036ｦ001, ｦ0090ｸ ｦ apple07006ｦ0 006ｦ0 oddball ｦ000030ｸ8 04 6ｦ apple07006ｦ0 (apple , 05) , 0ｹ ｸ90ｸ ｹ06ｦ ｦ0 oddball-06036ｦ000 6ｦ0apple- 030ｸ apple076 56ｦ000000ｸ ｦ00ｹ06ｦ ｸ ｹ0000ｸ ｦ000030ｸ ｦ000 6ｦ0apple030ｸ ｹ ｦ ｸ ｹ06ｦ006 (08070ｹ ｸ apple ), ｸ90ｸ ｦ0apple000ｸ806 6ｦ0006ｦ ｦ000000ｸ apple ( apple ｸ , 0ｹ apple apple6ｦ009000ｸ fl fl ｦ ｦ000 6ｦ0apple ) ｸ8fl6 502fl ｦ apple6ｦ ｹ009fl ｦ ｦ0 oddball ｦ000030ｸ8 09 apple006ｦ0010ｷ309fl ｦ0 apple ｸ fl apple6 56ｦ ｦ0006ｦ ｦ0 (08apple090300apple ｦ , 016ｦ fl apple ｸ apple apple apple0ｸ8) ｦ ｹ ｦ0006ｦ apple08 6ｦ ｹ ｦ ｸ90ｹ apple6ｦ00ｸ76ｦ0fl ｦ0006ｦ apple006ｦ0010ｷ apple07006ｦ0 6ｧ8 14 (apple ), apple , 0ｹ ｦ apple apple00 (apple ), 036ｦ ｦ ｸ ｸ ｸ ｸ ｦ ｹ0000ｸ apple07000ｸ8, apple ｸ ｸ9 07apple apple ｹ apple006ｦ0010ｷ ｦ apple0ｸ ｦ0 09fl ｦ apple07006ｦ0 006ｦ0 apple , ｦ000000ｸ ｦ ｸ ｸ8. ｨｴ apple ｸ806 6ｦ0006ｦ ｸ apple ｦ003086ｦ0. ｨｦ016ｦ0096ｦ (016ｦ fl apple apple apple0ｸ8), 0ｹ apple ｦ ｦ0 (37% ｹ ｦ ｦ000000ｸ803) ｦ0apple ｸ ｦ ｹ0000ｸ800 apple006ｦ0010ｷ ｦ ｦ0 oddball ｦ000030ｸ8 (036ｦ ). 056ｦ000000ｸ ｦ fl apple09apple040ｸ903, 0ｹ apple6ｦ009applefl (300ｨC ) 09006ｦ00ｹ ｹ ｦ ｦ0 oddball ｦ000030ｸ apple6ｦ ｸ9 06 apple006ｦ0010ｷ309fl ｦ apple ｸ ｦ ｹ ｦ0006ｦ ｦ0apple apple090300apple090ｸ ｦ ｦ0090ｸ86 56ｦ00003, 0ｹ apple ｹ ｦ ｦ apple6 56ｦ fl ｦ0 oddball ｦ000030ｸ ｦ00ｹ ｹ06ｦ fl 016ｦ ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8 (p < 0.01), 036ｦ ｦ ｹ0000ｸ ｦ00ｹ06ｦ ｸ ｹ0000ｸ ｦ000030ｸ ｦ00ｹ06ｦ ｸ ｹ0000ｸ ｦ000 6ｦ0apple030ｸ8 ( p < 0.001). 046ｦ00ｹ06ｦ ｸ806 apple6ｦ009applefl 6ｦ fl 0ｹ ｦ ｹ0000ｸ apple ｦ fl 070ｹ ｦ ｸ ｦ0apple apple6ｦ ｹ ｦ ｦ0 oddball ｦ000030ｸ8 09 apple ｸ ｦ apple6 56ｦ0026ｦ003 6ｦ0006ｦ ｦ0 200ｨC ｨC ｨｦ0305ｨｬ ｨ 00 ｨｺ00ｨ ｨｬ00ｨｲｨｪ02ｨｪｨｦ05 ｨｨapple ｸ ｸ8030ｸ ｦ0096ｦ00209, 0ｹ apple090300apple0007 1/3 026ｦ009000ｸ apple apple apple0ｸ8 (34%) ｦ003086ｦ0 (37%) 01apple ｦ ｦ0090ｸ86 56ｦ00ｸ ｹ ｦ0 006ｦ ｦ000030ｸ apple070ｷ300 04apple00 oddball ( ｹ06ｦ006000ｸ ｦ ｸ8). 0502fl 08026ｦ ｹ0000ｸ apple apple apple0ｸ ｹ apple006ｦ0010ｷ apple apple6 56ｦ0026ｦ ｨC ｨC (apple , 036ｦ ), 02fl apple ｦ003086ｦ0 ｨC 300ｨC (apple , 036ｦ ) ｦ0apple apple6ｦ ｹ0000ｸ apple09fl ｸ ｦ0-0ｹ06ｦ ｸ806 apple6ｦ009applefl (40ｨC ｦ00ｹ06ｦ0026ｦ ｦ0) 016ｦ001 02fl apple apple0ｸ8, 036ｦ fl CA ｦ003086ｦ ｦ0096ｦ ｦ00ｹ fl ｹ009000ｸ8 apple6ｦ009applefl 6ｦ0 006ｦ0 oddball ｦ000030ｸ apple6ｦ ｸ ｦ ｦ0 oddball-06036ｦ000 6ｦ0apple030ｸ8 (036ｦ ). 05 0ｷ ｦ00ｹ06ｦ ｸ806 apple6ｦ009applefl ｦ003070ｹ ｦ apple09fl ｸ ｦ001 02fl 07 0ｸ apple ｸ ｦ000000ｸ apple , 036ｦ fl 6 7apple ｸ ｦ ｹ0000ｸ ｦ001 0ｸ ｦ0apple apple00 0ｸ8 040ｸ70006 apple6ｦ [5], 0107apple ｦ ｸ ｦ00ｹ06ｦ ｸ806 apple6ｦ009applefl ｦ ｦ00300 apple6ｦ009applefl apple6ｦ ｹ009fl, apple6ｦ0086ｦ apple036ｦ00ｷ3090ｸ9 07 apple6ｦ ｹ009fl ｦ06 56ｦ000030ｸ ｦ ｦ ｸ ｦ ｸ apple , 0ｹ1036ｦ0 026ｦ0096ｦ ｦ apple036ｦ00ｷ3090ｸ ｦ0 [1].

7 13366 ｨｨ0702fl apple apple07000ｸ8 apple6ｦ ｹ009fl (09apple ) 0107apple0ｸ apple07000ｸ8 apple6ｦ ｹ009fl ｦ003086ｦ ｦ0 6ｦ00ｸ ｹ apple apple ｦ apple6ｦ009applefl 6ｦ0, apple ｸ ｹ06ｦ ｸ806 apple6ｦ009applefl fl6 502fl000306fl 0300apple apple6ｦ ｹ009fl [5]. ｨｴ ｦ00ｹ06ｦ00003, 0ｹ apple07000ｸ8 apple6ｦ ｹ009fl 08apple ｸ apple00, ｦ (08apple CA1). ｨｺ006ｦ0010ｷ apple apple6ｦ ｹ009fl ｦ0apple ｦ00ｸ903 apple6ｦ fl 08apple096 50ｸ8016ｦ00009fl, applefl 006ｦ apple6ｦ ｸ fl0ｷ3090ｸ9 ( ) fl apple006ｦ0010ｷ ｦ0 6 50ｸ apple07fl03000ｸ ｦ000 6ｦ0apple03000ｸ fl03 oddball ｦ0066ｦ00ｸ903, apple006ｦ0010ｷ ｦ0 apple ｦ ｸ ｸ ｹ ｦ0-0ｸ90306fl 09 0ｹ ｹ ｹ06ｦ ｹ apple6ｦ000fl000306fl 08apple apple076 56ｦ apple ｦ ｦ ｷ ｸ8036ｦ0. 016ｦ ｦ apple6ｦ006036ｦ00009fl apple006ｦ0010ｷ ｦ0 apple ｦ ｸ ｸ8 08apple apple096 50ｸ8016ｦ00009fl apple006ｦ0010ｷ ｦ0 086ｦ0apple6ｦ ｦ ｸ ｦ000 6ｦ0apple03000ｸ ｸ8? ｸ ｨBfl apple- 0309apple076 56ｦ ｦ003fl ｦ ｦ000 6ｦ0apple03000ｸ , ｦ06 56ｦ000030ｸ ｸ apple07fl ｸ ｦ0apple6ｦ oddball. ｨｨapple ｦ000 6ｦ0apple ｦ ｦ0 6ｦ , apple6ｦ apple6ｦ ｦ ｸ apple07006ｦ0 apple6ｦ ｹ009fl 0708apple fl000306fl apple6ｦ ｸ ｦ000 6ｦ0apple03000ｸ ｨｨapple ｦ ｦ , ｦ ｸ ｦ000-6ｦ0apple03000ｸ , ｦ0 6ｦ ｦ ｸ ｨ 03 apple6ｦ ｹ ｦ ｸ ｦ002, apple0ｸ apple fl ｦ apple0ｷ apple6ｦ ｹ009fl, 6 50ｸ ｦ000000ｸ ｦ ｦ apple , 0ｹ ｦ ｦ0 006ｦ ｦ ｸ fl03 oddball 6ｦ0006ｦ ｹ ｦ apple6ｦ ｦ ｦ0-0009fl, ｦ0apple apple ｨｺ. 046ｦ06ｦ0036ｦ ｦ0 [2, 12ｨC 17] 08apple ｦ ｸ ｨｨapple , 0ｹ ｹ ｦ apple6ｦ009applefl 6ｦ0 006ｦ ｦ ｸ ｸ8 6ｦ0006ｦ ｹ ｦ ｦ apple6ｦ ｦ ｦ00009fl, 06036ｦ ｸ apple apple ｸ ｦ ｦ apple0309apple ｦ00009fl ｦ apple ｦ ｦ ｹ fl ｦ0 086ｦ003fl ｸ90ｸ ｦ0 006ｦ ｦ fl03 oddball 6 50ｸ80ｹ ｦ apple076 56ｦ000000ｸ ｦ00003, ｹ ｹ apple6ｦ ｹ009fl ｦ ｦ000 6ｦ0apple ｦ ｨｮ ｹ ｦ000036ｦ ｦ000 6ｦ0apple036ｦ0, ｦ0016ｦ0fl apple6ｦ ｨｴ ｦ ｸ806 apple6ｦ009applefl apple6ｦ ｹ009fl ｦ000 6ｦ0apple ｦ ｦ00003 MMN (mismatch negativity, ｦ apple6ｦ ｦ ｦ0-0009fl), ｦ0apple ｸ9 ｨｺ. 046ｦ06ｦ0036ｦ [2] 08apple ｦ ｸ ｦ06 502flfl ｹ ｸ ｦ ｦ000000ｸ ｦ000000ｸ80309 ｨｺ. 046ｦ06ｦ0036ｦ ｦ0, ｦ ｸ ｦ003 apple6ｦ ｹ00906: 0800apple- 0ｷ ｸ800 apple6ｦ ｹ009fl applefl0ｸ90306fl apple6ｦ ｦ ｸ803 apple6ｦ009applefl 0703, apple0ｸ appleapple000209apple ｦ ｨｨ N ｦ ｸ806 apple6ｦ009applefl 250ｨC ｦ ｦ000030ｸ8 fl6 502fl000306fl apple090ｹ apple6ｦ009applefl 6ｦ0 09 apple6ｦ ｹ009fl ｦ000036ｦ ｦ000 6ｦ0apple036ｦ0, apple0700 apple6ｦ N2 + MMN. ｨｪ6ｦ apple6ｦ , 01apple apple0ｷ ｸ803 apple6ｦ ｹ00906 (N1) 6 50ｸ80ｹ fl000306fl 000ｸ700 apple6ｦ ｹ ｦ000 6ｦ0apple036ｦ0 09 6ｦ ｦ ｸ ｦ ｦ000000ｸ800, ｹ ｸ ｹ apple6ｦ apple07-006ｦ003, ｦ0 6ｦ00ｸ ｦ apple ｦ0-036ｦ00309, ｹ ｸ ｦ0006ｦ ｹ0000ｸ ｸ8036ｦ apple apple6ｦ00ｷ ｸ ｦ000000ｸ ｷ3096ｦ ｦ0 01apple [4]. ｨｺ6ｦ ｸ ｦ0096ｦ00007, 0ｹ apple apple ｦ0 apple ｦ000030ｸ8 08apple ｹ ｦ ｸ P1(56ｨC ) 09 P2(120ｨC ), 6ｦ ｦ ｨC P1, N1, P2 (80ｨC ) apple6ｦ ｸ ｦ ｦ0 oddball ｦ000030ｸ ｹ0000ｸ ｦ000030ｸ8. 016ｦ ｨBfl ｹ ｸ ? 020ｸ ｦ06 76ｦ00003, 0ｹ ｸ ｦ ｷ3096ｦ0026ｦ ｦ ｦ ｹ ｷ3096ｦ apple , apple07fl6 502fl030006fl fl0ｷ apple07000ｸ8 apple flapple000ｸ apple apple apple apple0003fl , 0ｹ ｦ ｸ806 apple6ｦ009applefl apple07006ｦ ｦ apple apple apple fl apple6ｦ ｷ3096ｦ0026ｦ0, ｦ00ｹ06ｦ ｸ ｦ009000ｸ806 apple6ｦ009applefl 0807fl fl, ｸ ｹ ｸ803, 0800apple00 6ｦ06 56ｦ0fl fl ｹ ｦ apple6ｦ ｹ009fl ｦ ｨｬ ｹ apple006ｦ0010ｷ apple ｦ0 apple ｦ ｸ ｸ apple070ｷ300 04apple00 oddball 016ｦ0066ｦ fl ｦ0 (200ｨC500, 200ｨC ), 0ｹ apple0903, , 07 apple apple flapple000ｸ803, ｹ apple apple ｸ fl ｦ0 6ｦ apple apple0ｸ ｦ003086ｦ0. ｨｴ fl030007, ｦ0090ｸ apple apple6ｦ0086ｦ0-0ｸ apple apple070ｷ ｸ8, fl0ｸ fl ｸ ｷ309fl03 [1]. ｨｨ fl , ｦ00309apple076 56ｦ00300, 0ｹ apple07000ｸ8 09apple apple0ｸ fl CA ｦ003086ｦ0 01apple ｦ ｦ0096ｦ apple6ｦ000090ｹ ｸ ｦ ｹ , 0ｹ ｸ8apple6ｦ0086ｦ fl, apple6 50ｸ803, , 0ｹ ｦ ｹ ｸ ｦ0 6ｦ00ｸ apple07000ｸ8, 6ｦ0 08apple090300apple apple , 09, apple0ｸ803, 04 0ｹ ｦ ｹ0000ｸ apple ｹ ｹ06ｦ ｦ0 apple ｦ ｸ ｸ apple070ｷ300 04apple00 oddball apple6ｦ ｸ ｦ ｦ0 0ｹ06ｦ006030ｸ ｦ000-6ｦ0apple030ｸ8. ｨｨ07 0ｷ applefl 04 08apple ｦ ｹ apple6ｦ009applefl 6ｦ apple07fl ｸ9 07apple apple ｹ apple ｦ0 006ｦ ｦ apple00 ｨBfl ｦ0 [7]. ｨｬ ｹ06ｦ ｦ apple6ｦ ｦ ｦ ｹ apple00apple6ｦ apple036ｦ00ｷ apple apple00 6ｦ00ｹ apple ｹ06ｦ ｦ ｦ ｸ apple6ｦ apple apple ｦ003fl ｨｦ ｨｨapple ｦ0apple6ｦ ｸ8 oddball ( ｹ06ｦ ｦ ｸ8) ｦ ｷ ｸ803 09apple ｸ apple6ｦ ｦ0096ｦ ｦ00ｹ ｹ apple006ｦ0010ｷ ｦ ｹ0000ｸ ｦ0 apple ｦ000030ｸ apple6ｦ ｸ ｦ ｦ0 0ｹ06ｦ006030ｸ ｦ000 6ｦ0apple030ｸ apple6 56ｦ0026ｦ ｨC500 09

8 13ｨｦ07ｨｺ0204ｨ ｨｮ ｨｹ ｨｨ000205ｨｪｨ ｨｮ04ｨｦ05ｨｪｨｲ ｨｺｨｦ04ｨｦ ｨC ｦ00ｹ06ｦ0026ｦ ｦ0 ｨC 02fl apple apple apple0ｸ ｨC ｨC 02fl apple fl CA ｦ003086ｦ0 01apple ｦ ｹ009006ｦ0 0800apple ｦ0 apple6ｦ009applefl 6ｦ apple (40ｨC ) apple09fl ｸ apple apple0ｸ ｦ003086ｦ ｦ ｦ0 oddball ｦ000030ｸ ｦ fl 06036ｦ ｦ ｦ ｦ0006ｦ ｦ ｹ ｸ803 6ｦ000000ｸ803 08apple ｦ06 76ｦ fl, 0ｹ ｹ06ｦ apple ｦ0 apple ｦ ｸ ｸ8 ( apple6ｦ000090ｹ ｦ0fl 08026ｦ ｹ ) 0703apple6ｦ0086ｦ ｸ90ｹ ｦ apple apple ｹ apple ｦ0. ｨｺ6ｦ ｦ0 6 50ｸ ｦ0 08apple apple ｨｺ ｦ ｦ ｦ ｸ ｦ (6 7apple6ｦ ｧ ), 6ｦ0 036ｦ apple apple6ｦ0036ｦ0 0207ｨｬ 02fl ｸ803 08apple ｦ06 56ｦ ｹ000000ｸ803 ( ). 05ｨｨｨ 05ｨｦ01 00ｨ ｨｪ00ｨｺ02ｨｪｨｬｨｺ01 1. ｨ 6 56ｦ000090ｷ ｸ ｦ ｸ8 070ｷ ｦ : ｷ309006ｦ0, ｦ06ｦ0036ｦ ｨｺ ｦ ｷ ｦ0. 02.: ｨ ｨｬ, ｨｨ0702fl , , ｨｺ apple04010ｷ309fl 0800apple0ｷ ｸ803 08apple070603apple6ｦ flapple ｷ ｦ0 01apple ｦ0 006ｦ apple ｸ ｸ ｦ000000ｸ ｷ3096ｦ apple6ｦ ｦ000000ｸ ｹ ｸ // 0504apple ｸ apple fl ｨｪ ｧ ｨC ｨｨ0702fl , , ｨｦ03apple6ｦ apple apple ｹ apple ｦ ｦ0096ｦ ｸ ｦ000000ｸ ｷ3096ｦ apple apple ｦ apple ｦ0 08apple09 096ｦ // 0504apple ｸ apple fl ｨｪ ｧ ｨC ｨｨ0702fl , , ｸ80ｹ ｷ ｸ ｦ003apple07036ｦ003090ｹ apple6ｦ ｹ apple07006ｦ apple apple0ｸ8 01apple ｦ0 // 0504apple ｸ apple fl ｨｪ ｧ8 1. C. 60ｨC ｨｨ0702fl , , 026ｦ0apple0ｹ ｨｪ.04., , ｨｺ04 00apple036ｦ ｨｨ00apple0ｷ ｷ apple070603apple6ｦ apple ｦ0 // 0504apple ｸ apple fl ｨｪ ｧ ｨC , ｦ0 04.ｨ., ｨｨ0702fl , ｨｦapple apple ｹ0000ｸ806 apple : apple006ｦ0010ｷ309fl 08apple090ｷ ｦ00009fl 09 08apple apple ｦ00009fl // 0504apple ｸ apple fl ｨｪ ｧ ｨC Astikainen P., Ruusivirta T., Korhonen T. Cortical and subcortical visual event-related potentials to oddball stimuli in rabbits // Neuroreport V ｧ8 7. P. 1515ｨC Astikainen P., Ruusivirta T., Korhonen T. Somatosensory event-related potentials in the rabbit cerebral and cerebellar cortices: a correspondence with mismatch responses in humans // Neurosci. Lett V P. 222ｨC Csepe V. On the origin and development of the mismatch negativity // Ear Hear V ｧ8 1. P. 91ｨC Csepe V., Karmos G., Molnar M. Effect of signal probability on sensory evoked potentials in cats // Int. J. Neurosci V ｧ8 1ｨC2. P. 61ｨC Kenemans J.L., Verbaten M.N., Melis C.J., Slangen J.L. Visual stimulus change and the orienting reaction: event-related potential evidence for a two-stage process // Biol. Physiol V ｧ8 2ｨC3. P. 97ｨC Kenemans J.L., Verbaten M.N., Roelofs J.W., Slangen J.L. ｰInitials ｱ and ｰchange-orienting reactions ｱ: an analysis based on visual single-trial event-related potentials // Biol. Physiol V ｧ8 3. P. 199ｨC Naatanen R. Mismatch negativity (MMN): perspectives for application // Int. J. Psychophysiol V ｧ8 1. P. 3ｨC Naatanen R., Alho K. Mismatch negativity ｨC a unique measure of sensory processing in audition // Int. J. Neurosci V ｧ8 1ｨC4. P. 317ｨC Naatanen R., Gaillard A.W.K., Mantysalo S. Early selective attention effect on evoked potential reinterpreted // Acta Psychol V. 42. P. 313ｨC Naatanen R., Paavilainen P., Tirtinen H., Jiang D., Alho K. Attention and mismatch negativity // Psychophysiology V ｧ8 5. P. 436ｨC Ruusivirta T., Korhonen T., Arikoski J., Kivirikko K. ERP to pitch changes: a result of reduced responses to standard tones in rabbits // Neuroreport V. 7. 6ｧ8 2. P. 413ｨC Ruusivirta T., Korhonen T., Arikoski J., Kivirikko K. Multiple ｨC unit responses to pitch changes in rabbits // Neuroreport V. 7. 6ｧ8 7. P. 1226ｨC Shinba T. Neuronal firing activity in the dorsal hippocampus during the auditory discrimination oddball task in awake rats: relation to event-related potential generation// Brain Res Cogn. Brain Res V. 8. 6ｧ8 3. P. 241ｨC Shinozaki N., Jabe H., Takeyuki S., Sutoh T., Hiruma T., Kaneki S. Somatosensory automatic responses to deviant stimuli // Brain Res Cogn. Brain Res V. 7. P. 165ｨC Tales A., Newton P., Troscianko T., Butler S. Mismatch negativity in the visual modality // Neuroreport V ｧ8 16. P. 3363ｨC Vinogradova O.S. Hippocampus as comparator: role of the two output systems of the hippocampus in selection and registration of information // Hippocampus V ｧ8 5. P. 578ｨC598.